Project 3448: C. Weirauch, R. T. Schuh, G. Cassis, W. C. Wheeler. 2019. Revisiting habitat and lifestyle transitions in Heteroptera (Insecta: Hemiptera): insights from a combined morphological and molecular phylogeny. Cladistics. 35 (1):67-105.
Abstract
Heteroptera, the true bugs, are part of the largest clade of non‐holometabolous insects, the Hemiptera, and include > 42 000 described species in about 90 families. Despite progress in resolving phylogenetic relationships between and within infraorders since the first combined morphological and molecular analysis published in 1993 (29 taxa, 669 bp, 31 morphological characters), recent hypotheses have relied entirely on molecular data. Weakly supported nodes along the backbone of Heteroptera made these published phylogenies unsuitable for investigations into the evolution of habitats and lifestyles across true bugs. Here we present the first combined morphological and molecular analyses of Heteroptera since 1993, using 135 taxa in 60 families, 4018 aligned bp of ribosomal DNA and 81 morphological characters, and various analytical approaches. The sister‐group relationship of the predominantly aquatic Nepomorpha with all remaining Heteroptera is supported in all analyses, and a clade formed by Enicocephalomorpha, Dipsocoromorpha and Gerromorpha in some. All analyses recover Leptopodomorpha + (Cimicomorpha + Pentatomomorpha), mostly with high support. Parsimony‐ and likelihood‐based ancestral state reconstructions of habitats and lifestyles on the combined likelihood phylogeny provide new insights into the evolution of true bugs. The results indicate that aquatic and semi‐aquatic true bugs invaded these habitats three times independently from terrestrial habitats in contrast to a recent hypothesis. They further suggest that the most recent common ancestor of Heteroptera was predacious, and that the two large predominantly phytophagous clades (Trichophora and Miroidea) are likely to have derived independently from predatory ancestors. We conclude that by combining morphological and molecular data and employing various analytical methods our analyses have converged on a relatively well‐supported hypothesis of heteropteran infraordinal relationships that now requires further testing using phylogenomic and more extensive morphological datasets.Read the article »
Article DOI: 10.1111/cla.12233
Project DOI: 10.7934/P3448, http://dx.doi.org/10.7934/P3448
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 10796 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 81 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 219 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:80/1 |
Currently Viewing:
MorphoBank Project 3448
MorphoBank Project 3448
- Creation Date:
04 April 2019 - Publication Date:
19 April 2019 - Matrix downloads: 11
This research
supported by
Authors' Institutions
- University of California Riverside
- American Museum of Natural History (AMNH)
- University of New South Wales
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| MorphoBank Curator Project Administrator | 135 | 1 | 1 | 81 | 0 | 0 | 10796 (10504, 0, 292) | 0 | 0 | 0 | ||||
| Gerasimos Cassis Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| Randall Schuh Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| Christiane Weirauch Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| Ward Wheeler Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M25988' (135 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] Aeneolamia contigua Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [2] Locris rubra Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [3] Tropidocephala sp Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [4] Amerzanna peruana Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [5] Putoniessa rivularis Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [6] Cicadatra atra Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [7] Hackeriella veitchi Taxon name last Modified on 04/04/19 | 2 | 72 | 72 | 0 | 7 | 0 | 0 | 5 |
| [8] Peloridium hammonniorum Taxon name last Modified on 04/04/19 | 2 | 72 | 72 | 0 | 7 | 0 | 0 | 5 |
| [9] Aenictopecheidae sp Taxon name last Modified on 04/04/19 | 2 | 76 | 76 | 0 | 3 | 0 | 0 | 5 |
| [10] Systelloderes sp Taxon name last Modified on 04/04/19 | 1 | 77 | 77 | 0 | 3 | 0 | 0 | 5 |
| [11] Enicocephalinae sp Taxon name last Modified on 04/04/19 | 2 | 76 | 76 | 0 | 3 | 0 | 0 | 5 |
| [12] Enicocephalinae nig 137 Taxon name last Modified on 04/04/19 | 2 | 76 | 76 | 0 | 3 | 0 | 0 | 5 |
| [13] Enicocephalus sp cf sturmi ecu 150 Taxon name last Modified on 04/04/19 | 2 | 76 | 76 | 0 | 3 | 0 | 0 | 5 |
| [14] Ceratocombus australiensis Taxon name last Modified on 04/04/19 | 3 | 76 | 76 | 0 | 2 | 0 | 0 | 5 |
| [15] Ceratocombus vagans 247 Taxon name last Modified on 04/04/19 | 3 | 76 | 76 | 0 | 2 | 0 | 0 | 5 |
| [16] Cryptostemma sp Taxon name last Modified on 04/04/19 | 3 | 77 | 77 | 0 | 1 | 0 | 0 | 5 |
| [17] Cryptostemma sp peru 251 Taxon name last Modified on 04/04/19 | 3 | 77 | 77 | 0 | 1 | 0 | 0 | 5 |
| [18] Hypselosoma hickmani Taxon name last Modified on 04/04/19 | 3 | 77 | 77 | 0 | 1 | 0 | 0 | 5 |
| [19] Pateena polymitarior Taxon name last Modified on 04/04/19 | 3 | 77 | 77 | 0 | 1 | 0 | 0 | 5 |
| [20] Voragocoris schuhi 254 Taxon name last Modified on 04/04/19 | 3 | 77 | 77 | 0 | 1 | 0 | 0 | 5 |
| [21] Chinannus sp cr 80 Taxon name last Modified on 04/04/19 | 3 | 77 | 77 | 0 | 1 | 0 | 0 | 5 |
| [22] Ptenidiophyes mirabilis cr 89 Taxon name last Modified on 04/04/19 | 3 | 75 | 75 | 0 | 3 | 0 | 0 | 5 |
| [23] Mesovelia mulsanti Taxon name last Modified on 04/04/19 | 3 | 75 | 75 | 0 | 3 | 0 | 0 | 5 |
| [24] Macrovelia horni Taxon name last Modified on 04/04/19 | 7 | 71 | 71 | 0 | 3 | 0 | 0 | 5 |
| [25] Microvelia sp Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [26] Aquarius remigis Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [27] Abedus indentatus Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [28] Ranatra sp Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [29] Nerthra adspersa Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [30] Megochterus occidentalis Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [31] Corixidae sp Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [32] Usingerina moapensis Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [33] Hydrotrephes sp Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [34] Notonectidae sp peru Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [35] Pleidae sp peru Taxon name last Modified on 04/04/19 | 0 | 77 | 77 | 0 | 4 | 0 | 0 | 5 |
| [36] Salda sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [37] Saldula brevicornis Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [38] Valleriola sp Taxon name last Modified on 04/04/19 | 4 | 76 | 76 | 0 | 1 | 0 | 0 | 5 |
| [39] Leotichius sp Taxon name last Modified on 04/04/19 | 4 | 76 | 76 | 0 | 1 | 0 | 0 | 5 |
| [40] Aphelonotus fraterculus 1586 Taxon name last Modified on 04/04/19 | 7 | 74 | 74 | 0 | 0 | 0 | 0 | 5 |
| [41] Camarochilus 2720 Taxon name last Modified on 04/04/19 | 7 | 74 | 74 | 0 | 0 | 0 | 0 | 5 |
| [42] Phymata pacifica Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [43] Macrocephalus barberi Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [44] Tapeinus sp Taxon name last Modified on 04/04/19 | 3 | 76 | 76 | 0 | 2 | 0 | 0 | 5 |
| [45] Zelurus sp Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [46] Triatoma protracta Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [47] Apiomerus lanipes Taxon name last Modified on 04/04/19 | 0 | 76 | 76 | 0 | 5 | 0 | 0 | 5 |
| [48] Rhynocoris segmentarius Taxon name last Modified on 04/04/19 | 0 | 76 | 76 | 0 | 5 | 0 | 0 | 5 |
| [49] Stenopoda sp Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [50] Racelda sp Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [51] Stenolemoides arizonensis Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [52] Loricula elegantula Taxon name last Modified on 04/04/19 | 6 | 74 | 74 | 0 | 1 | 0 | 0 | 5 |
| [53] Joppeicus paradoxus Taxon name last Modified on 04/04/19 | 4 | 76 | 76 | 0 | 1 | 0 | 0 | 5 |
| [54] Himacerus apterus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [55] Nabis ferus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [56] Nabis apicalis Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [57] Prostemma hilgendorfii Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [58] Lasiochilus japonicus Taxon name last Modified on 04/04/19 | 6 | 74 | 74 | 0 | 1 | 0 | 0 | 5 |
| [59] Lasiochilus luceonotatus Taxon name last Modified on 04/04/19 | 6 | 74 | 74 | 0 | 1 | 0 | 0 | 5 |
| [60] Lipokophila eberhardi Taxon name last Modified on 04/04/19 | 7 | 73 | 73 | 0 | 1 | 0 | 0 | 5 |
| [61] Lyctocoris beneficus Taxon name last Modified on 04/04/19 | 6 | 74 | 74 | 0 | 1 | 0 | 0 | 5 |
| [62] Anthocoris confusus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [63] Orius horvathi Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [64] Tetraphleps aterrimus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [65] Cimex lectularius Taxon name last Modified on 04/04/19 | 3 | 74 | 74 | 0 | 4 | 0 | 0 | 5 |
| [66] Curalium cronini Taxon name last Modified on 04/04/19 | 5 | 73 | 73 | 0 | 3 | 0 | 0 | 5 |
| [67] Baclozygum brachypterum Taxon name last Modified on 04/04/19 | 4 | 75 | 75 | 0 | 2 | 0 | 0 | 5 |
| [68] Thaumastocorinae thai 2846 Taxon name last Modified on 04/04/19 | 4 | 75 | 75 | 0 | 2 | 0 | 0 | 5 |
| [69] Discocoris drakei Taxon name last Modified on 04/04/19 | 4 | 75 | 75 | 0 | 2 | 0 | 0 | 5 |
| [70] Xylastodoris luteolus Taxon name last Modified on 04/04/19 | 4 | 75 | 75 | 0 | 2 | 0 | 0 | 5 |
| [71] Corythucha sp Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [72] Diplocysta sp Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [73] Eteoneus angulatus Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [74] Myiomma sp Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [75] Monalocoris americanus Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [76] Campyloneura virgula Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [77] Dicyphus pallidicornis Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [78] Tupiocoris sp Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [79] Phallospinophylus setosus Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [80] Plagiognathus chrysanthemi Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [81] Rhinacloa forticornis Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [82] Semium hirtum Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [83] Pilophorus discretus Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [84] Lopidea bullata Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [85] Pseudopsallus angularis Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [86] Capsus ater Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [87] Cyphopelta modesta Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [88] Irbisia brachycera Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [89] Lygus lineolaris Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [90] Trigonotylus sp Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [91] Deraeocoris brevis Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [92] Calisius hackeri Taxon name last Modified on 04/04/19 | 1 | 78 | 78 | 0 | 2 | 0 | 0 | 5 |
| [93] Dysodius sp Taxon name last Modified on 04/04/19 | 1 | 78 | 78 | 0 | 2 | 0 | 0 | 5 |
| [94] Isodermus planus Taxon name last Modified on 04/04/19 | 1 | 78 | 78 | 0 | 2 | 0 | 0 | 5 |
| [95] Mezira sayi Taxon name last Modified on 04/04/19 | 1 | 78 | 78 | 0 | 2 | 0 | 0 | 5 |
| [96] Anischys luteovarius Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [97] Stauralia chloracantha Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [98] Lestonia haustorifera Taxon name last Modified on 04/04/19 | 0 | 78 | 78 | 0 | 3 | 0 | 0 | 5 |
| [99] Parastrachia japonensis Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [100] Cydninae sp 3 Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [101] Brochymena sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [102] Oechalia schellenbergii Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [103] Poecilometis sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [104] Tholosanus sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [105] Austrotichus rugosus Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [106] Choerocoris variegatus Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [107] Coleotichus costatus Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [108] Thaumastella namaquensis Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [109] Urochela luteovaria Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [110] Urostylis westwoodi Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [111] Henicocoris monteithi Taxon name last Modified on 04/04/19 | 1 | 78 | 78 | 0 | 2 | 0 | 0 | 5 |
| [112] Trisecus sp Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [113] Kleidocerys sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [114] Lygaeus equestris Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [115] Oncopeltus fasciatus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [116] Malcus elongatus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [117] Pachygrontha sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [118] Mcateella sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [119] Neoneides muticus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [120] Pronotacantha annulata Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [121] Ischnodemus sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [122] Cymus sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [123] Germalus sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [124] Remaudiereana inornata Taxon name last Modified on 04/04/19 | 0 | 81 | 81 | 0 | 0 | 0 | 0 | 5 |
| [125] Laryngodus cervantes Taxon name last Modified on 04/04/19 | 0 | 81 | 81 | 0 | 0 | 0 | 0 | 5 |
| [126] Leptocorisa acuta Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [127] Cletus sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [128] Aulacosternum nigrorubrum Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [129] Narnia femorata Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [130] Maevius indecorus Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [131] Harmostes sp Taxon name last Modified on 04/04/19 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 5 |
| [132] Dysdercus fuscomaculatus Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [133] Probergrothius sexpunctatus Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [134] Largus rufipennis Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
| [135] Physopelta melanoptera Taxon name last Modified on 04/04/19 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 5 |
Project downloads
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 163 | |
| Project downloads | 152 | |
| Matrix downloads | 11 | Morphological matrix used for the phylogenetic analyses NEXUS file. (11 downloads); |