Project 3512: R. Robles, P. C. Dworschak, D. L. Felder, G. C. Poore, F. L. Mantelatto. 2020. A molecular phylogeny of Callianassidae and related families (Crustacea : Decapoda : Axiidea) with morphological support. Invertebrate Systematics. 34 (2):113.
Abstract
The axiidean families Callianassidae and Ctenochelidae (sensu Dworschak et al. 2012), whether or not combined as Callianassoidea (sensu Sakai 2005), are shown to represent a monophyletic taxon. It comprises 265 accepted species in 74 genera, twice this number of species if fossil taxa are included. The higher taxonomy of the group has proved difficult and fluid. In a molecular phylogenetic approach, we inferred evolutionary relationships from an ML and Bayesian analysis of four genes, mitochondrial 16S and 12S along with nuclear Histone 3 and 18S. Our sample consisted of 298 specimens representing 123 species plus two species each of Axiidae and Callianideidae serving as outgroups. This number represented about half of all known species, but included 26 species undescribed or not confidently identified, 9% of all known. In a parallel morphological approach, the published descriptions of all species were examined and detailed observations made on about two-thirds of the known fauna in museum collections. A DELTA (DEscription Language for TAxonomy) database of 135 characters was made for 195 putative species, 18 of which were undescribed. A PAUP analysis found small clades coincident with the terminal clades found in the molecular treatment. Bayesian analysis of a total-evidence data set combined elements of both molecular and morphological analyses. Clades were interpreted as seven families and 53 genera. Seventeen new genera are required to reflect the molecular and morphological phylograms. Relationships between the families and genera inferred from the two analyses differed between the two strategies in spite of retrospective searches for morphological features supporting intermediate clades. The family Ctenochelidae was recovered in both analyses but the monophyly of Paragourretia was not supported by molecular data. The hitherto well recognised family Eucalliacidae was found to be polyphyletic in the molecular analysis, but the family and its genera were well defined by morphological synapomorphies. The phylogram for Callianassidae suggested the isolation of several species from the genera to which they had traditionally been assigned and necessitated 12 new generic names. The same was true for Callichiridae, with stronger ML than Bayesian support, and five new genera are proposed. Morphological data did not reliably reflect generic relationships inferred from the molecular analysis though they did diagnose terminal taxa treated as genera. We conclude that discrepancies between molecular and morphological analyses are due at least in part to missing sequences for key species, but no less to our inability to recognise unambiguously informative morphological synapomorphies. The ML analysis revealed the presence of at least ten complexes wherein two to four cryptic species masquerade under single species names.Read the article »
Article DOI: 10.1071/IS19021
Project DOI: 10.7934/P3512, http://dx.doi.org/10.7934/P3512
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 11651 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 176 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 427 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:176/0 |
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MorphoBank Project 3512
MorphoBank Project 3512
- Creation Date:
09 August 2019 - Publication Date:
29 June 2020 - Project views: 15481
- Matrix downloads: 8
This research
supported by
Authors' Institutions
- Museums Victoria
- Natural History Museum, Vienna (Naturhistorisches Museum, Wien)
- Universidade de Sao Paulo (USP, University of Sao Paulo)
- University of Louisiana
- Universidad Autónoma de Campeche
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
GARY POORE Project Administrator | 1 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M26194' (75 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Pillsburyaxius clevai Last Modified in 08/12/19 | 1 | 38 | 38 | 0 | 6 | 0 | 0 | 1 |
[2] Eiconaxius agassizi Last Modified in 08/12/19 | 1 | 38 | 38 | 0 | 6 | 0 | 0 | 1 |
[3] Heardaxius rogerbamberi Last Modified in 08/12/19 | 1 | 43 | 43 | 0 | 1 | 0 | 0 | 1 |
[4] Thomassinia aimsae Last Modified in 08/12/19 | 1 | 43 | 43 | 0 | 1 | 0 | 0 | 1 |
[5] PRAEDATRYPAEA praedatrix Last Modified in 08/12/19 | 1 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[6] MOZ-34 Last Modified in 08/12/19 | 1 | 43 | 43 | 0 | 1 | 0 | 0 | 1 |
[7] Propinquus Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[8] Intermedia Last Modified in 06/26/20 | 9 | 35 | 35 | 0 | 1 | 0 | 0 | 1 |
[9] PESTARELLA tyrrhena Last Modified in 06/26/20 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[10] GILVOSSIUS setimanus Last Modified in 06/26/20 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[11] Candidus Last Modified in 06/26/20 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[12] Convexus Last Modified in 06/26/20 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[13] Whitei Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[14] JOCULLIANASSA joculatrix Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[15] Ceramica Last Modified in 06/26/20 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[16] FILHOLLIANASSA filholi Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[17] NOTIAX brachyophthalma Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[18] Santarita Last Modified in 08/12/19 | 1 | 42 | 42 | 0 | 2 | 0 | 0 | 1 |
[19] TRYPAEA australiensis Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[20] ARENALLIANASSA arenosa Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[21] PARATRYPAEA bouvieri Last Modified in 06/26/20 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[22] Maldivensis Last Modified in 08/12/19 | 0 | 44 | 43 | 0 | 2 | 0 | 0 | 1 |
[23] TASTRYPAEA poorei Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[24] CHERAMUS profundus Last Modified in 08/12/19 | 4 | 40 | 40 | 0 | 1 | 0 | 0 | 1 |
[25] CHERAMOIDES marginata Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[26] Oblonga Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[27] POTI gaucho Last Modified in 08/12/19 | 8 | 35 | 35 | 0 | 2 | 0 | 0 | 1 |
[28] LIPKECALLIANASSA abyssa Last Modified in 08/12/19 | 17 | 25 | 25 | 0 | 3 | 0 | 0 | 1 |
[29] PUGNATRYPAEA pugnatrix Last Modified in 08/12/19 | 4 | 40 | 40 | 0 | 1 | 0 | 0 | 1 |
[30] GMX Last Modified in 08/12/19 | 1 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[31] Lobetobensis Last Modified in 08/12/19 | 7 | 37 | 37 | 0 | 1 | 0 | 0 | 1 |
[32] Sibogae Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[33] MOZ-31 Last Modified in 08/12/19 | 2 | 41 | 41 | 0 | 2 | 0 | 0 | 1 |
[34] MOZ-33 Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[35] CORIOLLIANASSA coriolisae Last Modified in 08/12/19 | 2 | 41 | 41 | 0 | 2 | 0 | 0 | 1 |
[36] BIFFARIUS biformis Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[37] Limosus Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[38] Delicatulus Last Modified in 06/26/20 | 5 | 39 | 39 | 0 | 1 | 0 | 0 | 1 |
[39] FRAGILLIANASSA fragilis Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[40] Debilis Last Modified in 06/26/20 | 1 | 42 | 42 | 0 | 2 | 0 | 0 | 1 |
[41] NECALLIANASSA berylae Last Modified in 08/12/19 | 1 | 42 | 42 | 0 | 2 | 0 | 0 | 1 |
[42] Acanthura Last Modified in 06/26/20 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[43] Truncata Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[44] CALLIANASSA subterranea Last Modified in 06/26/20 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[45] Diaphora Last Modified in 06/26/20 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[46] Australis Last Modified in 06/26/20 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[47] Marchali Last Modified in 08/12/19 | 0 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[48] NEOTRYPAEA californiensis Last Modified in 06/26/20 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[49] NIHONOTRYPAEA japonica Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[50] PSEUDOBIFFARIUS caesari Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[51] Harmandi Last Modified in 06/26/20 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[52] Petalura Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[53] Makarovi Last Modified in 08/12/19 | 1 | 43 | 43 | 0 | 1 | 0 | 0 | 1 |
[54] Gigas Last Modified in 06/26/20 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[55] Tabogensis Last Modified in 08/12/19 | 1 | 43 | 43 | 0 | 1 | 0 | 0 | 1 |
[56] Uncinata Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[57] Thermophila Last Modified in 08/12/19 | 1 | 43 | 43 | 0 | 1 | 0 | 0 | 1 |
[58] SPINICALLIANASSA spinicauda Last Modified in 08/12/19 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[59] Ohurana Last Modified in 08/12/19 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[60] Acutirostella Last Modified in 08/12/19 | 5 | 39 | 39 | 0 | 1 | 0 | 0 | 1 |
[61] PNG-757 Last Modified in 08/12/19 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[62] FP-10 Last Modified in 08/12/19 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[63] Aff. acutirostella Last Modified in 08/12/19 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[64] AQABALLIANASSA aqabaensis Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[65] Brevirostris Last Modified in 06/26/20 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[66] Lewtonae Last Modified in 08/12/19 | 0 | 45 | 45 | 0 | 0 | 0 | 0 | 1 |
[67] PNG-1166 Last Modified in 08/12/19 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[68] FP-11 Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[69] PNG-1172 Last Modified in 08/12/19 | 2 | 41 | 41 | 0 | 2 | 0 | 0 | 1 |
[70] CAVIALLIANASSA cavifrons Last Modified in 08/12/19 | 6 | 37 | 37 | 0 | 2 | 0 | 0 | 1 |
[71] RAYLLIANASSA amboinensis Last Modified in 08/12/19 | 0 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[72] SCALLASIS amboinae Last Modified in 08/12/19 | 0 | 45 | 45 | 0 | 0 | 0 | 0 | 1 |
[73] Spinophthalmus Last Modified in 08/12/19 | 2 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[74] RUDISULLIANASSA rudisulcus Last Modified in 08/12/19 | 4 | 39 | 39 | 0 | 2 | 0 | 0 | 1 |
[75] PNG-369 Last Modified in 08/12/19 | 2 | 41 | 41 | 0 | 2 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M26195' (72 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Pillsburyaxius clevai Last Modified in 08/12/19 | 2 | 43 | 43 | 0 | 9 | 0 | 0 | 1 |
[2] Eiconaxius agassizi Last Modified in 08/12/19 | 2 | 43 | 43 | 0 | 9 | 0 | 0 | 1 |
[3] Heardaxius rogerbamberi Last Modified in 08/12/19 | 3 | 49 | 49 | 0 | 2 | 0 | 0 | 1 |
[4] Thomassinia aimsae Last Modified in 08/12/19 | 2 | 48 | 48 | 0 | 4 | 0 | 0 | 1 |
[5] MOCALLICHIRUS mocambiquensis Last Modified in 08/12/19 | 0 | 50 | 50 | 0 | 4 | 0 | 0 | 1 |
[6] AUDACALLICHIRUS audax Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[7] Mirim Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[8] Monodi Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[9] CALLICHIRUS major Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[10] Aff. major Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[11] Islagrande Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[12] Adamas Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[13] Seilacheri Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[14] KRAUSSILLICHIRUS kraussi Last Modified in 08/12/19 | 0 | 53 | 53 | 0 | 1 | 0 | 0 | 1 |
[15] LATICALLICHIRUS grandis Last Modified in 08/12/19 | 0 | 53 | 53 | 0 | 1 | 0 | 0 | 1 |
[16] GRYNAMINNA tamakii Last Modified in 08/12/19 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[17] CALLIAPAGUROPS charcoti Last Modified in 08/12/19 | 5 | 43 | 43 | 0 | 6 | 0 | 0 | 1 |
[18] ForestiC Last Modified in 06/26/20 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[19] GLYPTURUS acanthochirus Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[20] Laurae Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[21] Armatus Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[22] Rabalaisae Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[23] CORALLICHIRUS xutha Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[24] CORALLIANASSA longiventris Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[25] Martensi Last Modified in 08/12/19 | 1 | 51 | 51 | 0 | 2 | 0 | 0 | 1 |
[26] Coutierei Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[27] Intesi Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[28] Haswelli Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[29] Hartmeyeri Last Modified in 06/26/20 | 8 | 41 | 41 | 0 | 5 | 0 | 0 | 1 |
[30] Borradailei Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[31] MUCROLLICHIRUS mucronatus Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[32] BALSSCALLICHIRUS balssi Last Modified in 08/12/19 | 4 | 45 | 45 | 0 | 5 | 0 | 0 | 1 |
[33] TIRMIZICALLICHIRUS masoomi Last Modified in 08/12/19 | 1 | 51 | 51 | 0 | 2 | 0 | 0 | 1 |
[34] CAPECALLIAX pixii Last Modified in 08/12/19 | 3 | 49 | 49 | 0 | 2 | 0 | 0 | 1 |
[35] Guineensis Last Modified in 06/26/20 | 3 | 47 | 47 | 0 | 4 | 0 | 0 | 1 |
[36] FORESTCALLICHIRUS foresti Last Modified in 08/12/19 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[37] BARNARDCALLICHIRUS tenuimanus Last Modified in 08/12/19 | 2 | 49 | 49 | 0 | 3 | 0 | 0 | 1 |
[38] NEOCALLICHIRUS horneri Last Modified in 06/26/20 | 2 | 50 | 50 | 0 | 2 | 0 | 0 | 1 |
[39] Darwinensis Last Modified in 06/26/20 | 8 | 42 | 42 | 0 | 4 | 0 | 0 | 1 |
[40] Pachydactylus Last Modified in 08/12/19 | 1 | 51 | 51 | 0 | 2 | 0 | 0 | 1 |
[41] Vaugelasi Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[42] Lemaitrei Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[43] Jousseaumei Last Modified in 06/26/20 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[44] Variabilis Last Modified in 08/12/19 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[45] Nickellae Last Modified in 08/12/19 | 5 | 46 | 46 | 0 | 3 | 0 | 0 | 1 |
[46] Grandimana Last Modified in 06/26/20 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[47] Maryae Last Modified in 08/12/19 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[48] Angelikae Last Modified in 06/26/20 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[49] CALLICHIROPSIS spiridonovi Last Modified in 08/12/19 | 8 | 40 | 40 | 0 | 6 | 0 | 0 | 1 |
[50] Cacahuate Last Modified in 06/26/20 | 1 | 50 | 50 | 0 | 3 | 0 | 0 | 1 |
[51] Natalensis Last Modified in 08/12/19 | 3 | 46 | 46 | 0 | 5 | 0 | 0 | 1 |
[52] Calmani Last Modified in 06/26/20 | 1 | 51 | 51 | 0 | 2 | 0 | 0 | 1 |
[53] PNG-1589 Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[54] SERGIO guassutinga Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[55] Mericeae Last Modified in 08/12/19 | 1 | 51 | 51 | 0 | 2 | 0 | 0 | 1 |
[56] Sulfureus Last Modified in 08/12/19 | 0 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[57] Vigilax Last Modified in 08/12/19 | 1 | 50 | 50 | 0 | 3 | 0 | 0 | 1 |
[58] MICHAELCALLIANASSA indica Last Modified in 08/12/19 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[59] Sinica Last Modified in 08/12/19 | 0 | 51 | 51 | 0 | 3 | 0 | 0 | 1 |
[60] GLYPTUROIDES trilobata Last Modified in 08/12/19 | 0 | 53 | 53 | 0 | 1 | 0 | 0 | 1 |
[61] KARUMBALLICHIRUS karumba Last Modified in 08/12/19 | 0 | 54 | 54 | 0 | 0 | 0 | 0 | 1 |
[62] LEPIDOPHTHALMUS eiseni Last Modified in 08/12/19 | 0 | 54 | 54 | 0 | 0 | 0 | 0 | 1 |
[63] Bocourti Last Modified in 06/26/20 | 0 | 53 | 53 | 0 | 1 | 0 | 0 | 1 |
[64] Jamaicensis Last Modified in 06/26/20 | 0 | 54 | 54 | 0 | 0 | 0 | 0 | 1 |
[65] Louisianensis Last Modified in 08/12/19 | 0 | 54 | 54 | 0 | 0 | 0 | 0 | 1 |
[66] Rosae Last Modified in 08/12/19 | 1 | 53 | 53 | 0 | 0 | 0 | 0 | 1 |
[67] Tridentatus Last Modified in 06/26/20 | 0 | 54 | 54 | 0 | 0 | 0 | 0 | 1 |
[68] Grandidieri Last Modified in 06/26/20 | 4 | 47 | 47 | 0 | 3 | 0 | 0 | 1 |
[69] Turneranus Last Modified in 08/12/19 | 0 | 54 | 54 | 0 | 0 | 0 | 0 | 1 |
[70] PODOCALLICHIRUS madagassa Last Modified in 08/12/19 | 0 | 53 | 53 | 0 | 1 | 0 | 0 | 1 |
[71] THAILANDCALLICHIRUS ranongensis Last Modified in 08/12/19 | 0 | 53 | 53 | 0 | 1 | 0 | 0 | 1 |
[72] FAMILY callianassidae Last Modified in 08/12/19 | 4 | 75 | 49 | 0 | 1 | 0 | 0 | 1 |
Taxonomic Overview for Matrix 'M26196' (63 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Pillsburyaxius clevai Last Modified in 08/12/19 | 5 | 61 | 61 | 0 | 11 | 0 | 0 | 1 |
[2] Eiconaxius agassizi Last Modified in 08/12/19 | 5 | 61 | 61 | 0 | 11 | 0 | 0 | 1 |
[3] Heardaxius rogerbamberi Last Modified in 08/12/19 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[4] Thomassinia aimsae Last Modified in 08/12/19 | 5 | 66 | 66 | 0 | 6 | 0 | 0 | 1 |
[5] ANACALLIAX argentinensis Last Modified in 08/12/19 | 7 | 68 | 68 | 0 | 2 | 0 | 0 | 1 |
[6] ANACALLIAOPSIS agassizi Last Modified in 08/12/19 | 6 | 69 | 69 | 0 | 2 | 0 | 0 | 1 |
[7] BATHYCALLIAX geomar Last Modified in 08/12/19 | 7 | 67 | 67 | 0 | 3 | 0 | 0 | 1 |
[8] VULCANOCALLIAX arutyunovi Last Modified in 08/12/19 | 6 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[9] CALLIANOPSIS goniophthalma Last Modified in 08/12/19 | 7 | 69 | 69 | 0 | 1 | 0 | 0 | 1 |
[10] PLEUROCALLIAX caecigena Last Modified in 08/12/19 | 13 | 57 | 57 | 0 | 7 | 0 | 0 | 1 |
[11] PHAETONCALLIAX mauritana Last Modified in 08/12/19 | 6 | 70 | 70 | 0 | 1 | 0 | 0 | 1 |
[12] NEOCALLIANOPSIS anovalis Last Modified in 08/12/19 | 8 | 68 | 68 | 0 | 1 | 0 | 0 | 1 |
[13] PARACALLIAX bollorei Last Modified in 08/12/19 | 8 | 64 | 64 | 0 | 5 | 0 | 0 | 1 |
[14] CTENOCHELES balssi Last Modified in 08/12/19 | 3 | 74 | 73 | 0 | 1 | 0 | 0 | 1 |
[15] CTENOCHELOIDES attenboroughi Last Modified in 08/12/19 | 6 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[16] Boucheti Last Modified in 06/26/20 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[17] Almeidai Last Modified in 06/26/20 | 6 | 65 | 65 | 0 | 6 | 0 | 0 | 1 |
[18] KIICTENOCHELOIDES nomurai Last Modified in 08/12/19 | 6 | 70 | 70 | 0 | 1 | 0 | 0 | 1 |
[19] LAURENTGOURRETIA rhopalommata Last Modified in 08/12/19 | 13 | 61 | 61 | 0 | 3 | 0 | 0 | 1 |
[20] DAWSONIUS latispina Last Modified in 08/12/19 | 2 | 73 | 73 | 0 | 2 | 0 | 0 | 1 |
[21] Aff. latispina Last Modified in 08/12/19 | 2 | 73 | 73 | 0 | 2 | 0 | 0 | 1 |
[22] PARAGOURRETIA phuketensis Last Modified in 08/12/19 | 9 | 67 | 67 | 0 | 1 | 0 | 0 | 1 |
[23] Lahouensis Last Modified in 08/12/19 | 2 | 74 | 74 | 0 | 1 | 0 | 0 | 1 |
[24] Laevidactyla Last Modified in 06/26/20 | 5 | 67 | 67 | 0 | 5 | 0 | 0 | 1 |
[25] Coolibah Last Modified in 06/26/20 | 1 | 75 | 75 | 0 | 1 | 0 | 0 | 1 |
[26] TUERKAYGOURRETIA galathea Last Modified in 08/12/19 | 8 | 68 | 68 | 0 | 1 | 0 | 0 | 1 |
[27] HETEROGOURRETIA aungtonyae Last Modified in 08/12/19 | 3 | 71 | 71 | 0 | 3 | 0 | 0 | 1 |
[28] Biffari Last Modified in 06/26/20 | 2 | 72 | 72 | 0 | 3 | 0 | 0 | 1 |
[29] ULLZ-10134 Last Modified in 08/12/19 | 8 | 66 | 66 | 0 | 3 | 0 | 0 | 1 |
[30] PSEUDOGOURRETIA portsudanensis Last Modified in 08/12/19 | 23 | 52 | 52 | 0 | 2 | 0 | 0 | 1 |
[31] GOURRETIA denticulata Last Modified in 08/12/19 | 1 | 75 | 75 | 0 | 1 | 0 | 0 | 1 |
[32] IVORYGOURRETIA barracuda Last Modified in 08/12/19 | 1 | 75 | 75 | 0 | 1 | 0 | 0 | 1 |
[33] PLANTESGOURRETIA nosybeensis Last Modified in 08/12/19 | 2 | 73 | 73 | 0 | 2 | 0 | 0 | 1 |
[34] RUIYULIUGOURRETIA sinica Last Modified in 08/12/19 | 1 | 74 | 74 | 0 | 2 | 0 | 0 | 1 |
[35] Zarenkovi Last Modified in 08/12/19 | 13 | 61 | 61 | 0 | 3 | 0 | 0 | 1 |
[36] Laresi Last Modified in 08/12/19 | 2 | 74 | 74 | 0 | 1 | 0 | 0 | 1 |
[37] Manihinae Last Modified in 08/12/19 | 8 | 64 | 64 | 0 | 5 | 0 | 0 | 1 |
[38] ULLZ-9707 Last Modified in 08/12/19 | 18 | 51 | 51 | 0 | 8 | 0 | 0 | 1 |
[39] ANDAMANCALLIAX andamanica Last Modified in 08/12/19 | 12 | 65 | 64 | 0 | 1 | 0 | 0 | 1 |
[40] PSEUDOCALLIAX tooradin Last Modified in 08/12/19 | 5 | 69 | 69 | 0 | 3 | 0 | 0 | 1 |
[41] EUCALLIAX quadracuta Last Modified in 08/12/19 | 5 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[42] PARAGLYPTURUS calderus Last Modified in 08/12/19 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[43] Tonganus Last Modified in 08/12/19 | 6 | 70 | 70 | 0 | 1 | 0 | 0 | 1 |
[44] CALLIAXINA punica Last Modified in 08/12/19 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[45] Sakaii Last Modified in 08/12/19 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[46] Novaebritanniae Last Modified in 08/12/19 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[47] Kensleyi Last Modified in 06/26/20 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[48] Bulimba Last Modified in 08/12/19 | 5 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[49] Xishaensis Last Modified in 08/12/19 | 10 | 61 | 61 | 0 | 6 | 0 | 0 | 1 |
[50] EUCALLIAXIOPSIS cearaensis Last Modified in 08/12/19 | 8 | 67 | 67 | 0 | 2 | 0 | 0 | 1 |
[51] Panglaoensis Last Modified in 08/12/19 | 5 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[52] Aff. panglaoensis Last Modified in 08/12/19 | 5 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[53] BAKERCALLIAX aequimana Last Modified in 08/12/19 | 5 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[54] Inaequimana Last Modified in 08/12/19 | 5 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[55] PNG-1153C Last Modified in 08/12/19 | 5 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[56] CALLIAXIOPSIS madagassa Last Modified in 08/12/19 | 8 | 68 | 68 | 0 | 1 | 0 | 0 | 1 |
[57] HEARDCALLIAX jonesi Last Modified in 08/12/19 | 6 | 69 | 69 | 0 | 2 | 0 | 0 | 1 |
[58] MANNINGCALLIAX mcilhennyi Last Modified in 08/12/19 | 5 | 70 | 70 | 0 | 2 | 0 | 0 | 1 |
[59] CALLIAX lobata Last Modified in 08/12/19 | 4 | 70 | 70 | 0 | 3 | 0 | 0 | 1 |
[60] Doerjesti Last Modified in 08/12/19 | 4 | 70 | 70 | 0 | 3 | 0 | 0 | 1 |
[61] Tulearensis Last Modified in 08/12/19 | 8 | 66 | 66 | 0 | 3 | 0 | 0 | 1 |
[62] FAMILY callichiridae Last Modified in 08/12/19 | 4 | 121 | 72 | 0 | 1 | 0 | 0 | 1 |
[63] FAMILY callianassidae Last Modified in 08/12/19 | 6 | 92 | 67 | 0 | 4 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 15481 | |
Project overview | 1081 | |
Matrix views | 875 | CALLIC.nex (103 views); BASAL.nex (112 views); CALASS.nex (101 views); Matrix landing page (559 views); |
Media views | 2287 | Media search (1974 views); M692396 (313 views); |
Documents list | 581 | |
Views for media list | 729 | |
Specimen list | 1193 | |
Taxon list | 8251 | |
Bibliography | 484 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 119 | |
Project downloads | 110 | |
Document downloads | 1 | Full character labels (1 download); |
Matrix downloads | 8 | BASAL.nex (3 downloads); CALASS.nex (4 downloads); CALLIC.nex (1 download); |