Project 352: E. V. Bärmann. Towards a comprehensive phylogeny of Bovidae, Chapter 2: A total evidence analysis of bovid phylogeny - Dissertation by Eva Bärmann, University of Cambridge UK.
Specimen: Raphicerus melanotis (ZMB/Mammals:2125)

Abstract

Combined-data analyses, also termed total evidence analyses, have proven to be beneficial for phylogeny reconstructions for a range of taxa. For Bovidae, the most diverse clade of ruminants (Artiodactyla, Mammalia), such an analysis was last performed a decade ago. Since then, the range of possible analysis techniques has expanded, now including model-based Bayesian inference that can also be used to incorporate non-molecular data. Furthermore, the possibility of making morphological data widely accessible online, providing detailed information on character states via photographs, facilitates the use and re-use of this type of data. This study presents a data set of 88 cranial, 19 soft-tissue, and 27 behavioural characters, sampled from 58 bovid species. These data are analysed in a simultaneous analysis with molecular data from nine genes (four mitochondrial and five nuclear markers), sampled for 57 bovid species. A few morphological and behavioural characters are identified as apomorphies of bovid taxa. Despite the fact that the majority of characters show homoplasies and cannot unambiguously be optimized on the phylogenetic tree, they allow for inferring the phylogenetic position of two fossil bovid species: †Turcoceros grangeri and †Gazella lydekkeri. The study is designed as a starting point for further combined data analyses of bovids or ruminants in general.


Read the article »

Project DOI: 10.7934/P352, http://dx.doi.org/10.7934/P352
This project containsMatrices
  • 806 Media
  • 3 Matrices
  • 86 Taxa
  • 212 Specimens
  • 134 Characters
Total size of project's media files: 1.13G

Download Project SDD File
Total scored cells: 8111
Total media associated with cells: 6
Total labels associated with cell media: 0
Characters
Total characters: 134
Total characters with associated media: 10
Total characters with media with labels: 0
Total character states: 332
Total character states with associated media: 170
Total character states with media with labels:122
Total unordered/ordered characters:134/0
Currently Viewing:
MorphoBank Project 352

    Authors' Institutions

    • University of Cambridge



    Members

    member name taxa specimens media chars character
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    notes  medialabels
    rules
    Eva Baermann
    Project Administrator
    Last logged in 05/08/13
    86212806134112801588111
    (7554, 429, 128)
    434600
    Robert Asher
    Full membership
    Last logged in 05/18/10
    00000000
    (0, 0, 0)
    0000


    Taxonomic Overview for Matrix 'M817' (61 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    0878620002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    0898500002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    0847931002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    0878511002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    0878710002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    0888700002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0838032002
    [8] Capra falconeri
    Last Modified in 12/05/13
    1848231002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0868111002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    0847731002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    0817752002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    0878601002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    0878501002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    0878311002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    0878401002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    0878601002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1828151002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    0878501002
    [19] Gazella gazella
    Last Modified in 12/05/13
    1838131002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0878401002
    [21] Gazella marica
    Last Modified in 12/05/13
    0808071002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0848131002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    0878301002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    0868411002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    0898501002
    [26] Kobus kob
    Last Modified in 12/05/13
    0827761002
    [27] Kobus leche
    Last Modified in 12/05/13
    0837741002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    0797881002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    0838241002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    0888500002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    0858521002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    0868511002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    0868611002
    [34] Nanger dama
    Last Modified in 12/05/13
    0878301002
    [35] Nanger granti
    Last Modified in 12/05/13
    0878201002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    0888301002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0858412002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    0878701002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    0878302002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    0898601002
    [41] Oryx beisa
    Last Modified in 12/05/13
    1868221002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    0878511202
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0848331002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    0878601002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    0908101002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0878301002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    0837841002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    0818061002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    0878402002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0868602002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0828152002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    0868221002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    0878321202
    [54] Redunca redunca
    Last Modified in 12/05/13
    0817861002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    0888112002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0878301002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    0898301202
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0847432002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    0878701002
    [60] † Gazella lydekkeri
    Last Modified in 12/05/13
    0680191002
    [61] † Turcoceros grangeri
    Last Modified in 12/05/13
    1691180002


    Taxonomic Overview for Matrix 'M848' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    022523002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    027100002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    026401002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    025020002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    027000002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    026010002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    026711002
    [8] Capra falconeri
    Last Modified in 12/05/13
    022023002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0221241002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    024921002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    029401002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    024221002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    017082002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018090002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    027000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    081190002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1161100002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    023140002
    [19] Gazella gazella
    Last Modified in 12/05/13
    027100002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0161110002
    [21] Gazella marica
    Last Modified in 12/05/13
    025720002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0131140002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    025120002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    026101002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    023004002
    [26] Kobus kob
    Last Modified in 12/05/13
    027402002
    [27] Kobus leche
    Last Modified in 12/05/13
    026304002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    018564002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    617504002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    025020002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    025020002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    027100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    026110002
    [34] Nanger dama
    Last Modified in 12/05/13
    019080002
    [35] Nanger granti
    Last Modified in 12/05/13
    026010002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    019080002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0100161002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    025220002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    080190002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    025211002
    [41] Oryx beisa
    Last Modified in 12/05/13
    026603002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    025120002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0140112002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    024803002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    026701002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0100170002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    020561002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    018581002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    025020002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0120150002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0100170002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    024203002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    024003002
    [54] Redunca redunca
    Last Modified in 12/05/13
    024303002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019153002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0152111002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    026101002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0143121002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    022123002


    Taxonomic Overview for Matrix 'M849' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    019800002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    019000002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    019900002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    018510002
    [5] Antilope cervicapra
    Last Modified in 12/05/13
    017520002
    [6] Antidorcas marsupialis
    Last Modified in 12/05/13
    019000002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0191200002
    [8] Capra falconeri
    Last Modified in 12/05/13
    018010002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    118600002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    019600002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    019700002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    019000002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    019000002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018610002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    019000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    017520002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    018510002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    018510002
    [19] Gazella gazella
    Last Modified in 12/05/13
    019600002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    017520002
    [21] Gazella marica
    Last Modified in 12/05/13
    019800002
    [22] Gazella spekei
    Last Modified in 12/05/13
    018510002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    019600002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    019000002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    019000002
    [26] Kobus kob
    Last Modified in 12/05/13
    019500002
    [27] Kobus leche
    Last Modified in 12/05/13
    019500002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    019500002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    019600002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    019000002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    019000002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    019100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    019000002
    [34] Nanger dama
    Last Modified in 12/05/13
    018510002
    [35] Nanger granti
    Last Modified in 12/05/13
    019000002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    018510002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    018210002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    018010002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    018410002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    019000002
    [41] Oryx beisa
    Last Modified in 12/05/13
    019900002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    019000002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    019000002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    019400002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    019600002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    019400002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    019300002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    019400002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    019000002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    018010002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    019000002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    019500002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    019000002
    [54] Redunca redunca
    Last Modified in 12/05/13
    019500002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019700002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    018010002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    019000002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0191300002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    019000002


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M107807 (141 views); M107817 (212 views); M107818 (189 views); M107821 (145 views); M142302 (116 views); M142304 (156 views); M142305 (166 views); M142307 (218 views); M142308 (129 views); M142309 (189 views); M148781 (188 views); M149147 (34 views); M148170 (114 views); M148782 (112 views); M148795 (123 views); M284576 (7 views); M149111 (38 views); M148171 (104 views); M46500 (221 views); M46842 (172 views); M148784 (119 views); M46495 (139 views); M46825 (197 views); M148785 (114 views); M148780 (120 views); M149225 (34 views); M98198 (4 views); M148799 (121 views); M148794 (119 views); M148152 (158 views); M148778 (164 views); M46343 (379 views); M46187 (398 views); M46167 (397 views); M46172 (453 views); M148173 (158 views); M46179 (440 views); M148820 (34 views); M148175 (139 views); M148788 (116 views); M149219 (33 views); M148814 (40 views); M149230 (22 views); M149193 (53 views); M148800 (115 views); M46335 (447 views); M148818 (29 views); M46148 (403 views); M46155 (440 views); M149170 (36 views); M107804 (130 views); M148169 (107 views); M98201 (7 views); M149208 (54 views); M148790 (160 views); M149145 (34 views); M148166 (114 views); M149141 (76 views); M149292 (32 views); M149259 (6 views); M148813 (35 views); M149150 (34 views); M148826 (31 views); M149122 (34 views); M148823 (48 views); M148798 (200 views); M149211 (37 views); M148827 (32 views); M149174 (27 views); M149160 (27 views); M148824 (31 views); M149135 (47 views); M148779 (149 views); M47883 (5 views); M149149 (29 views); M148812 (27 views); M149168 (32 views); M149192 (39 views); M149137 (26 views); M149189 (57 views); M149180 (49 views); M46348 (426 views); M149213 (30 views); M148803 (28 views); M149275 (5 views); M46852 (194 views); M47020 (5 views); M149253 (6 views); M149118 (34 views); M148805 (30 views); M149194 (33 views); M149143 (38 views); M148811 (52 views); M149223 (25 views); M149222 (33 views); M149159 (29 views); M149172 (29 views); M148801 (30 views); M148802 (31 views); M148804 (30 views); M148806 (27 views); M148807 (50 views); M148808 (72 views); M148809 (34 views); M148810 (26 views); M148815 (36 views); M148816 (54 views); M148817 (37 views); M148819 (26 views); M148821 (28 views); M148825 (36 views); M149110 (49 views); M149112 (26 views); M149113 (28 views); M149114 (36 views); M149115 (32 views); M149116 (29 views); M149117 (38 views); M149119 (37 views); M149123 (38 views); M149134 (71 views); M149136 (28 views); M149138 (27 views); M149139 (29 views); M149140 (34 views); M149142 (40 views); M149144 (33 views); M149146 (28 views); M149148 (26 views); M149151 (28 views); M149152 (38 views); M149153 (46 views); M149154 (36 views); M149155 (27 views); M149156 (28 views); M149157 (29 views); M149158 (39 views); M149161 (36 views); M149162 (26 views); M149163 (34 views); M149164 (30 views); M149165 (35 views); M149166 (34 views); M149167 (28 views); M149169 (26 views); M149173 (33 views); M149175 (35 views); M149176 (55 views); M149177 (57 views); M149179 (37 views); M149181 (54 views); M149182 (41 views); M149183 (38 views); M149184 (44 views); M149185 (39 views); M149187 (51 views); M149188 (64 views); M149190 (42 views); M149191 (89 views); M149196 (35 views); M149197 (45 views); M149198 (30 views); M149209 (34 views); M149210 (35 views); M149212 (29 views); M149214 (29 views); M149218 (30 views); M149220 (30 views); M149221 (23 views); M149224 (24 views); M149226 (26 views); M149227 (23 views); M149228 (29 views); M149229 (23 views); M149242 (8 views); M46411 (197 views); M46193 (402 views); M46333 (381 views); M46449 (157 views); M286147 (5 views); M46405 (177 views); M149233 (5 views); M48516 (6 views); M46824 (196 views); M47787 (4 views); M46919 (5 views); M46431 (165 views); M46905 (6 views); M47950 (5 views); M47788 (8 views); M47955 (6 views); M46188 (391 views); M46202 (379 views); M149289 (7 views); M47887 (4 views); M149282 (4 views); M46326 (371 views); M46421 (155 views); M46415 (197 views); M46453 (151 views); M46153 (426 views); M46904 (5 views); M46843 (200 views); M149286 (5 views); M151747 (4 views); M46851 (174 views); M46118 (392 views); M46127 (408 views); M46128 (397 views); M46134 (407 views); M46135 (405 views); M46144 (411 views); M46145 (420 views); M46146 (440 views); M46147 (420 views); M46151 (396 views); M46152 (400 views); M46154 (413 views); M46166 (405 views); M46169 (379 views); M46171 (458 views); M46176 (429 views); M46177 (405 views); M46178 (426 views); M46180 (405 views); M46184 (386 views); M46185 (395 views); M46186 (396 views); M46189 (410 views); M46190 (441 views); M46191 (392 views); M46192 (429 views); M46194 (369 views); M46195 (401 views); M46197 (401 views); M46198 (373 views); M46199 (404 views); M46200 (397 views); M46204 (364 views); M46205 (347 views); M46206 (372 views); M46207 (417 views); M46208 (359 views); M46210 (374 views); M46211 (372 views); M46212 (438 views); M46230 (447 views); M46327 (397 views); M46328 (460 views); M46329 (399 views); M46330 (396 views); M46331 (407 views); M46334 (426 views); M46337 (405 views); M46340 (401 views); M46341 (385 views); M46342 (387 views); M46344 (409 views); M46345 (375 views); M46350 (387 views); M46351 (356 views); M46352 (410 views); M46353 (392 views); M46354 (403 views); M46355 (414 views); M46356 (372 views); M46357 (409 views); M46358 (391 views); M46359 (396 views); M46361 (392 views); M46362 (399 views); M46403 (364 views); M46406 (200 views); M46407 (207 views); M46412 (190 views); M46413 (160 views); M46414 (162 views); M46416 (178 views); M46417 (150 views); M46418 (201 views); M46419 (203 views); M46420 (160 views); M46422 (149 views); M46423 (150 views); M46424 (190 views); M46425 (198 views); M46426 (140 views); M46427 (183 views); M46428 (164 views); M46429 (150 views); M46430 (178 views); M46432 (152 views); M46434 (182 views); M46436 (155 views); M46437 (165 views); M46438 (183 views); M46440 (137 views); M46441 (166 views); M46442 (189 views); M46443 (193 views); M46444 (202 views); M46446 (161 views); M46447 (130 views); M46448 (155 views); M46450 (150 views); M46451 (177 views); M46452 (196 views); M46454 (205 views); M46455 (181 views); M46456 (151 views); M46457 (149 views); M46458 (177 views); M46459 (154 views); M46471 (184 views); M46472 (156 views); M46475 (153 views); M46476 (202 views); M46479 (196 views); M46480 (178 views); M46481 (213 views); M46482 (141 views); M46483 (171 views); M46484 (177 views); M46485 (158 views); M46486 (149 views); M46487 (185 views); M46488 (166 views); M46489 (164 views); M46490 (155 views); M46492 (168 views); M46493 (156 views); M46494 (180 views); M46496 (151 views); M46497 (156 views); M46498 (180 views); M46499 (182 views); M46809 (143 views); M46810 (148 views); M46812 (144 views); M46814 (162 views); M46815 (148 views); M46817 (154 views); M46818 (180 views); M46819 (194 views); M46820 (166 views); M46822 (179 views); M46823 (185 views); M46827 (144 views); M46828 (176 views); M46829 (146 views); M46830 (163 views); M46831 (139 views); M46832 (163 views); M46833 (146 views); M46834 (157 views); M46835 (199 views); M46837 (167 views); M46838 (160 views); M46839 (134 views); M46840 (161 views); M46841 (143 views); M46844 (177 views); M46845 (144 views); M46846 (135 views); M46847 (146 views); M46848 (134 views); M46849 (156 views); M46850 (169 views); M46853 (175 views); M46854 (130 views); M46855 (162 views); M46856 (137 views); M46857 (159 views); M46858 (146 views); M46859 (147 views); M46860 (188 views); M46861 (177 views); M46862 (174 views); M46863 (142 views); M46864 (140 views); M284578 (9 views); M149294 (5 views); M149243 (4 views); M149268 (4 views); M149271 (6 views); M46946 (5 views); M47791 (8 views); M98218 (4 views); M149285 (4 views); M284582 (5 views); M46950 (5 views); M98192 (5 views); M284577 (8 views); M46986 (7 views); M286153 (6 views); M149231 (7 views); M286145 (4 views); M286141 (4 views); M149248 (6 views); M149291 (5 views); M47018 (7 views); M48485 (7 views); M47022 (5 views); M46995 (6 views); M47794 (4 views); M47030 (4 views); M47009 (4 views); M47948 (4 views); M47786 (4 views); M46998 (4 views); M47007 (4 views); M47779 (4 views); M47954 (4 views); M47785 (4 views); M47965 (7 views); M47899 (8 views); M47986 (4 views); M47778 (4 views); M47790 (4 views); M48056 (6 views); M47976 (4 views); M48488 (5 views); M48058 (4 views); M48029 (4 views); M47970 (5 views); M46988 (4 views); M98208 (5 views); M47892 (4 views); M47781 (5 views); M47953 (7 views); M48509 (4 views); M98204 (4 views); M47011 (4 views); M47915 (25 views); M47014 (4 views); M47010 (5 views); M47797 (5 views); M48512 (5 views); M47016 (4 views); M47905 (5 views); M47906 (4 views); M47886 (5 views); M98212 (5 views); M98213 (5 views); M47795 (4 views); M98196 (4 views); M47951 (4 views); M47978 (6 views); M48047 (45 views); M48048 (4 views); M48487 (8 views); M48059 (4 views); M98195 (6 views); M47003 (5 views); M48503 (5 views); M47964 (5 views); M48514 (4 views); M46921 (6 views); M51943 (7 views); M47917 (5 views); M286140 (4 views); M48062 (5 views); M98209 (5 views); M149260 (8 views); M149297 (7 views); M46957 (10 views); M47017 (8 views); M149283 (4 views); M149281 (3 views); M149264 (4 views); M149288 (5 views); M46966 (7 views); M149279 (4 views); M47796 (5 views); M47789 (4 views); M98215 (4 views); M47884 (5 views); M149290 (4 views); M48053 (8 views); M48511 (4 views); M46923 (6 views); M47903 (4 views); M46939 (6 views); M47890 (4 views); M46926 (7 views); M47977 (6 views); M46952 (8 views); M47031 (5 views); M46924 (6 views); M46960 (7 views); M47775 (4 views); M48508 (4 views); M48499 (5 views); M46969 (7 views); M46929 (6 views); M46976 (20 views); M46964 (6 views); M98203 (4 views); M46967 (7 views); M46937 (7 views); M46972 (8 views); M47782 (4 views); M46971 (6 views); M46981 (7 views); M98217 (4 views); M46982 (7 views); M46955 (6 views); M98214 (4 views); M46959 (9 views); M47005 (4 views); M47858 (4 views); M48507 (4 views); M46951 (7 views); M46910 (6 views); M48497 (7 views); M46961 (6 views); M47981 (5 views); M46956 (8 views); M46979 (6 views); M47885 (4 views); M46968 (8 views); M46965 (8 views); M48057 (5 views); M46940 (6 views); M46934 (6 views); M46945 (6 views); M46980 (11 views); M46973 (6 views); M46978 (6 views); M46928 (8 views); M46970 (6 views); M46920 (7 views); M46975 (9 views); M48502 (4 views); M46953 (6 views); M98197 (5 views); M46962 (6 views); M46984 (6 views); M98210 (5 views); M46911 (7 views); M46925 (6 views); M48049 (5 views); M46917 (7 views); M47975 (5 views); M46932 (6 views); M46909 (6 views); M48515 (5 views); M47877 (5 views); M98194 (4 views); M48500 (4 views); M47949 (6 views); M149246 (5 views); M98207 (5 views); M47006 (4 views); M149235 (4 views); M47904 (4 views); M47889 (4 views); M98219 (4 views); M286142 (6 views); M46913 (6 views); M48505 (5 views); M47888 (4 views); M47776 (4 views); M46915 (5 views); M47974 (4 views); M149295 (4 views); M47780 (4 views); M284579 (6 views); M149269 (4 views); M149239 (6 views); M47792 (4 views); M47777 (4 views); M47021 (4 views); M46914 (5 views); M47019 (4 views); M47004 (4 views); M47012 (4 views); M47916 (6 views); M47871 (4 views); M98211 (5 views); M48061 (4 views); M98202 (5 views); M48510 (7 views); M48506 (5 views); M48055 (4 views); M151745 (4 views); M149267 (4 views); M149252 (4 views); M149255 (5 views); M149298 (4 views); M46943 (5 views); M286149 (4 views); M286154 (6 views); M46927 (5 views); M149238 (4 views); M149266 (4 views); M149244 (5 views); M149280 (4 views); M286143 (4 views); M286139 (4 views); M286146 (4 views); M286152 (7 views); M149250 (4 views); M149274 (4 views); M149299 (4 views); M149277 (4 views); M286155 (6 views); M286156 (6 views); M149287 (5 views); M149241 (4 views); M151743 (7 views); M149232 (4 views); M286150 (4 views); M46941 (6 views); M47882 (4 views); M47963 (4 views); M46907 (5 views); M46963 (7 views); M46912 (6 views); M48486 (5 views); M48050 (6 views); M47952 (3 views); M46935 (6 views); M46918 (4 views); M48054 (3 views); M48498 (3 views); M46938 (4 views); M48051 (3 views); M284581 (5 views); M149237 (3 views); M149296 (3 views); M149293 (3 views); M284584 (4 views); M149234 (4 views); M284580 (5 views); M149258 (8 views); M149256 (3 views); M149263 (3 views); M149240 (3 views); M149257 (4 views); M149236 (5 views); M46991 (3 views); M46989 (3 views); M47783 (3 views); M46922 (7 views); M47784 (3 views); M149278 (3 views); M46865 (6 views); M47034 (3 views); M46993 (3 views); M47774 (3 views); M46983 (5 views); M47874 (3 views); M46908 (5 views); M46996 (3 views); M286148 (3 views); M47896 (3 views); M149245 (3 views); M47914 (29 views); M286144 (3 views); M286151 (4 views); M48501 (3 views); M47956 (6 views); M149247 (3 views); M149254 (3 views); M149249 (3 views); M46997 (5 views); M151744 (3 views); M151746 (3 views); M149270 (3 views); M98193 (38 views); M46994 (3 views); M51944 (5 views); M47013 (4 views); M48052 (4 views); M149273 (3 views); M47895 (3 views); M47033 (3 views); M149265 (3 views); M46977 (8 views); M47015 (3 views); M46992 (4 views); M149261 (4 views); M47793 (3 views); M48513 (3 views); M149276 (3 views); M149284 (3 views); M98206 (5 views); M46958 (6 views); M47875 (4 views); M47897 (3 views); M149272 (3 views); M46936 (6 views); M46974 (5 views); M46947 (8 views); M47001 (3 views); M149251 (3 views); M47032 (4 views); M47876 (3 views); M47002 (3 views); M47029 (3 views); M47859 (3 views); M48060 (3 views); M47880 (3 views); M47973 (3 views); M46930 (6 views); M47891 (3 views); M48504 (3 views); M284583 (6 views); M98216 (4 views); M46933 (5 views);
    Bibliography11471
    Taxon list13749
    Specimen list87399




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project1161
    Project downloads264
    Matrix downloads76Skull characters Bovidae (57 downloads); Behavior (9 downloads); Glands and Colouration (10 downloads);
    Media downloads821M142313 (2 downloads); M142296 (1 download); M148795 (4 downloads); M148798 (3 downloads); M46337 (2 downloads); M46421 (2 downloads); M46492 (1 download); M47899 (2 downloads); M48485 (2 downloads); M48486 (2 downloads); M48487 (2 downloads); M48488 (2 downloads); M46418 (2 downloads); M46440 (2 downloads); M46820 (2 downloads); M149138 (2 downloads); M149143 (2 downloads); M107819 (2 downloads); M46187 (1 download); M46438 (1 download); M46907 (1 download); M46953 (1 download); M47789 (1 download); M47884 (1 download); M47978 (1 download); M149134 (1 download); M149135 (1 download); M149136 (1 download); M149137 (1 download); M149139 (2 downloads); M149140 (1 download); M149141 (2 downloads); M149142 (1 download); M149144 (1 download); M46117 (1 download); M46178 (1 download); M46179 (1 download); M46180 (1 download); M46446 (1 download); M46471 (1 download); M46472 (1 download); M46812 (1 download); M46815 (1 download); M46950 (1 download); M46986 (1 download); M47034 (1 download); M47875 (1 download); M48048 (1 download); M148176 (1 download); M151746 (1 download); M151747 (1 download); M148792 (1 download); M148793 (1 download); M148794 (1 download); M148796 (1 download); M148797 (1 download); M148799 (1 download); M148800 (1 download); M148801 (2 downloads); M148802 (1 download); M148803 (1 download); M148804 (1 download); M148805 (1 download); M148806 (1 download); M148807 (1 download); M148808 (1 download); M148809 (1 download); M148810 (1 download); M148811 (1 download); M46139 (1 download); M46943 (1 download); M46952 (1 download); M46992 (1 download); M47033 (1 download); M47793 (1 download); M47880 (1 download); M47963 (1 download); M46134 (1 download); M46135 (1 download); M46176 (1 download); M46177 (1 download); M46184 (1 download); M46444 (1 download); M46809 (1 download); M46810 (1 download); M46817 (1 download); M46843 (1 download); M46844 (1 download); M46976 (1 download); M46989 (1 download); M47032 (1 download); M47876 (1 download); M47975 (1 download); M98192 (1 download); M98193 (1 download); M98194 (1 download); M98195 (1 download); M98196 (1 download); M98197 (1 download); M98198 (1 download); M98201 (1 download); M98202 (1 download); M98203 (1 download); M98204 (1 download); M98206 (1 download); M98207 (1 download); M98208 (1 download); M98209 (1 download); M98210 (1 download); M98211 (1 download); M98212 (1 download); M98213 (1 download); M98214 (1 download); M98215 (1 download); M98216 (1 download); M98217 (1 download); M98218 (1 download); M98219 (1 download); M149298 (1 download); M46185 (1 download); M46186 (1 download); M46443 (1 download); M46449 (1 download); M46475 (1 download); M46476 (1 download); M46814 (1 download); M46842 (1 download); M46904 (1 download); M46988 (1 download); M47882 (1 download); M47883 (1 download); M47950 (1 download); M48055 (1 download); M46148 (1 download); M46151 (1 download); M46188 (1 download); M46451 (1 download); M46452 (1 download); M46823 (1 download); M46845 (1 download); M46910 (1 download); M46994 (1 download); M149299 (1 download); M46144 (1 download); M46145 (1 download); M46146 (1 download); M46147 (1 download); M46437 (1 download); M46822 (1 download); M46908 (1 download); M46909 (1 download); M46978 (1 download); M46993 (1 download); M46152 (2 downloads); M46153 (1 download); M46154 (1 download); M46155 (1 download); M46166 (1 download); M46167 (1 download); M46189 (1 download); M46414 (1 download); M46434 (1 download); M46435 (1 download); M46436 (1 download); M46824 (1 download); M46825 (1 download); M46846 (1 download); M46847 (1 download); M46848 (1 download); M46911 (2 downloads); M46912 (1 download); M46913 (1 download); M46914 (1 download); M46915 (1 download); M46955 (1 download); M46956 (1 download); M46974 (1 download); M46995 (1 download); M46996 (1 download); M46997 (1 download); M46998 (1 download); M47794 (1 download); M47795 (1 download); M47885 (1 download); M47986 (1 download); M48029 (1 download); M48049 (1 download); M48058 (1 download); M48059 (1 download); M48060 (1 download); M48061 (1 download); M48062 (1 download); M107818 (1 download); M107820 (1 download); M107821 (1 download); M107822 (1 download); M107823 (1 download); M142314 (1 download); M142315 (1 download); M142316 (1 download); M142317 (1 download); M142318 (1 download); M142319 (1 download); M142320 (1 download); M142321 (1 download); M142322 (1 download); M149197 (1 download); M149208 (1 download); M149209 (1 download); M149210 (1 download); M149211 (1 download); M149212 (1 download); M149213 (1 download); M149214 (1 download); M286139 (1 download); M286140 (1 download); M286141 (1 download); M286142 (1 download); M286143 (1 download); M286144 (1 download); M286145 (1 download); M286146 (1 download); M286147 (1 download); M286148 (1 download); M286149 (1 download); M286150 (1 download); M286151 (1 download); M286152 (1 download); M286153 (1 download); M286154 (1 download); M286155 (1 download); M286156 (1 download); M46191 (1 download); M46361 (1 download); M46479 (1 download); M46480 (1 download); M46849 (1 download); M47774 (1 download); M47791 (1 download); M47797 (1 download); M47914 (1 download); M47915 (1 download); M48047 (1 download); M48056 (1 download); M48516 (1 download); M46192 (1 download); M46193 (1 download); M46194 (1 download); M46195 (1 download); M46197 (1 download); M46198 (1 download); M46199 (1 download); M46200 (1 download); M46202 (1 download); M46362 (1 download); M46412 (1 download); M46415 (1 download); M46430 (1 download); M46431 (1 download); M46450 (1 download); M46828 (1 download); M46829 (1 download); M46917 (1 download); M46946 (1 download); M46958 (1 download); M46959 (1 download); M46960 (1 download); M46983 (1 download); M47002 (1 download); M47003 (1 download); M47004 (1 download); M47005 (1 download); M47031 (1 download); M47775 (1 download); M47792 (1 download); M47858 (1 download); M47887 (1 download); M47888 (1 download); M47965 (1 download); M47974 (1 download); M48497 (1 download); M48498 (1 download); M48499 (1 download); M48500 (1 download); M48501 (1 download); M48502 (1 download); M48503 (1 download); M48504 (1 download); M48505 (1 download); M48506 (1 download); M48507 (1 download); M48508 (1 download); M48509 (1 download); M48510 (1 download); M48511 (1 download); M48512 (1 download); M48513 (1 download); M46203 (1 download); M46204 (1 download); M46205 (1 download); M46206 (1 download); M46207 (1 download); M46208 (1 download); M46209 (1 download); M46210 (1 download); M46211 (1 download); M46212 (1 download); M46403 (1 download); M46407 (1 download); M46424 (1 download); M46429 (1 download); M46484 (1 download); M46485 (1 download); M46830 (1 download); M46850 (1 download); M46851 (1 download); M46852 (1 download); M46918 (1 download); M46919 (1 download); M46920 (1 download); M46921 (1 download); M46961 (1 download); M47006 (1 download); M47007 (1 download); M47009 (1 download); M47010 (1 download); M47029 (1 download); M47889 (1 download); M47890 (1 download); M47954 (1 download); 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