Project 352: E. V. Bärmann. Towards a comprehensive phylogeny of Bovidae, Chapter 2: A total evidence analysis of bovid phylogeny - Dissertation by Eva Bärmann, University of Cambridge UK.
Specimen: Raphicerus melanotis (ZMB/Mammals:2125)

Abstract

Combined-data analyses, also termed total evidence analyses, have proven to be beneficial for phylogeny reconstructions for a range of taxa. For Bovidae, the most diverse clade of ruminants (Artiodactyla, Mammalia), such an analysis was last performed a decade ago. Since then, the range of possible analysis techniques has expanded, now including model-based Bayesian inference that can also be used to incorporate non-molecular data. Furthermore, the possibility of making morphological data widely accessible online, providing detailed information on character states via photographs, facilitates the use and re-use of this type of data. This study presents a data set of 88 cranial, 19 soft-tissue, and 27 behavioural characters, sampled from 58 bovid species. These data are analysed in a simultaneous analysis with molecular data from nine genes (four mitochondrial and five nuclear markers), sampled for 57 bovid species. A few morphological and behavioural characters are identified as apomorphies of bovid taxa. Despite the fact that the majority of characters show homoplasies and cannot unambiguously be optimized on the phylogenetic tree, they allow for inferring the phylogenetic position of two fossil bovid species: †Turcoceros grangeri and †Gazella lydekkeri. The study is designed as a starting point for further combined data analyses of bovids or ruminants in general.


Read the article »

Project DOI: 10.7934/P352, http://dx.doi.org/10.7934/P352
This project containsMatrices
  • 806 Media
  • 3 Matrices
  • 86 Taxa
  • 212 Specimens
  • 134 Characters
Total size of project's media files: 1.13G

Download Project SDD File
Total scored cells: 8111
Total media associated with cells: 6
Total labels associated with cell media: 0
Characters
Total characters: 134
Total characters with associated media: 10
Total characters with media with labels: 0
Total character states: 332
Total character states with associated media: 170
Total character states with media with labels:122
Total unordered/ordered characters:134/0
Currently Viewing:
MorphoBank Project 352

    Authors' Institutions

    • University of Cambridge



    Members

    member name taxa specimens media chars character
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    notes  medialabels
    rules
    Eva Baermann
    Project Administrator
    86212806134112801588111
    (7554, 429, 128)
    434600
    Robert Asher
    Full membership
    00000000
    (0, 0, 0)
    0000


    Taxonomic Overview for Matrix 'M817' (61 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    0878620002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    0898500002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    0847931002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    0878511002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    0878710002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    0888700002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0838032002
    [8] Capra falconeri
    Last Modified in 12/05/13
    1848231002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0868111002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    0847731002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    0817752002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    0878601002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    0878501002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    0878311002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    0878401002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    0878601002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1828151002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    0878501002
    [19] Gazella gazella
    Last Modified in 12/05/13
    1838131002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0878401002
    [21] Gazella marica
    Last Modified in 12/05/13
    0808071002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0848131002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    0878301002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    0868411002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    0898501002
    [26] Kobus kob
    Last Modified in 12/05/13
    0827761002
    [27] Kobus leche
    Last Modified in 12/05/13
    0837741002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    0797881002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    0838241002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    0888500002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    0858521002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    0868511002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    0868611002
    [34] Nanger dama
    Last Modified in 12/05/13
    0878301002
    [35] Nanger granti
    Last Modified in 12/05/13
    0878201002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    0888301002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0858412002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    0878701002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    0878302002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    0898601002
    [41] Oryx beisa
    Last Modified in 12/05/13
    1868221002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    0878511202
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0848331002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    0878601002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    0908101002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0878301002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    0837841002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    0818061002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    0878402002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0868602002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0828152002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    0868221002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    0878321202
    [54] Redunca redunca
    Last Modified in 12/05/13
    0817861002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    0888112002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0878301002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    0898301202
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0847432002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    0878701002
    [60] † Gazella lydekkeri
    Last Modified in 12/05/13
    0680191002
    [61] † Turcoceros grangeri
    Last Modified in 12/05/13
    1691180002


    Taxonomic Overview for Matrix 'M848' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    022523002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    027100002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    026401002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    025020002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    027000002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    026010002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    026711002
    [8] Capra falconeri
    Last Modified in 12/05/13
    022023002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0221241002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    024921002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    029401002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    024221002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    017082002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018090002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    027000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    081190002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1161100002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    023140002
    [19] Gazella gazella
    Last Modified in 12/05/13
    027100002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0161110002
    [21] Gazella marica
    Last Modified in 12/05/13
    025720002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0131140002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    025120002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    026101002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    023004002
    [26] Kobus kob
    Last Modified in 12/05/13
    027402002
    [27] Kobus leche
    Last Modified in 12/05/13
    026304002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    018564002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    617504002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    025020002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    025020002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    027100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    026110002
    [34] Nanger dama
    Last Modified in 12/05/13
    019080002
    [35] Nanger granti
    Last Modified in 12/05/13
    026010002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    019080002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0100161002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    025220002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    080190002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    025211002
    [41] Oryx beisa
    Last Modified in 12/05/13
    026603002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    025120002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0140112002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    024803002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    026701002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0100170002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    020561002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    018581002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    025020002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0120150002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0100170002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    024203002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    024003002
    [54] Redunca redunca
    Last Modified in 12/05/13
    024303002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019153002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0152111002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    026101002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0143121002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    022123002


    Taxonomic Overview for Matrix 'M849' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    019800002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    019000002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    019900002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    018510002
    [5] Antilope cervicapra
    Last Modified in 12/05/13
    017520002
    [6] Antidorcas marsupialis
    Last Modified in 12/05/13
    019000002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0191200002
    [8] Capra falconeri
    Last Modified in 12/05/13
    018010002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    118600002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    019600002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    019700002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    019000002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    019000002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018610002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    019000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    017520002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    018510002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    018510002
    [19] Gazella gazella
    Last Modified in 12/05/13
    019600002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    017520002
    [21] Gazella marica
    Last Modified in 12/05/13
    019800002
    [22] Gazella spekei
    Last Modified in 12/05/13
    018510002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    019600002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    019000002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    019000002
    [26] Kobus kob
    Last Modified in 12/05/13
    019500002
    [27] Kobus leche
    Last Modified in 12/05/13
    019500002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    019500002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    019600002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    019000002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    019000002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    019100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    019000002
    [34] Nanger dama
    Last Modified in 12/05/13
    018510002
    [35] Nanger granti
    Last Modified in 12/05/13
    019000002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    018510002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    018210002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    018010002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    018410002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    019000002
    [41] Oryx beisa
    Last Modified in 12/05/13
    019900002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    019000002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    019000002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    019400002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    019600002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    019400002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    019300002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    019400002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    019000002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    018010002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    019000002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    019500002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    019000002
    [54] Redunca redunca
    Last Modified in 12/05/13
    019500002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019700002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    018010002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    019000002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0191300002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    019000002


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M107807 (154 views); M107817 (230 views); M107818 (190 views); M107821 (158 views); M142302 (130 views); M142304 (166 views); M142305 (185 views); M142307 (250 views); M142308 (138 views); M142309 (198 views); M148781 (189 views); M149147 (34 views); M148170 (124 views); M148782 (112 views); M148795 (124 views); M284576 (7 views); M149111 (38 views); M148171 (120 views); M46500 (292 views); M46842 (254 views); M148784 (120 views); M46495 (218 views); M46825 (262 views); M148785 (114 views); M148780 (121 views); M149225 (34 views); M98198 (4 views); M148799 (122 views); M148794 (120 views); M148152 (159 views); M148778 (164 views); M46343 (484 views); M46187 (536 views); M46167 (555 views); M46172 (599 views); M148173 (167 views); M46179 (580 views); M148820 (34 views); M148175 (142 views); M148788 (117 views); M149219 (33 views); M148814 (40 views); M149230 (22 views); M149193 (64 views); M148800 (116 views); M46335 (577 views); M148818 (29 views); M46148 (552 views); M46155 (590 views); M149170 (36 views); M107804 (141 views); M148169 (116 views); M98201 (7 views); M149208 (54 views); M148790 (161 views); M149145 (34 views); M148166 (115 views); M149141 (110 views); M149292 (41 views); M149259 (6 views); M148813 (35 views); M149150 (34 views); M148826 (31 views); M149122 (34 views); M148823 (48 views); M148798 (221 views); M149211 (37 views); M148827 (32 views); M149174 (27 views); M149160 (27 views); M148824 (31 views); M149135 (47 views); M148779 (150 views); M47883 (5 views); M149149 (29 views); M148812 (27 views); M149168 (32 views); M149192 (40 views); M149137 (26 views); M149189 (68 views); M149180 (60 views); M46348 (544 views); M149213 (30 views); M148803 (28 views); M149275 (5 views); M46852 (261 views); M47020 (5 views); M149253 (6 views); M149118 (34 views); M148805 (30 views); M149194 (33 views); M149143 (38 views); M148811 (52 views); M149223 (25 views); M149222 (33 views); M149159 (29 views); M149172 (29 views); M148801 (30 views); M148802 (31 views); M148804 (30 views); M148806 (27 views); M148807 (50 views); M148808 (72 views); M148809 (34 views); M148810 (26 views); M148815 (36 views); M148816 (54 views); M148817 (37 views); M148819 (26 views); M148821 (28 views); M148825 (36 views); M149110 (49 views); M149112 (26 views); M149113 (28 views); M149114 (36 views); M149115 (32 views); M149116 (29 views); M149117 (38 views); M149119 (37 views); M149123 (38 views); M149134 (71 views); M149136 (28 views); M149138 (27 views); M149139 (29 views); M149140 (34 views); M149142 (40 views); M149144 (33 views); M149146 (28 views); M149148 (26 views); M149151 (28 views); M149152 (38 views); M149153 (46 views); M149154 (36 views); M149155 (27 views); M149156 (28 views); M149157 (29 views); M149158 (39 views); M149161 (36 views); M149162 (26 views); M149163 (34 views); M149164 (30 views); M149165 (35 views); M149166 (34 views); M149167 (28 views); M149169 (26 views); M149173 (33 views); M149175 (35 views); M149176 (66 views); M149177 (68 views); M149179 (48 views); M149181 (65 views); M149182 (52 views); M149183 (38 views); M149184 (55 views); M149185 (50 views); M149187 (62 views); M149188 (75 views); M149190 (42 views); M149191 (99 views); M149196 (35 views); M149197 (47 views); M149198 (30 views); M149209 (34 views); M149210 (35 views); M149212 (29 views); M149214 (29 views); M149218 (30 views); M149220 (30 views); M149221 (23 views); M149224 (24 views); M149226 (26 views); M149227 (23 views); M149228 (29 views); M149229 (23 views); M149242 (8 views); M46411 (261 views); M46193 (553 views); M46333 (499 views); M46449 (232 views); M286147 (5 views); M46405 (253 views); M149233 (5 views); M48516 (6 views); M46824 (263 views); M47787 (4 views); M46919 (5 views); M46431 (235 views); M46905 (6 views); M47950 (5 views); M47788 (8 views); M47955 (6 views); M46188 (533 views); M46202 (494 views); M149289 (7 views); M47887 (4 views); M149282 (4 views); M46326 (475 views); M46421 (230 views); M46415 (274 views); M46453 (224 views); M46153 (569 views); M46904 (5 views); M46843 (276 views); M149286 (5 views); M151747 (4 views); M46851 (260 views); M46118 (522 views); M46127 (551 views); M46128 (535 views); M46134 (551 views); M46135 (550 views); M46144 (552 views); M46145 (562 views); M46146 (580 views); M46147 (558 views); M46151 (532 views); M46152 (538 views); M46154 (552 views); M46166 (554 views); M46169 (509 views); M46171 (589 views); M46176 (556 views); M46177 (539 views); M46178 (566 views); M46180 (534 views); M46184 (517 views); M46185 (530 views); M46186 (528 views); M46189 (533 views); M46190 (567 views); M46191 (535 views); M46192 (627 views); M46194 (483 views); M46195 (539 views); M46197 (514 views); M46198 (489 views); M46199 (519 views); M46200 (517 views); M46204 (489 views); M46205 (456 views); M46206 (489 views); M46207 (573 views); M46208 (473 views); M46210 (490 views); M46211 (487 views); M46212 (580 views); M46230 (608 views); M46327 (502 views); M46328 (566 views); M46329 (507 views); M46330 (491 views); M46331 (512 views); M46334 (540 views); M46337 (507 views); M46340 (514 views); M46341 (498 views); M46342 (502 views); M46344 (514 views); M46345 (487 views); M46350 (487 views); M46351 (457 views); M46352 (514 views); M46353 (493 views); M46354 (510 views); M46355 (522 views); M46356 (473 views); M46357 (514 views); M46358 (504 views); M46359 (512 views); M46361 (491 views); M46362 (498 views); M46403 (496 views); M46406 (276 views); M46407 (285 views); M46412 (264 views); M46413 (229 views); M46414 (232 views); M46416 (248 views); M46417 (220 views); M46418 (272 views); M46419 (272 views); M46420 (234 views); M46422 (221 views); M46423 (220 views); M46424 (265 views); M46425 (275 views); M46426 (207 views); M46427 (252 views); M46428 (250 views); M46429 (226 views); M46430 (250 views); M46432 (226 views); M46434 (257 views); M46436 (230 views); M46437 (236 views); M46438 (252 views); M46440 (221 views); M46441 (237 views); M46442 (265 views); M46443 (266 views); M46444 (277 views); M46446 (230 views); M46447 (204 views); M46448 (232 views); M46450 (220 views); M46451 (245 views); M46452 (269 views); M46454 (276 views); M46455 (247 views); M46456 (221 views); M46457 (214 views); M46458 (256 views); M46459 (230 views); M46471 (255 views); M46472 (240 views); M46475 (221 views); M46476 (278 views); M46479 (267 views); M46480 (248 views); M46481 (283 views); M46482 (226 views); M46483 (238 views); M46484 (245 views); M46485 (240 views); M46486 (218 views); M46487 (254 views); M46488 (239 views); M46489 (233 views); M46490 (226 views); M46492 (247 views); M46493 (232 views); M46494 (257 views); M46496 (221 views); M46497 (230 views); M46498 (252 views); M46499 (260 views); M46809 (215 views); M46810 (222 views); M46812 (211 views); M46814 (240 views); M46815 (226 views); M46817 (227 views); M46818 (255 views); M46819 (258 views); M46820 (246 views); M46822 (250 views); M46823 (263 views); M46827 (222 views); M46828 (248 views); M46829 (215 views); M46830 (233 views); M46831 (213 views); M46832 (233 views); M46833 (217 views); M46834 (236 views); M46835 (278 views); M46837 (249 views); M46838 (239 views); M46839 (203 views); M46840 (232 views); M46841 (225 views); M46844 (251 views); M46845 (228 views); M46846 (217 views); M46847 (238 views); M46848 (210 views); M46849 (233 views); M46850 (246 views); M46853 (251 views); M46854 (206 views); M46855 (229 views); M46856 (214 views); M46857 (230 views); M46858 (219 views); M46859 (228 views); M46860 (277 views); M46861 (253 views); M46862 (254 views); M46863 (214 views); M46864 (217 views); M284578 (9 views); M149294 (5 views); M149243 (4 views); M149268 (4 views); M149271 (6 views); M46946 (5 views); M47791 (8 views); M98218 (4 views); M149285 (4 views); M284582 (5 views); M46950 (5 views); M98192 (7 views); M284577 (8 views); M46986 (7 views); M286153 (6 views); M149231 (7 views); M286145 (4 views); M286141 (4 views); M149248 (6 views); M149291 (5 views); M47018 (7 views); M48485 (7 views); M47022 (5 views); M46995 (6 views); M47794 (4 views); M47030 (4 views); M47009 (4 views); M47948 (4 views); M47786 (4 views); M46998 (4 views); M47007 (4 views); M47779 (4 views); M47954 (4 views); M47785 (4 views); M47965 (7 views); M47899 (8 views); M47986 (4 views); M47778 (4 views); M47790 (4 views); M48056 (6 views); M47976 (4 views); M48488 (5 views); M48058 (4 views); M48029 (4 views); M47970 (5 views); M46988 (4 views); M98208 (5 views); M47892 (4 views); M47781 (5 views); M47953 (7 views); M48509 (4 views); M98204 (4 views); M47011 (4 views); M47915 (25 views); M47014 (4 views); M47010 (5 views); M47797 (5 views); M48512 (5 views); M47016 (4 views); M47905 (5 views); M47906 (4 views); M47886 (5 views); M98212 (5 views); M98213 (5 views); M47795 (4 views); M98196 (4 views); M47951 (4 views); M47978 (6 views); M48047 (45 views); M48048 (4 views); M48487 (8 views); M48059 (4 views); M98195 (6 views); M47003 (5 views); M48503 (5 views); M47964 (5 views); M48514 (4 views); M46921 (6 views); M51943 (7 views); M47917 (5 views); M286140 (4 views); M48062 (5 views); M98209 (5 views); M149260 (8 views); M149297 (7 views); M46957 (10 views); M47017 (8 views); M149283 (4 views); M149281 (3 views); M149264 (4 views); M149288 (5 views); M46966 (7 views); M149279 (4 views); M47796 (5 views); M47789 (4 views); M98215 (4 views); M47884 (5 views); M149290 (4 views); M48053 (8 views); M48511 (4 views); M46923 (6 views); M47903 (4 views); M46939 (6 views); M47890 (4 views); M46926 (7 views); M47977 (6 views); M46952 (8 views); M47031 (5 views); M46924 (6 views); M46960 (7 views); M47775 (4 views); M48508 (4 views); M48499 (5 views); M46969 (7 views); M46929 (6 views); M46976 (20 views); M46964 (6 views); M98203 (4 views); M46967 (7 views); M46937 (7 views); M46972 (8 views); M47782 (4 views); M46971 (6 views); M46981 (7 views); M98217 (4 views); M46982 (7 views); M46955 (6 views); M98214 (4 views); M46959 (9 views); M47005 (4 views); M47858 (4 views); M48507 (4 views); M46951 (7 views); M46910 (6 views); M48497 (7 views); M46961 (6 views); M47981 (5 views); M46956 (8 views); M46979 (6 views); M47885 (4 views); M46968 (8 views); M46965 (8 views); M48057 (5 views); M46940 (6 views); M46934 (7 views); M46945 (6 views); M46980 (11 views); M46973 (6 views); M46978 (6 views); M46928 (8 views); M46970 (6 views); M46920 (7 views); M46975 (9 views); M48502 (4 views); M46953 (6 views); M98197 (5 views); M46962 (6 views); M46984 (6 views); M98210 (5 views); M46911 (9 views); M46925 (6 views); M48049 (5 views); M46917 (7 views); M47975 (5 views); M46932 (6 views); M46909 (6 views); M48515 (5 views); M47877 (5 views); M98194 (4 views); M48500 (4 views); M47949 (6 views); M149246 (5 views); M98207 (5 views); M47006 (4 views); M149235 (4 views); M47904 (4 views); M47889 (4 views); M98219 (4 views); M286142 (6 views); M46913 (6 views); M48505 (5 views); M47888 (4 views); M47776 (4 views); M46915 (5 views); M47974 (4 views); M149295 (4 views); M47780 (4 views); M284579 (6 views); M149269 (4 views); M149239 (6 views); M47792 (4 views); M47777 (4 views); M47021 (4 views); M46914 (5 views); M47019 (4 views); M47004 (4 views); M47012 (4 views); M47916 (6 views); M47871 (4 views); M98211 (5 views); M48061 (4 views); M98202 (5 views); M48510 (7 views); M48506 (5 views); M48055 (4 views); M151745 (4 views); M149267 (4 views); M149252 (4 views); M149255 (5 views); M149298 (4 views); M46943 (5 views); M286149 (4 views); M286154 (6 views); M46927 (5 views); M149238 (4 views); M149266 (4 views); M149244 (5 views); M149280 (4 views); M286143 (4 views); M286139 (4 views); M286146 (4 views); M286152 (7 views); M149250 (4 views); M149274 (4 views); M149299 (4 views); M149277 (4 views); M286155 (6 views); M286156 (6 views); M149287 (5 views); M149241 (4 views); M151743 (7 views); M149232 (4 views); M286150 (4 views); M46941 (6 views); M47882 (4 views); M47963 (4 views); M46907 (5 views); M46963 (7 views); M46912 (6 views); M48486 (5 views); M48050 (6 views); M47952 (3 views); M46935 (6 views); M46918 (4 views); M48054 (3 views); M48498 (3 views); M46938 (4 views); M48051 (3 views); M284581 (5 views); M149237 (3 views); M149296 (3 views); M149293 (3 views); M284584 (4 views); M149234 (4 views); M284580 (5 views); M149258 (8 views); M149256 (3 views); M149263 (3 views); M149240 (3 views); M149257 (4 views); M149236 (5 views); M46991 (3 views); M46989 (3 views); M47783 (3 views); M46922 (7 views); M47784 (3 views); M149278 (3 views); M46865 (6 views); M47034 (3 views); M46993 (3 views); M47774 (3 views); M46983 (5 views); M47874 (3 views); M46908 (6 views); M46996 (3 views); M286148 (3 views); M47896 (3 views); M149245 (3 views); M47914 (29 views); M286144 (3 views); M286151 (4 views); M48501 (3 views); M47956 (6 views); M149247 (3 views); M149254 (3 views); M149249 (3 views); M46997 (5 views); M151744 (3 views); M151746 (3 views); M149270 (3 views); M98193 (47 views); M46994 (3 views); M51944 (5 views); M47013 (4 views); M48052 (4 views); M149273 (3 views); M47895 (3 views); M47033 (3 views); M149265 (3 views); M46977 (8 views); M47015 (3 views); M46992 (4 views); M149261 (4 views); M47793 (3 views); M48513 (3 views); M149276 (3 views); M149284 (3 views); M98206 (5 views); M46958 (6 views); M47875 (4 views); M47897 (3 views); M149272 (3 views); M46936 (6 views); M46974 (5 views); M46947 (10 views); M47001 (3 views); M149251 (3 views); M47032 (4 views); M47876 (3 views); M47002 (3 views); M47029 (3 views); M47859 (3 views); M48060 (3 views); M47880 (3 views); M47973 (3 views); M46930 (6 views); M47891 (3 views); M48504 (3 views); M284583 (6 views); M98216 (4 views); M46933 (5 views);
    Bibliography14768
    Taxon list16977
    Specimen list111632




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project1262
    Project downloads359
    Matrix downloads78Skull characters Bovidae (59 downloads); Behavior (9 downloads); Glands and Colouration (10 downloads);
    Media downloads825M142313 (2 downloads); M142296 (1 download); M148795 (4 downloads); M148798 (3 downloads); M46337 (2 downloads); M46421 (2 downloads); M46492 (1 download); M47899 (2 downloads); M48485 (2 downloads); M48486 (2 downloads); M48487 (2 downloads); M48488 (2 downloads); M46418 (2 downloads); M46440 (2 downloads); M46820 (2 downloads); M149138 (2 downloads); M149143 (2 downloads); M107819 (2 downloads); M46187 (1 download); M46438 (1 download); M46907 (1 download); M46953 (1 download); M47789 (1 download); M47884 (1 download); M47978 (1 download); M149134 (1 download); M149135 (1 download); M149136 (1 download); M149137 (1 download); M149139 (2 downloads); M149140 (1 download); M149141 (2 downloads); M149142 (1 download); M149144 (1 download); M46117 (1 download); M46178 (1 download); M46179 (1 download); M46180 (1 download); M46446 (1 download); M46471 (1 download); M46472 (1 download); M46812 (1 download); M46815 (1 download); M46950 (1 download); M46986 (1 download); M47034 (1 download); M47875 (1 download); M48048 (1 download); M148176 (1 download); M151746 (1 download); M151747 (1 download); M148792 (1 download); M148793 (1 download); M148794 (1 download); M148796 (1 download); M148797 (1 download); M148799 (1 download); M148800 (1 download); M148801 (2 downloads); M148802 (1 download); M148803 (1 download); M148804 (1 download); M148805 (1 download); M148806 (1 download); M148807 (1 download); M148808 (1 download); M148809 (1 download); M148810 (1 download); M148811 (1 download); M46139 (1 download); M46943 (1 download); M46952 (1 download); M46992 (1 download); M47033 (1 download); M47793 (1 download); M47880 (1 download); M47963 (1 download); M46134 (1 download); M46135 (1 download); M46176 (1 download); M46177 (1 download); M46184 (1 download); M46444 (1 download); M46809 (1 download); M46810 (1 download); M46817 (1 download); M46843 (1 download); M46844 (1 download); M46976 (1 download); M46989 (1 download); M47032 (1 download); M47876 (1 download); M47975 (1 download); M98192 (1 download); M98193 (1 download); M98194 (1 download); M98195 (1 download); M98196 (1 download); M98197 (1 download); M98198 (1 download); M98201 (1 download); M98202 (1 download); M98203 (1 download); M98204 (1 download); M98206 (1 download); M98207 (1 download); M98208 (1 download); M98209 (1 download); M98210 (1 download); M98211 (1 download); M98212 (1 download); M98213 (1 download); M98214 (1 download); M98215 (1 download); M98216 (1 download); M98217 (1 download); M98218 (1 download); M98219 (1 download); M149298 (1 download); M46185 (1 download); M46186 (1 download); M46443 (1 download); M46449 (1 download); M46475 (1 download); M46476 (1 download); M46814 (1 download); M46842 (1 download); M46904 (1 download); M46988 (1 download); M47882 (1 download); M47883 (1 download); M47950 (1 download); M48055 (1 download); M46148 (1 download); M46151 (1 download); M46188 (1 download); M46451 (1 download); M46452 (1 download); M46823 (1 download); M46845 (1 download); M46910 (1 download); M46994 (1 download); M149299 (1 download); M46144 (1 download); M46145 (1 download); M46146 (1 download); M46147 (1 download); M46437 (1 download); M46822 (1 download); M46908 (1 download); M46909 (1 download); M46978 (1 download); M46993 (1 download); M46152 (2 downloads); M46153 (1 download); M46154 (1 download); M46155 (1 download); M46166 (1 download); M46167 (1 download); M46189 (1 download); M46414 (1 download); M46434 (1 download); M46435 (1 download); M46436 (1 download); M46824 (1 download); M46825 (1 download); M46846 (1 download); M46847 (1 download); M46848 (1 download); M46911 (2 downloads); M46912 (1 download); M46913 (1 download); M46914 (1 download); M46915 (1 download); M46955 (1 download); M46956 (1 download); M46974 (1 download); M46995 (1 download); M46996 (1 download); M46997 (1 download); M46998 (1 download); M47794 (1 download); M47795 (1 download); M47885 (1 download); M47986 (1 download); M48029 (1 download); M48049 (1 download); M48058 (1 download); M48059 (1 download); M48060 (1 download); M48061 (1 download); M48062 (1 download); M107818 (1 download); M107820 (1 download); M107821 (1 download); M107822 (1 download); M107823 (1 download); M142314 (1 download); M142315 (1 download); M142316 (1 download); M142317 (1 download); M142318 (1 download); M142319 (1 download); M142320 (1 download); M142321 (1 download); M142322 (1 download); M149197 (1 download); M149208 (1 download); M149209 (1 download); M149210 (1 download); M149211 (1 download); M149212 (1 download); M149213 (1 download); M149214 (1 download); M286139 (1 download); M286140 (1 download); M286141 (1 download); M286142 (1 download); M286143 (1 download); M286144 (1 download); M286145 (1 download); M286146 (1 download); M286147 (1 download); M286148 (1 download); M286149 (1 download); M286150 (1 download); M286151 (1 download); M286152 (1 download); M286153 (1 download); M286154 (1 download); M286155 (1 download); M286156 (1 download); M46191 (1 download); M46361 (1 download); M46479 (1 download); M46480 (1 download); M46849 (1 download); M47774 (1 download); M47791 (1 download); M47797 (1 download); M47914 (1 download); M47915 (1 download); M48047 (1 download); M48056 (1 download); M48516 (1 download); M46192 (1 download); M46193 (1 download); M46194 (1 download); M46195 (1 download); M46197 (1 download); M46198 (1 download); M46199 (1 download); M46200 (1 download); M46202 (1 download); M46362 (1 download); M46412 (1 download); M46415 (1 download); M46430 (1 download); M46431 (1 download); M46450 (1 download); M46828 (1 download); M46829 (1 download); M46917 (1 download); M46946 (1 download); M46958 (1 download); M46959 (1 download); M46960 (1 download); M46983 (1 download); M47002 (1 download); M47003 (1 download); M47004 (1 download); M47005 (1 download); M47031 (1 download); M47775 (1 download); M47792 (1 download); M47858 (1 download); M47887 (1 download); M47888 (1 download); M47965 (1 download); M47974 (1 download); M48497 (1 download); M48498 (1 download); M48499 (1 download); M48500 (1 download); M48501 (1 download); M48502 (1 download); M48503 (1 download); M48504 (1 download); M48505 (1 download); M48506 (1 download); M48507 (1 download); M48508 (1 download); M48509 (1 download); M48510 (1 download); M48511 (1 download); M48512 (1 download); M48513 (1 download); M46203 (1 download); M46204 (1 download); M46205 (1 download); M46206 (1 download); M46207 (1 download); M46208 (1 download); M46209 (1 download); M46210 (1 download); M46211 (1 download); M46212 (1 download); M46403 (1 download); M46407 (1 download); M46424 (1 download); M46429 (1 download); M46484 (1 download); M46485 (1 download); M46830 (1 download); M46850 (1 download); M46851 (1 download); M46852 (1 download); M46918 (1 download); M46919 (1 download); M46920 (1 download); M46921 (1 download); M46961 (1 download); M47006 (1 download); M47007 (1 download); M47009 (1 download); M47010 (1 download); M47029 (1 download); M47889 (1 download); M47890 (1 download); M47954 (1 download); M106565 (1 download); M46172 (1 download); M46213 (1 download); M46214 (1 download); M46224 (1 download); M46230 (1 download); M46427 (1 download); M46428 (1 download); M46454 (1 download); M46486 (1 download); M46487 (1 download); M46853 (1 download); M46854 (1 download); M46922 (1 download); M46923 (1 download); M46945 (1 download); M46962 (1 download); M46979 (1 download); M47011 (1 download); M47786 (1 download); M47891 (1 download); M107814 (1 download); M107815 (1 download); M107816 (1 download); M107817 (1 download); M149145 (1 download); M149146 (1 download); M149147 (1 download); M149148 (1 download); M149149 (1 download); M149150 (1 download); M149151 (1 download); M149152 (1 download); M149153 (1 download); M149154 (1 download); M149155 (1 download); M149156 (1 download); M149157 (1 download); M46326 (1 download); M46963 (1 download); M47892 (1 download); M47874 (1 download); M47787 (1 download); M47785 (1 download); M47784 (1 download); M47012 (1 download); M46924 (1 download); 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