Project 352: E. V. Bärmann. Towards a comprehensive phylogeny of Bovidae, Chapter 2: A total evidence analysis of bovid phylogeny - Dissertation by Eva Bärmann, University of Cambridge UK.
Specimen: Raphicerus melanotis (ZMB/Mammals:2125)

Abstract

Combined-data analyses, also termed total evidence analyses, have proven to be beneficial for phylogeny reconstructions for a range of taxa. For Bovidae, the most diverse clade of ruminants (Artiodactyla, Mammalia), such an analysis was last performed a decade ago. Since then, the range of possible analysis techniques has expanded, now including model-based Bayesian inference that can also be used to incorporate non-molecular data. Furthermore, the possibility of making morphological data widely accessible online, providing detailed information on character states via photographs, facilitates the use and re-use of this type of data. This study presents a data set of 88 cranial, 19 soft-tissue, and 27 behavioural characters, sampled from 58 bovid species. These data are analysed in a simultaneous analysis with molecular data from nine genes (four mitochondrial and five nuclear markers), sampled for 57 bovid species. A few morphological and behavioural characters are identified as apomorphies of bovid taxa. Despite the fact that the majority of characters show homoplasies and cannot unambiguously be optimized on the phylogenetic tree, they allow for inferring the phylogenetic position of two fossil bovid species: †Turcoceros grangeri and †Gazella lydekkeri. The study is designed as a starting point for further combined data analyses of bovids or ruminants in general.


Read the article »

Project DOI: 10.7934/P352, http://dx.doi.org/10.7934/P352
This project containsMatrices
  • 806 Media
  • 3 Matrices
  • 86 Taxa
  • 212 Specimens
  • 134 Characters
Total size of project's media files: 1.13G

Download Project SDD File
Total scored cells: 8111
Total media associated with cells: 6
Total labels associated with cell media: 0
Characters
Total characters: 134
Total characters with associated media: 10
Total characters with media with labels: 0
Total character states: 332
Total character states with associated media: 170
Total character states with media with labels:122
Total unordered/ordered characters:134/0
Currently Viewing:
MorphoBank Project 352

    Authors' Institutions

    • University of Cambridge



    Members

    member name taxa specimens media chars character
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    notes  medialabels
    rules
    Eva Baermann
    Project Administrator
    86212806134112801588111
    (7554, 429, 128)
    434600
    Robert Asher
    Full membership
    00000000
    (0, 0, 0)
    0000


    Taxonomic Overview for Matrix 'M817' (61 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    0878620002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    0898500002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    0847931002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    0878511002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    0878710002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    0888700002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0838032002
    [8] Capra falconeri
    Last Modified in 12/05/13
    1848231002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0868111002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    0847731002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    0817752002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    0878601002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    0878501002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    0878311002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    0878401002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    0878601002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1828151002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    0878501002
    [19] Gazella gazella
    Last Modified in 12/05/13
    1838131002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0878401002
    [21] Gazella marica
    Last Modified in 12/05/13
    0808071002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0848131002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    0878301002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    0868411002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    0898501002
    [26] Kobus kob
    Last Modified in 12/05/13
    0827761002
    [27] Kobus leche
    Last Modified in 12/05/13
    0837741002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    0797881002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    0838241002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    0888500002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    0858521002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    0868511002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    0868611002
    [34] Nanger dama
    Last Modified in 12/05/13
    0878301002
    [35] Nanger granti
    Last Modified in 12/05/13
    0878201002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    0888301002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0858412002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    0878701002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    0878302002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    0898601002
    [41] Oryx beisa
    Last Modified in 12/05/13
    1868221002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    0878511202
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0848331002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    0878601002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    0908101002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0878301002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    0837841002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    0818061002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    0878402002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0868602002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0828152002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    0868221002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    0878321202
    [54] Redunca redunca
    Last Modified in 12/05/13
    0817861002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    0888112002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0878301002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    0898301202
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0847432002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    0878701002
    [60] † Gazella lydekkeri
    Last Modified in 12/05/13
    0680191002
    [61] † Turcoceros grangeri
    Last Modified in 12/05/13
    1691180002


    Taxonomic Overview for Matrix 'M848' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    022523002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    027100002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    026401002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    025020002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    027000002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    026010002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    026711002
    [8] Capra falconeri
    Last Modified in 12/05/13
    022023002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0221241002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    024921002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    029401002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    024221002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    017082002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018090002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    027000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    081190002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1161100002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    023140002
    [19] Gazella gazella
    Last Modified in 12/05/13
    027100002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0161110002
    [21] Gazella marica
    Last Modified in 12/05/13
    025720002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0131140002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    025120002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    026101002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    023004002
    [26] Kobus kob
    Last Modified in 12/05/13
    027402002
    [27] Kobus leche
    Last Modified in 12/05/13
    026304002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    018564002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    617504002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    025020002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    025020002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    027100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    026110002
    [34] Nanger dama
    Last Modified in 12/05/13
    019080002
    [35] Nanger granti
    Last Modified in 12/05/13
    026010002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    019080002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0100161002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    025220002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    080190002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    025211002
    [41] Oryx beisa
    Last Modified in 12/05/13
    026603002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    025120002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0140112002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    024803002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    026701002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0100170002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    020561002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    018581002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    025020002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0120150002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0100170002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    024203002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    024003002
    [54] Redunca redunca
    Last Modified in 12/05/13
    024303002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019153002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0152111002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    026101002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0143121002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    022123002


    Taxonomic Overview for Matrix 'M849' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    019800002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    019000002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    019900002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    018510002
    [5] Antilope cervicapra
    Last Modified in 12/05/13
    017520002
    [6] Antidorcas marsupialis
    Last Modified in 12/05/13
    019000002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0191200002
    [8] Capra falconeri
    Last Modified in 12/05/13
    018010002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    118600002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    019600002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    019700002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    019000002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    019000002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018610002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    019000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    017520002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    018510002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    018510002
    [19] Gazella gazella
    Last Modified in 12/05/13
    019600002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    017520002
    [21] Gazella marica
    Last Modified in 12/05/13
    019800002
    [22] Gazella spekei
    Last Modified in 12/05/13
    018510002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    019600002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    019000002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    019000002
    [26] Kobus kob
    Last Modified in 12/05/13
    019500002
    [27] Kobus leche
    Last Modified in 12/05/13
    019500002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    019500002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    019600002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    019000002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    019000002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    019100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    019000002
    [34] Nanger dama
    Last Modified in 12/05/13
    018510002
    [35] Nanger granti
    Last Modified in 12/05/13
    019000002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    018510002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    018210002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    018010002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    018410002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    019000002
    [41] Oryx beisa
    Last Modified in 12/05/13
    019900002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    019000002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    019000002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    019400002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    019600002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    019400002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    019300002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    019400002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    019000002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    018010002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    019000002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    019500002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    019000002
    [54] Redunca redunca
    Last Modified in 12/05/13
    019500002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019700002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    018010002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    019000002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0191300002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    019000002


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M107807 (150 views); M107817 (225 views); M107818 (189 views); M107821 (153 views); M142302 (125 views); M142304 (161 views); M142305 (179 views); M142307 (234 views); M142308 (134 views); M142309 (194 views); M148781 (188 views); M149147 (34 views); M148170 (121 views); M148782 (112 views); M148795 (123 views); M284576 (7 views); M149111 (38 views); M148171 (112 views); M46500 (255 views); M46842 (214 views); M148784 (119 views); M46495 (179 views); M46825 (227 views); M148785 (114 views); M148780 (120 views); M149225 (34 views); M98198 (4 views); M148799 (121 views); M148794 (119 views); M148152 (158 views); M148778 (164 views); M46343 (438 views); M46187 (469 views); M46167 (488 views); M46172 (523 views); M148173 (165 views); M46179 (523 views); M148820 (34 views); M148175 (142 views); M148788 (116 views); M149219 (33 views); M148814 (40 views); M149230 (22 views); M149193 (60 views); M148800 (115 views); M46335 (523 views); M148818 (29 views); M46148 (484 views); M46155 (527 views); M149170 (36 views); M107804 (137 views); M148169 (113 views); M98201 (7 views); M149208 (54 views); M148790 (160 views); M149145 (34 views); M148166 (114 views); M149141 (91 views); M149292 (34 views); M149259 (6 views); M148813 (35 views); M149150 (34 views); M148826 (31 views); M149122 (34 views); M148823 (48 views); M148798 (214 views); M149211 (37 views); M148827 (32 views); M149174 (27 views); M149160 (27 views); M148824 (31 views); M149135 (47 views); M148779 (149 views); M47883 (5 views); M149149 (29 views); M148812 (27 views); M149168 (32 views); M149192 (39 views); M149137 (26 views); M149189 (64 views); M149180 (56 views); M46348 (494 views); M149213 (30 views); M148803 (28 views); M149275 (5 views); M46852 (226 views); M47020 (5 views); M149253 (6 views); M149118 (34 views); M148805 (30 views); M149194 (33 views); M149143 (38 views); M148811 (52 views); M149223 (25 views); M149222 (33 views); M149159 (29 views); M149172 (29 views); M148801 (30 views); M148802 (31 views); M148804 (30 views); M148806 (27 views); M148807 (50 views); M148808 (72 views); M148809 (34 views); M148810 (26 views); M148815 (36 views); M148816 (54 views); M148817 (37 views); M148819 (26 views); M148821 (28 views); M148825 (36 views); M149110 (49 views); M149112 (26 views); M149113 (28 views); M149114 (36 views); M149115 (32 views); M149116 (29 views); M149117 (38 views); M149119 (37 views); M149123 (38 views); M149134 (71 views); M149136 (28 views); M149138 (27 views); M149139 (29 views); M149140 (34 views); M149142 (40 views); M149144 (33 views); M149146 (28 views); M149148 (26 views); M149151 (28 views); M149152 (38 views); M149153 (46 views); M149154 (36 views); M149155 (27 views); M149156 (28 views); M149157 (29 views); M149158 (39 views); M149161 (36 views); M149162 (26 views); M149163 (34 views); M149164 (30 views); M149165 (35 views); M149166 (34 views); M149167 (28 views); M149169 (26 views); M149173 (33 views); M149175 (35 views); M149176 (62 views); M149177 (64 views); M149179 (44 views); M149181 (61 views); M149182 (48 views); M149183 (38 views); M149184 (51 views); M149185 (46 views); M149187 (58 views); M149188 (71 views); M149190 (42 views); M149191 (95 views); M149196 (35 views); M149197 (47 views); M149198 (30 views); M149209 (34 views); M149210 (35 views); M149212 (29 views); M149214 (29 views); M149218 (30 views); M149220 (30 views); M149221 (23 views); M149224 (24 views); M149226 (26 views); M149227 (23 views); M149228 (29 views); M149229 (23 views); M149242 (8 views); M46411 (229 views); M46193 (480 views); M46333 (439 views); M46449 (195 views); M286147 (5 views); M46405 (216 views); M149233 (5 views); M48516 (6 views); M46824 (226 views); M47787 (4 views); M46919 (5 views); M46431 (198 views); M46905 (6 views); M47950 (5 views); M47788 (8 views); M47955 (6 views); M46188 (466 views); M46202 (438 views); M149289 (7 views); M47887 (4 views); M149282 (4 views); M46326 (429 views); M46421 (193 views); M46415 (236 views); M46453 (188 views); M46153 (508 views); M46904 (5 views); M46843 (235 views); M149286 (5 views); M151747 (4 views); M46851 (222 views); M46118 (460 views); M46127 (488 views); M46128 (466 views); M46134 (483 views); M46135 (484 views); M46144 (488 views); M46145 (495 views); M46146 (515 views); M46147 (494 views); M46151 (473 views); M46152 (474 views); M46154 (483 views); M46166 (486 views); M46169 (447 views); M46171 (525 views); M46176 (494 views); M46177 (470 views); M46178 (497 views); M46180 (473 views); M46184 (460 views); M46185 (468 views); M46186 (463 views); M46189 (472 views); M46190 (503 views); M46191 (468 views); M46192 (517 views); M46194 (424 views); M46195 (476 views); M46197 (457 views); M46198 (433 views); M46199 (462 views); M46200 (456 views); M46204 (428 views); M46205 (407 views); M46206 (430 views); M46207 (497 views); M46208 (421 views); M46210 (437 views); M46211 (435 views); M46212 (511 views); M46230 (522 views); M46327 (455 views); M46328 (515 views); M46329 (459 views); M46330 (448 views); M46331 (459 views); M46334 (493 views); M46337 (457 views); M46340 (460 views); M46341 (442 views); M46342 (451 views); M46344 (462 views); M46345 (438 views); M46350 (435 views); M46351 (411 views); M46352 (463 views); M46353 (444 views); M46354 (458 views); M46355 (468 views); M46356 (425 views); M46357 (469 views); M46358 (459 views); M46359 (461 views); M46361 (444 views); M46362 (452 views); M46403 (424 views); M46406 (240 views); M46407 (248 views); M46412 (227 views); M46413 (195 views); M46414 (197 views); M46416 (212 views); M46417 (183 views); M46418 (236 views); M46419 (238 views); M46420 (195 views); M46422 (184 views); M46423 (184 views); M46424 (226 views); M46425 (240 views); M46426 (174 views); M46427 (217 views); M46428 (209 views); M46429 (189 views); M46430 (214 views); M46432 (190 views); M46434 (221 views); M46436 (194 views); M46437 (201 views); M46438 (219 views); M46440 (183 views); M46441 (203 views); M46442 (228 views); M46443 (235 views); M46444 (244 views); M46446 (195 views); M46447 (167 views); M46448 (192 views); M46450 (185 views); M46451 (210 views); M46452 (233 views); M46454 (241 views); M46455 (213 views); M46456 (185 views); M46457 (182 views); M46458 (213 views); M46459 (190 views); M46471 (216 views); M46472 (200 views); M46475 (187 views); M46476 (242 views); M46479 (231 views); M46480 (212 views); M46481 (247 views); M46482 (187 views); M46483 (205 views); M46484 (209 views); M46485 (202 views); M46486 (182 views); M46487 (218 views); M46488 (204 views); M46489 (196 views); M46490 (190 views); M46492 (210 views); M46493 (192 views); M46494 (220 views); M46496 (183 views); M46497 (191 views); M46498 (219 views); M46499 (218 views); M46809 (179 views); M46810 (183 views); M46812 (174 views); M46814 (202 views); M46815 (184 views); M46817 (189 views); M46818 (214 views); M46819 (221 views); M46820 (206 views); M46822 (213 views); M46823 (224 views); M46827 (180 views); M46828 (208 views); M46829 (177 views); M46830 (195 views); M46831 (173 views); M46832 (195 views); M46833 (179 views); M46834 (199 views); M46835 (240 views); M46837 (209 views); M46838 (201 views); M46839 (170 views); M46840 (194 views); M46841 (183 views); M46844 (212 views); M46845 (185 views); M46846 (175 views); M46847 (195 views); M46848 (171 views); M46849 (196 views); M46850 (207 views); M46853 (215 views); M46854 (167 views); M46855 (195 views); M46856 (174 views); M46857 (194 views); M46858 (183 views); M46859 (189 views); M46860 (232 views); M46861 (214 views); M46862 (215 views); M46863 (176 views); M46864 (179 views); M284578 (9 views); M149294 (5 views); M149243 (4 views); M149268 (4 views); M149271 (6 views); M46946 (5 views); M47791 (8 views); M98218 (4 views); M149285 (4 views); M284582 (5 views); M46950 (5 views); M98192 (7 views); M284577 (8 views); M46986 (7 views); M286153 (6 views); M149231 (7 views); M286145 (4 views); M286141 (4 views); M149248 (6 views); M149291 (5 views); M47018 (7 views); M48485 (7 views); M47022 (5 views); M46995 (6 views); M47794 (4 views); M47030 (4 views); M47009 (4 views); M47948 (4 views); M47786 (4 views); M46998 (4 views); M47007 (4 views); M47779 (4 views); M47954 (4 views); M47785 (4 views); M47965 (7 views); M47899 (8 views); M47986 (4 views); M47778 (4 views); M47790 (4 views); M48056 (6 views); M47976 (4 views); M48488 (5 views); M48058 (4 views); M48029 (4 views); M47970 (5 views); M46988 (4 views); M98208 (5 views); M47892 (4 views); M47781 (5 views); M47953 (7 views); M48509 (4 views); M98204 (4 views); M47011 (4 views); M47915 (25 views); M47014 (4 views); M47010 (5 views); M47797 (5 views); M48512 (5 views); M47016 (4 views); M47905 (5 views); M47906 (4 views); M47886 (5 views); M98212 (5 views); M98213 (5 views); M47795 (4 views); M98196 (4 views); M47951 (4 views); M47978 (6 views); M48047 (45 views); M48048 (4 views); M48487 (8 views); M48059 (4 views); M98195 (6 views); M47003 (5 views); M48503 (5 views); M47964 (5 views); M48514 (4 views); M46921 (6 views); M51943 (7 views); M47917 (5 views); M286140 (4 views); M48062 (5 views); M98209 (5 views); M149260 (8 views); M149297 (7 views); M46957 (10 views); M47017 (8 views); M149283 (4 views); M149281 (3 views); M149264 (4 views); M149288 (5 views); M46966 (7 views); M149279 (4 views); M47796 (5 views); M47789 (4 views); M98215 (4 views); M47884 (5 views); M149290 (4 views); M48053 (8 views); M48511 (4 views); M46923 (6 views); M47903 (4 views); M46939 (6 views); M47890 (4 views); M46926 (7 views); M47977 (6 views); M46952 (8 views); M47031 (5 views); M46924 (6 views); M46960 (7 views); M47775 (4 views); M48508 (4 views); M48499 (5 views); M46969 (7 views); M46929 (6 views); M46976 (20 views); M46964 (6 views); M98203 (4 views); M46967 (7 views); M46937 (7 views); M46972 (8 views); M47782 (4 views); M46971 (6 views); M46981 (7 views); M98217 (4 views); M46982 (7 views); M46955 (6 views); M98214 (4 views); M46959 (9 views); M47005 (4 views); M47858 (4 views); M48507 (4 views); M46951 (7 views); M46910 (6 views); M48497 (7 views); M46961 (6 views); M47981 (5 views); M46956 (8 views); M46979 (6 views); M47885 (4 views); M46968 (8 views); M46965 (8 views); M48057 (5 views); M46940 (6 views); M46934 (6 views); M46945 (6 views); M46980 (11 views); M46973 (6 views); M46978 (6 views); M46928 (8 views); M46970 (6 views); M46920 (7 views); M46975 (9 views); M48502 (4 views); M46953 (6 views); M98197 (5 views); M46962 (6 views); M46984 (6 views); M98210 (5 views); M46911 (7 views); M46925 (6 views); M48049 (5 views); M46917 (7 views); M47975 (5 views); M46932 (6 views); M46909 (6 views); M48515 (5 views); M47877 (5 views); M98194 (4 views); M48500 (4 views); M47949 (6 views); M149246 (5 views); M98207 (5 views); M47006 (4 views); M149235 (4 views); M47904 (4 views); M47889 (4 views); M98219 (4 views); M286142 (6 views); M46913 (6 views); M48505 (5 views); M47888 (4 views); M47776 (4 views); M46915 (5 views); M47974 (4 views); M149295 (4 views); M47780 (4 views); M284579 (6 views); M149269 (4 views); M149239 (6 views); M47792 (4 views); M47777 (4 views); M47021 (4 views); M46914 (5 views); M47019 (4 views); M47004 (4 views); M47012 (4 views); M47916 (6 views); M47871 (4 views); M98211 (5 views); M48061 (4 views); M98202 (5 views); M48510 (7 views); M48506 (5 views); M48055 (4 views); M151745 (4 views); M149267 (4 views); M149252 (4 views); M149255 (5 views); M149298 (4 views); M46943 (5 views); M286149 (4 views); M286154 (6 views); M46927 (5 views); M149238 (4 views); M149266 (4 views); M149244 (5 views); M149280 (4 views); M286143 (4 views); M286139 (4 views); M286146 (4 views); M286152 (7 views); M149250 (4 views); M149274 (4 views); M149299 (4 views); M149277 (4 views); M286155 (6 views); M286156 (6 views); M149287 (5 views); M149241 (4 views); M151743 (7 views); M149232 (4 views); M286150 (4 views); M46941 (6 views); M47882 (4 views); M47963 (4 views); M46907 (5 views); M46963 (7 views); M46912 (6 views); M48486 (5 views); M48050 (6 views); M47952 (3 views); M46935 (6 views); M46918 (4 views); M48054 (3 views); M48498 (3 views); M46938 (4 views); M48051 (3 views); M284581 (5 views); M149237 (3 views); M149296 (3 views); M149293 (3 views); M284584 (4 views); M149234 (4 views); M284580 (5 views); M149258 (8 views); M149256 (3 views); M149263 (3 views); M149240 (3 views); M149257 (4 views); M149236 (5 views); M46991 (3 views); M46989 (3 views); M47783 (3 views); M46922 (7 views); M47784 (3 views); M149278 (3 views); M46865 (6 views); M47034 (3 views); M46993 (3 views); M47774 (3 views); M46983 (5 views); M47874 (3 views); M46908 (5 views); M46996 (3 views); M286148 (3 views); M47896 (3 views); M149245 (3 views); M47914 (29 views); M286144 (3 views); M286151 (4 views); M48501 (3 views); M47956 (6 views); M149247 (3 views); M149254 (3 views); M149249 (3 views); M46997 (5 views); M151744 (3 views); M151746 (3 views); M149270 (3 views); M98193 (40 views); M46994 (3 views); M51944 (5 views); M47013 (4 views); M48052 (4 views); M149273 (3 views); M47895 (3 views); M47033 (3 views); M149265 (3 views); M46977 (8 views); M47015 (3 views); M46992 (4 views); M149261 (4 views); M47793 (3 views); M48513 (3 views); M149276 (3 views); M149284 (3 views); M98206 (5 views); M46958 (6 views); M47875 (4 views); M47897 (3 views); M149272 (3 views); M46936 (6 views); M46974 (5 views); M46947 (8 views); M47001 (3 views); M149251 (3 views); M47032 (4 views); M47876 (3 views); M47002 (3 views); M47029 (3 views); M47859 (3 views); M48060 (3 views); M47880 (3 views); M47973 (3 views); M46930 (6 views); M47891 (3 views); M48504 (3 views); M284583 (6 views); M98216 (4 views); M46933 (5 views);
    Bibliography13249
    Taxon list15646
    Specimen list101445




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project1214
    Project downloads311
    Matrix downloads78Skull characters Bovidae (59 downloads); Behavior (9 downloads); Glands and Colouration (10 downloads);
    Media downloads825M142313 (2 downloads); M142296 (1 download); M148795 (4 downloads); M148798 (3 downloads); M46337 (2 downloads); M46421 (2 downloads); M46492 (1 download); M47899 (2 downloads); M48485 (2 downloads); M48486 (2 downloads); M48487 (2 downloads); M48488 (2 downloads); M46418 (2 downloads); M46440 (2 downloads); M46820 (2 downloads); M149138 (2 downloads); M149143 (2 downloads); M107819 (2 downloads); M46187 (1 download); M46438 (1 download); M46907 (1 download); M46953 (1 download); M47789 (1 download); M47884 (1 download); M47978 (1 download); M149134 (1 download); M149135 (1 download); M149136 (1 download); M149137 (1 download); M149139 (2 downloads); M149140 (1 download); M149141 (2 downloads); M149142 (1 download); M149144 (1 download); M46117 (1 download); M46178 (1 download); M46179 (1 download); M46180 (1 download); M46446 (1 download); M46471 (1 download); M46472 (1 download); M46812 (1 download); M46815 (1 download); M46950 (1 download); M46986 (1 download); M47034 (1 download); M47875 (1 download); M48048 (1 download); M148176 (1 download); M151746 (1 download); M151747 (1 download); M148792 (1 download); M148793 (1 download); M148794 (1 download); M148796 (1 download); M148797 (1 download); M148799 (1 download); M148800 (1 download); M148801 (2 downloads); M148802 (1 download); M148803 (1 download); M148804 (1 download); M148805 (1 download); M148806 (1 download); M148807 (1 download); M148808 (1 download); M148809 (1 download); M148810 (1 download); M148811 (1 download); M46139 (1 download); M46943 (1 download); M46952 (1 download); M46992 (1 download); M47033 (1 download); M47793 (1 download); M47880 (1 download); M47963 (1 download); M46134 (1 download); M46135 (1 download); M46176 (1 download); M46177 (1 download); M46184 (1 download); M46444 (1 download); M46809 (1 download); M46810 (1 download); M46817 (1 download); M46843 (1 download); M46844 (1 download); M46976 (1 download); M46989 (1 download); M47032 (1 download); M47876 (1 download); M47975 (1 download); M98192 (1 download); M98193 (1 download); M98194 (1 download); M98195 (1 download); M98196 (1 download); M98197 (1 download); M98198 (1 download); M98201 (1 download); M98202 (1 download); M98203 (1 download); M98204 (1 download); M98206 (1 download); M98207 (1 download); M98208 (1 download); M98209 (1 download); M98210 (1 download); M98211 (1 download); M98212 (1 download); M98213 (1 download); M98214 (1 download); M98215 (1 download); M98216 (1 download); M98217 (1 download); M98218 (1 download); M98219 (1 download); M149298 (1 download); M46185 (1 download); M46186 (1 download); M46443 (1 download); M46449 (1 download); M46475 (1 download); M46476 (1 download); M46814 (1 download); M46842 (1 download); M46904 (1 download); M46988 (1 download); M47882 (1 download); M47883 (1 download); M47950 (1 download); M48055 (1 download); M46148 (1 download); M46151 (1 download); M46188 (1 download); M46451 (1 download); M46452 (1 download); M46823 (1 download); M46845 (1 download); M46910 (1 download); M46994 (1 download); M149299 (1 download); M46144 (1 download); M46145 (1 download); M46146 (1 download); M46147 (1 download); M46437 (1 download); M46822 (1 download); M46908 (1 download); M46909 (1 download); M46978 (1 download); M46993 (1 download); M46152 (2 downloads); M46153 (1 download); M46154 (1 download); M46155 (1 download); M46166 (1 download); M46167 (1 download); M46189 (1 download); M46414 (1 download); M46434 (1 download); M46435 (1 download); M46436 (1 download); M46824 (1 download); M46825 (1 download); M46846 (1 download); M46847 (1 download); M46848 (1 download); M46911 (2 downloads); M46912 (1 download); M46913 (1 download); M46914 (1 download); M46915 (1 download); M46955 (1 download); M46956 (1 download); M46974 (1 download); M46995 (1 download); M46996 (1 download); M46997 (1 download); M46998 (1 download); M47794 (1 download); M47795 (1 download); M47885 (1 download); M47986 (1 download); M48029 (1 download); M48049 (1 download); M48058 (1 download); M48059 (1 download); M48060 (1 download); M48061 (1 download); M48062 (1 download); M107818 (1 download); M107820 (1 download); M107821 (1 download); M107822 (1 download); M107823 (1 download); M142314 (1 download); M142315 (1 download); M142316 (1 download); M142317 (1 download); M142318 (1 download); M142319 (1 download); M142320 (1 download); M142321 (1 download); M142322 (1 download); M149197 (1 download); M149208 (1 download); M149209 (1 download); M149210 (1 download); M149211 (1 download); M149212 (1 download); M149213 (1 download); M149214 (1 download); M286139 (1 download); M286140 (1 download); M286141 (1 download); M286142 (1 download); M286143 (1 download); M286144 (1 download); M286145 (1 download); M286146 (1 download); M286147 (1 download); M286148 (1 download); M286149 (1 download); M286150 (1 download); M286151 (1 download); M286152 (1 download); M286153 (1 download); M286154 (1 download); M286155 (1 download); M286156 (1 download); M46191 (1 download); M46361 (1 download); M46479 (1 download); M46480 (1 download); M46849 (1 download); M47774 (1 download); M47791 (1 download); M47797 (1 download); M47914 (1 download); M47915 (1 download); M48047 (1 download); M48056 (1 download); M48516 (1 download); M46192 (1 download); M46193 (1 download); M46194 (1 download); M46195 (1 download); M46197 (1 download); M46198 (1 download); M46199 (1 download); M46200 (1 download); M46202 (1 download); M46362 (1 download); M46412 (1 download); M46415 (1 download); M46430 (1 download); M46431 (1 download); M46450 (1 download); M46828 (1 download); M46829 (1 download); M46917 (1 download); M46946 (1 download); M46958 (1 download); M46959 (1 download); M46960 (1 download); M46983 (1 download); M47002 (1 download); M47003 (1 download); M47004 (1 download); M47005 (1 download); M47031 (1 download); M47775 (1 download); M47792 (1 download); M47858 (1 download); M47887 (1 download); M47888 (1 download); M47965 (1 download); M47974 (1 download); M48497 (1 download); M48498 (1 download); M48499 (1 download); M48500 (1 download); M48501 (1 download); M48502 (1 download); M48503 (1 download); M48504 (1 download); M48505 (1 download); M48506 (1 download); M48507 (1 download); M48508 (1 download); M48509 (1 download); M48510 (1 download); M48511 (1 download); M48512 (1 download); M48513 (1 download); M46203 (1 download); M46204 (1 download); M46205 (1 download); M46206 (1 download); M46207 (1 download); M46208 (1 download); M46209 (1 download); M46210 (1 download); M46211 (1 download); M46212 (1 download); M46403 (1 download); M46407 (1 download); M46424 (1 download); M46429 (1 download); M46484 (1 download); M46485 (1 download); M46830 (1 download); M46850 (1 download); M46851 (1 download); M46852 (1 download); M46918 (1 download); M46919 (1 download); M46920 (1 download); M46921 (1 download); M46961 (1 download); M47006 (1 download); M47007 (1 download); M47009 (1 download); M47010 (1 download); M47029 (1 download); M47889 (1 download); M47890 (1 download); M47954 (1 download); M106565 (1 download); M46172 (1 download); M46213 (1 download); M46214 (1 download); M46224 (1 download); M46230 (1 download); M46427 (1 download); M46428 (1 download); M46454 (1 download); M46486 (1 download); M46487 (1 download); M46853 (1 download); M46854 (1 download); M46922 (1 download); M46923 (1 download); M46945 (1 download); M46962 (1 download); M46979 (1 download); M47011 (1 download); M47786 (1 download); M47891 (1 download); M107814 (1 download); M107815 (1 download); M107816 (1 download); M107817 (1 download); M149145 (1 download); M149146 (1 download); M149147 (1 download); M149148 (1 download); M149149 (1 download); M149150 (1 download); M149151 (1 download); M149152 (1 download); M149153 (1 download); M149154 (1 download); M149155 (1 download); M149156 (1 download); M149157 (1 download); M46326 (1 download); M46963 (1 download); M47892 (1 download); M47874 (1 download); M47787 (1 download); M47785 (1 download); M47784 (1 download); M47012 (1 download); M46924 (1 download); 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