Project 352: E. V. Bärmann. Towards a comprehensive phylogeny of Bovidae, Chapter 2: A total evidence analysis of bovid phylogeny - Dissertation by Eva Bärmann, University of Cambridge UK.
Specimen: Raphicerus melanotis (ZMB/Mammals:2125)

Abstract

Combined-data analyses, also termed total evidence analyses, have proven to be beneficial for phylogeny reconstructions for a range of taxa. For Bovidae, the most diverse clade of ruminants (Artiodactyla, Mammalia), such an analysis was last performed a decade ago. Since then, the range of possible analysis techniques has expanded, now including model-based Bayesian inference that can also be used to incorporate non-molecular data. Furthermore, the possibility of making morphological data widely accessible online, providing detailed information on character states via photographs, facilitates the use and re-use of this type of data. This study presents a data set of 88 cranial, 19 soft-tissue, and 27 behavioural characters, sampled from 58 bovid species. These data are analysed in a simultaneous analysis with molecular data from nine genes (four mitochondrial and five nuclear markers), sampled for 57 bovid species. A few morphological and behavioural characters are identified as apomorphies of bovid taxa. Despite the fact that the majority of characters show homoplasies and cannot unambiguously be optimized on the phylogenetic tree, they allow for inferring the phylogenetic position of two fossil bovid species: †Turcoceros grangeri and †Gazella lydekkeri. The study is designed as a starting point for further combined data analyses of bovids or ruminants in general.


Read the article »

Project DOI: 10.7934/P352, http://dx.doi.org/10.7934/P352
This project containsMatrices
  • 806 Media
  • 3 Matrices
  • 86 Taxa
  • 212 Specimens
  • 134 Characters
Total size of project's media files: 1468 MB

Download Project SDD File
Total scored cells: 8111
Total media associated with cells: 6
Total labels associated with cell media: 0
Characters
Total characters: 134
Total characters with associated media: 10
Total characters with media with labels: 0
Total character states: 332
Total character states with associated media: 170
Total character states with media with labels:122
Total unordered/ordered characters:134/0
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MorphoBank Project 352

    Authors' Institutions

    • the University of Cambridge



    Members

    member name taxa specimens media chars character
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    notes  medialabels
    rules
    Eva Baermann
    Project Administrator
    Last logged in 05/08/13
    86212806134112801588111
    (7554, 429, 128)
    443600
    Robert Asher
    Full membership
    Last logged in 05/18/10
    00000000
    (0, 0, 0)
    0000


    Taxonomic Overview for Matrix 'M817' (61 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    0878620002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    0898500002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    0847931002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    0878511002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    0878710002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    0888700002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0838032002
    [8] Capra falconeri
    Last Modified in 12/05/13
    1848231002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0868111002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    0847731002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    0817752002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    0878601002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    0878501002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    0878311002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    0878401002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    0878601002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1828151002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    0878501002
    [19] Gazella gazella
    Last Modified in 12/05/13
    1838131002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0878401002
    [21] Gazella marica
    Last Modified in 12/05/13
    0808071002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0848131002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    0878301002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    0868411002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    0898501002
    [26] Kobus kob
    Last Modified in 12/05/13
    0827761002
    [27] Kobus leche
    Last Modified in 12/05/13
    0837741002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    0797881002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    0838241002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    0888500002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    0858521002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    0868511002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    0868611002
    [34] Nanger dama
    Last Modified in 12/05/13
    0878301002
    [35] Nanger granti
    Last Modified in 12/05/13
    0878201002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    0888301002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0858412002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    0878701002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    0878302002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    0898601002
    [41] Oryx beisa
    Last Modified in 12/05/13
    1868221002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    0878511202
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0848331002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    0878601002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    0908101002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0878301002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    0837841002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    0818061002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    0878402002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0868602002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0828152002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    0868221002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    0878321202
    [54] Redunca redunca
    Last Modified in 12/05/13
    0817861002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    0888112002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0878301002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    0898301202
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0847432002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    0878701002
    [60] † Gazella lydekkeri
    Last Modified in 12/05/13
    0680191002
    [61] † Turcoceros grangeri
    Last Modified in 12/05/13
    1691180002


    Taxonomic Overview for Matrix 'M848' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    022523002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    027100002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    026401002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    025020002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    027000002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    026010002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    026711002
    [8] Capra falconeri
    Last Modified in 12/05/13
    022023002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0221241002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    024921002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    029401002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    024221002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    017082002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018090002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    027000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    081190002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1161100002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    023140002
    [19] Gazella gazella
    Last Modified in 12/05/13
    027100002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0161110002
    [21] Gazella marica
    Last Modified in 12/05/13
    025720002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0131140002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    025120002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    026101002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    023004002
    [26] Kobus kob
    Last Modified in 12/05/13
    027402002
    [27] Kobus leche
    Last Modified in 12/05/13
    026304002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    018564002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    617504002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    025020002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    025020002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    027100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    026110002
    [34] Nanger dama
    Last Modified in 12/05/13
    019080002
    [35] Nanger granti
    Last Modified in 12/05/13
    026010002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    019080002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0100161002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    025220002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    080190002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    025211002
    [41] Oryx beisa
    Last Modified in 12/05/13
    026603002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    025120002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0140112002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    024803002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    026701002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0100170002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    020561002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    018581002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    025020002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0120150002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0100170002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    024203002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    024003002
    [54] Redunca redunca
    Last Modified in 12/05/13
    024303002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019153002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0152111002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    026101002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0143121002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    022123002


    Taxonomic Overview for Matrix 'M849' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    019800002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    019000002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    019900002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    018510002
    [5] Antilope cervicapra
    Last Modified in 12/05/13
    017520002
    [6] Antidorcas marsupialis
    Last Modified in 12/05/13
    019000002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0191200002
    [8] Capra falconeri
    Last Modified in 12/05/13
    018010002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    118600002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    019600002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    019700002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    019000002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    019000002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018610002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    019000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    017520002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    018510002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    018510002
    [19] Gazella gazella
    Last Modified in 12/05/13
    019600002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    017520002
    [21] Gazella marica
    Last Modified in 12/05/13
    019800002
    [22] Gazella spekei
    Last Modified in 12/05/13
    018510002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    019600002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    019000002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    019000002
    [26] Kobus kob
    Last Modified in 12/05/13
    019500002
    [27] Kobus leche
    Last Modified in 12/05/13
    019500002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    019500002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    019600002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    019000002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    019000002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    019100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    019000002
    [34] Nanger dama
    Last Modified in 12/05/13
    018510002
    [35] Nanger granti
    Last Modified in 12/05/13
    019000002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    018510002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    018210002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    018010002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    018410002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    019000002
    [41] Oryx beisa
    Last Modified in 12/05/13
    019900002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    019000002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    019000002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    019400002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    019600002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    019400002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    019300002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    019400002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    019000002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    018010002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    019000002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    019500002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    019000002
    [54] Redunca redunca
    Last Modified in 12/05/13
    019500002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019700002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    018010002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    019000002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0191300002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    019000002


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M107807 (137 views); M107817 (205 views); M107818 (189 views); M107821 (141 views); M142302 (112 views); M142304 (153 views); M142305 (160 views); M142307 (210 views); M142308 (126 views); M142309 (186 views); M148781 (188 views); M149147 (34 views); M148170 (114 views); M148782 (112 views); M148795 (123 views); M284576 (7 views); M149111 (38 views); M148171 (104 views); M46500 (207 views); M46842 (157 views); M148784 (119 views); M46495 (124 views); M46825 (185 views); M148785 (114 views); M148780 (120 views); M149225 (34 views); M98198 (4 views); M148799 (121 views); M148794 (119 views); M148152 (158 views); M148778 (164 views); M46343 (356 views); M46187 (361 views); M46167 (366 views); M46172 (421 views); M148173 (158 views); M46179 (405 views); M148820 (34 views); M148175 (139 views); M148788 (116 views); M149219 (33 views); M148814 (40 views); M149230 (22 views); M149193 (50 views); M148800 (115 views); M46335 (413 views); M148818 (29 views); M46148 (372 views); M46155 (411 views); M149170 (36 views); M107804 (126 views); M148169 (107 views); M98201 (7 views); M149208 (54 views); M148790 (160 views); M149145 (34 views); M148166 (114 views); M149141 (74 views); M149292 (31 views); M149259 (6 views); M148813 (35 views); M149150 (34 views); M148826 (31 views); M149122 (34 views); M148823 (48 views); M148798 (196 views); M149211 (37 views); M148827 (32 views); M149174 (27 views); M149160 (27 views); M148824 (31 views); M149135 (47 views); M148779 (149 views); M47883 (5 views); M149149 (29 views); M148812 (27 views); M149168 (32 views); M149192 (39 views); M149137 (26 views); M149189 (54 views); M149180 (46 views); M46348 (394 views); M149213 (30 views); M148803 (28 views); M149275 (5 views); M46852 (180 views); M47020 (5 views); M149253 (6 views); M149118 (34 views); M148805 (30 views); M149194 (33 views); M149143 (38 views); M148811 (52 views); M149223 (25 views); M149222 (33 views); M149159 (29 views); M149172 (29 views); M148801 (30 views); M148802 (31 views); M148804 (30 views); M148806 (27 views); M148807 (50 views); M148808 (72 views); M148809 (34 views); M148810 (26 views); M148815 (36 views); M148816 (54 views); M148817 (37 views); M148819 (26 views); M148821 (28 views); M148825 (36 views); M149110 (49 views); M149112 (26 views); M149113 (28 views); M149114 (36 views); M149115 (32 views); M149116 (29 views); M149117 (38 views); M149119 (37 views); M149123 (38 views); M149134 (71 views); M149136 (28 views); M149138 (27 views); M149139 (29 views); M149140 (34 views); M149142 (40 views); M149144 (33 views); M149146 (28 views); M149148 (26 views); M149151 (28 views); M149152 (38 views); M149153 (46 views); M149154 (36 views); M149155 (27 views); M149156 (28 views); M149157 (29 views); M149158 (39 views); M149161 (36 views); M149162 (26 views); M149163 (34 views); M149164 (30 views); M149165 (35 views); M149166 (34 views); M149167 (28 views); M149169 (26 views); M149173 (33 views); M149175 (35 views); M149176 (52 views); M149177 (54 views); M149179 (34 views); M149181 (51 views); M149182 (38 views); M149183 (38 views); M149184 (41 views); M149185 (36 views); M149187 (48 views); M149188 (61 views); M149190 (42 views); M149191 (86 views); M149196 (35 views); M149197 (45 views); M149198 (30 views); M149209 (34 views); M149210 (35 views); M149212 (29 views); M149214 (29 views); M149218 (30 views); M149220 (30 views); M149221 (23 views); M149224 (24 views); M149226 (26 views); M149227 (23 views); M149228 (29 views); M149229 (23 views); M149242 (8 views); M46411 (182 views); M46193 (375 views); M46333 (352 views); M46449 (142 views); M286147 (5 views); M46405 (160 views); M149233 (5 views); M48516 (6 views); M46824 (182 views); M47787 (4 views); M46919 (5 views); M46431 (147 views); M46905 (6 views); M47950 (5 views); M47788 (8 views); M47955 (6 views); M46188 (361 views); M46202 (355 views); M149289 (7 views); M47887 (4 views); M149282 (4 views); M46326 (345 views); M46421 (139 views); M46415 (180 views); M46453 (134 views); M46153 (396 views); M46904 (5 views); M46843 (185 views); M149286 (5 views); M151747 (4 views); M46851 (157 views); M46118 (365 views); M46127 (378 views); M46128 (368 views); M46134 (378 views); M46135 (377 views); M46144 (379 views); M46145 (388 views); M46146 (410 views); M46147 (390 views); M46151 (365 views); M46152 (370 views); M46154 (382 views); M46166 (375 views); M46169 (350 views); M46171 (427 views); M46176 (395 views); M46177 (373 views); M46178 (400 views); M46180 (368 views); M46184 (360 views); M46185 (367 views); M46186 (362 views); M46189 (380 views); M46190 (406 views); M46191 (361 views); M46192 (393 views); M46194 (340 views); M46195 (371 views); M46197 (363 views); M46198 (346 views); M46199 (376 views); M46200 (371 views); M46204 (338 views); M46205 (324 views); M46206 (348 views); M46207 (388 views); M46208 (331 views); M46210 (350 views); M46211 (348 views); M46212 (411 views); M46230 (415 views); M46327 (367 views); M46328 (435 views); M46329 (366 views); M46330 (368 views); M46331 (381 views); M46334 (398 views); M46337 (374 views); M46340 (367 views); M46341 (359 views); M46342 (358 views); M46344 (382 views); M46345 (347 views); M46350 (363 views); M46351 (328 views); M46352 (378 views); M46353 (364 views); M46354 (374 views); M46355 (389 views); M46356 (343 views); M46357 (382 views); M46358 (362 views); M46359 (369 views); M46361 (362 views); M46362 (371 views); M46403 (335 views); M46406 (184 views); M46407 (192 views); M46412 (172 views); M46413 (144 views); M46414 (146 views); M46416 (161 views); M46417 (134 views); M46418 (185 views); M46419 (187 views); M46420 (145 views); M46422 (133 views); M46423 (134 views); M46424 (173 views); M46425 (181 views); M46426 (124 views); M46427 (167 views); M46428 (149 views); M46429 (134 views); M46430 (162 views); M46432 (137 views); M46434 (166 views); M46436 (140 views); M46437 (149 views); M46438 (167 views); M46440 (122 views); M46441 (150 views); M46442 (174 views); M46443 (176 views); M46444 (185 views); M46446 (146 views); M46447 (115 views); M46448 (140 views); M46450 (134 views); M46451 (161 views); M46452 (180 views); M46454 (189 views); M46455 (165 views); M46456 (135 views); M46457 (134 views); M46458 (162 views); M46459 (141 views); M46471 (167 views); M46472 (140 views); M46475 (138 views); M46476 (186 views); M46479 (180 views); M46480 (161 views); M46481 (197 views); M46482 (126 views); M46483 (156 views); M46484 (161 views); M46485 (141 views); M46486 (133 views); M46487 (171 views); M46488 (150 views); M46489 (148 views); M46490 (140 views); M46492 (153 views); M46493 (140 views); M46494 (165 views); M46496 (136 views); M46497 (141 views); M46498 (165 views); M46499 (167 views); M46809 (129 views); M46810 (135 views); M46812 (131 views); M46814 (148 views); M46815 (133 views); M46817 (141 views); M46818 (166 views); M46819 (180 views); M46820 (154 views); M46822 (168 views); M46823 (173 views); M46827 (131 views); M46828 (162 views); M46829 (133 views); M46830 (150 views); M46831 (128 views); M46832 (153 views); M46833 (132 views); M46834 (143 views); M46835 (185 views); M46837 (153 views); M46838 (147 views); M46839 (120 views); M46840 (147 views); M46841 (128 views); M46844 (163 views); M46845 (129 views); M46846 (120 views); M46847 (130 views); M46848 (118 views); M46849 (141 views); M46850 (155 views); M46853 (161 views); M46854 (116 views); M46855 (147 views); M46856 (121 views); M46857 (145 views); M46858 (131 views); M46859 (133 views); M46860 (174 views); M46861 (164 views); M46862 (159 views); M46863 (128 views); M46864 (126 views); M284578 (9 views); M149294 (5 views); M149243 (4 views); M149268 (4 views); M149271 (6 views); M46946 (5 views); M47791 (8 views); M98218 (4 views); M149285 (4 views); M284582 (5 views); M46950 (5 views); M98192 (5 views); M284577 (8 views); M46986 (7 views); M286153 (6 views); M149231 (7 views); M286145 (4 views); M286141 (4 views); M149248 (6 views); M149291 (5 views); M47018 (7 views); M48485 (7 views); M47022 (5 views); M46995 (6 views); M47794 (4 views); M47030 (4 views); M47009 (4 views); M47948 (4 views); M47786 (4 views); M46998 (4 views); M47007 (4 views); M47779 (4 views); M47954 (4 views); M47785 (4 views); M47965 (7 views); M47899 (8 views); M47986 (4 views); M47778 (4 views); M47790 (4 views); M48056 (6 views); M47976 (4 views); M48488 (5 views); M48058 (4 views); M48029 (4 views); M47970 (5 views); M46988 (4 views); M98208 (5 views); M47892 (4 views); M47781 (5 views); M47953 (7 views); M48509 (4 views); M98204 (4 views); M47011 (4 views); M47915 (25 views); M47014 (4 views); M47010 (5 views); M47797 (5 views); M48512 (5 views); M47016 (4 views); M47905 (5 views); M47906 (4 views); M47886 (5 views); M98212 (5 views); M98213 (5 views); M47795 (4 views); M98196 (4 views); M47951 (4 views); M47978 (6 views); M48047 (45 views); M48048 (4 views); M48487 (8 views); M48059 (4 views); M98195 (6 views); M47003 (5 views); M48503 (5 views); M47964 (5 views); M48514 (4 views); M46921 (6 views); M51943 (7 views); M47917 (5 views); M286140 (4 views); M48062 (5 views); M98209 (5 views); M149260 (8 views); M149297 (7 views); M46957 (10 views); M47017 (8 views); M149283 (4 views); M149281 (3 views); M149264 (4 views); M149288 (5 views); M46966 (7 views); M149279 (4 views); M47796 (5 views); M47789 (4 views); M98215 (4 views); M47884 (5 views); M149290 (4 views); M48053 (8 views); M48511 (4 views); M46923 (6 views); M47903 (4 views); M46939 (6 views); M47890 (4 views); M46926 (7 views); M47977 (6 views); M46952 (8 views); M47031 (5 views); M46924 (6 views); M46960 (7 views); M47775 (4 views); M48508 (4 views); M48499 (5 views); M46969 (7 views); M46929 (6 views); M46976 (20 views); M46964 (6 views); M98203 (4 views); M46967 (7 views); M46937 (7 views); M46972 (8 views); M47782 (4 views); M46971 (6 views); M46981 (7 views); M98217 (4 views); M46982 (7 views); M46955 (6 views); M98214 (4 views); M46959 (9 views); M47005 (4 views); M47858 (4 views); M48507 (4 views); M46951 (7 views); M46910 (6 views); M48497 (7 views); M46961 (6 views); M47981 (5 views); M46956 (8 views); M46979 (6 views); M47885 (4 views); M46968 (8 views); M46965 (8 views); M48057 (5 views); M46940 (6 views); M46934 (6 views); M46945 (6 views); M46980 (11 views); M46973 (6 views); M46978 (6 views); M46928 (8 views); M46970 (6 views); M46920 (7 views); M46975 (9 views); M48502 (4 views); M46953 (6 views); M98197 (5 views); M46962 (6 views); M46984 (6 views); M98210 (5 views); M46911 (7 views); M46925 (6 views); M48049 (5 views); M46917 (7 views); M47975 (5 views); M46932 (6 views); M46909 (6 views); M48515 (5 views); M47877 (5 views); M98194 (4 views); M48500 (4 views); M47949 (6 views); M149246 (5 views); M98207 (5 views); M47006 (4 views); M149235 (4 views); M47904 (4 views); M47889 (4 views); M98219 (4 views); M286142 (6 views); M46913 (6 views); M48505 (5 views); M47888 (4 views); M47776 (4 views); M46915 (5 views); M47974 (4 views); M149295 (4 views); M47780 (4 views); M284579 (6 views); M149269 (4 views); M149239 (6 views); M47792 (4 views); M47777 (4 views); M47021 (4 views); M46914 (5 views); M47019 (4 views); M47004 (4 views); M47012 (4 views); M47916 (6 views); M47871 (4 views); M98211 (5 views); M48061 (4 views); M98202 (5 views); M48510 (7 views); M48506 (5 views); M48055 (4 views); M151745 (4 views); M149267 (4 views); M149252 (4 views); M149255 (5 views); M149298 (4 views); M46943 (5 views); M286149 (4 views); M286154 (6 views); M46927 (5 views); M149238 (4 views); M149266 (4 views); M149244 (5 views); M149280 (4 views); M286143 (4 views); M286139 (4 views); M286146 (4 views); M286152 (7 views); M149250 (4 views); M149274 (4 views); M149299 (4 views); M149277 (4 views); M286155 (6 views); M286156 (6 views); M149287 (5 views); M149241 (4 views); M151743 (7 views); M149232 (4 views); M286150 (4 views); M46941 (6 views); M47882 (4 views); M47963 (4 views); M46907 (5 views); M46963 (7 views); M46912 (6 views); M48486 (5 views); M48050 (6 views); M47952 (3 views); M46935 (6 views); M46918 (4 views); M48054 (3 views); M48498 (3 views); M46938 (4 views); M48051 (3 views); M284581 (5 views); M149237 (3 views); M149296 (3 views); M149293 (3 views); M284584 (4 views); M149234 (4 views); M284580 (5 views); M149258 (8 views); M149256 (3 views); M149263 (3 views); M149240 (3 views); M149257 (4 views); M149236 (5 views); M46991 (3 views); M46989 (3 views); M47783 (3 views); M46922 (7 views); M47784 (3 views); M149278 (3 views); M46865 (6 views); M47034 (3 views); M46993 (3 views); M47774 (3 views); M46983 (5 views); M47874 (3 views); M46908 (5 views); M46996 (3 views); M286148 (3 views); M47896 (3 views); M149245 (3 views); M47914 (29 views); M286144 (3 views); M286151 (4 views); M48501 (3 views); M47956 (6 views); M149247 (3 views); M149254 (3 views); M149249 (3 views); M46997 (5 views); M151744 (3 views); M151746 (3 views); M149270 (3 views); M98193 (37 views); M46994 (3 views); M51944 (5 views); M47013 (4 views); M48052 (4 views); M149273 (3 views); M47895 (3 views); M47033 (3 views); M149265 (3 views); M46977 (8 views); M47015 (3 views); M46992 (4 views); M149261 (4 views); M47793 (3 views); M48513 (3 views); M149276 (3 views); M149284 (3 views); M98206 (5 views); M46958 (6 views); M47875 (4 views); M47897 (3 views); M149272 (3 views); M46936 (6 views); M46974 (5 views); M46947 (8 views); M47001 (3 views); M149251 (3 views); M47032 (4 views); M47876 (3 views); M47002 (3 views); M47029 (3 views); M47859 (3 views); M48060 (3 views); M47880 (3 views); M47973 (3 views); M46930 (6 views); M47891 (3 views); M48504 (3 views); M284583 (6 views); M98216 (4 views); M46933 (5 views);
    Bibliography10507
    Taxon list11935
    Specimen list81410




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project1125
    Project downloads240
    Matrix downloads64Skull characters Bovidae (45 downloads); Behavior (9 downloads); Glands and Colouration (10 downloads);
    Media downloads821M142313 (2 downloads); M142296 (1 download); M148795 (4 downloads); M148798 (3 downloads); M46337 (2 downloads); M46421 (2 downloads); M46492 (1 download); M47899 (2 downloads); M48485 (2 downloads); M48486 (2 downloads); M48487 (2 downloads); M48488 (2 downloads); M46418 (2 downloads); M46440 (2 downloads); M46820 (2 downloads); M149138 (2 downloads); M149143 (2 downloads); M107819 (2 downloads); M46187 (1 download); M46438 (1 download); M46907 (1 download); M46953 (1 download); M47789 (1 download); M47884 (1 download); M47978 (1 download); M149134 (1 download); M149135 (1 download); M149136 (1 download); M149137 (1 download); M149139 (2 downloads); M149140 (1 download); M149141 (2 downloads); M149142 (1 download); M149144 (1 download); M46117 (1 download); M46178 (1 download); M46179 (1 download); M46180 (1 download); M46446 (1 download); M46471 (1 download); M46472 (1 download); M46812 (1 download); M46815 (1 download); M46950 (1 download); M46986 (1 download); M47034 (1 download); M47875 (1 download); M48048 (1 download); M148176 (1 download); M151746 (1 download); M151747 (1 download); M148792 (1 download); M148793 (1 download); M148794 (1 download); M148796 (1 download); M148797 (1 download); M148799 (1 download); M148800 (1 download); M148801 (2 downloads); M148802 (1 download); M148803 (1 download); M148804 (1 download); M148805 (1 download); M148806 (1 download); M148807 (1 download); M148808 (1 download); M148809 (1 download); M148810 (1 download); M148811 (1 download); M46139 (1 download); M46943 (1 download); M46952 (1 download); M46992 (1 download); M47033 (1 download); M47793 (1 download); M47880 (1 download); M47963 (1 download); M46134 (1 download); M46135 (1 download); M46176 (1 download); M46177 (1 download); M46184 (1 download); M46444 (1 download); M46809 (1 download); M46810 (1 download); M46817 (1 download); M46843 (1 download); M46844 (1 download); M46976 (1 download); M46989 (1 download); M47032 (1 download); M47876 (1 download); M47975 (1 download); M98192 (1 download); M98193 (1 download); M98194 (1 download); M98195 (1 download); M98196 (1 download); M98197 (1 download); M98198 (1 download); M98201 (1 download); M98202 (1 download); M98203 (1 download); M98204 (1 download); M98206 (1 download); M98207 (1 download); M98208 (1 download); M98209 (1 download); M98210 (1 download); M98211 (1 download); M98212 (1 download); M98213 (1 download); M98214 (1 download); M98215 (1 download); M98216 (1 download); M98217 (1 download); M98218 (1 download); M98219 (1 download); M149298 (1 download); M46185 (1 download); M46186 (1 download); M46443 (1 download); M46449 (1 download); M46475 (1 download); M46476 (1 download); M46814 (1 download); M46842 (1 download); M46904 (1 download); M46988 (1 download); M47882 (1 download); M47883 (1 download); M47950 (1 download); M48055 (1 download); M46148 (1 download); M46151 (1 download); M46188 (1 download); M46451 (1 download); M46452 (1 download); M46823 (1 download); M46845 (1 download); M46910 (1 download); M46994 (1 download); M149299 (1 download); M46144 (1 download); M46145 (1 download); M46146 (1 download); M46147 (1 download); M46437 (1 download); M46822 (1 download); M46908 (1 download); M46909 (1 download); M46978 (1 download); M46993 (1 download); M46152 (2 downloads); M46153 (1 download); M46154 (1 download); M46155 (1 download); M46166 (1 download); M46167 (1 download); M46189 (1 download); M46414 (1 download); M46434 (1 download); M46435 (1 download); M46436 (1 download); M46824 (1 download); M46825 (1 download); M46846 (1 download); M46847 (1 download); M46848 (1 download); M46911 (2 downloads); M46912 (1 download); M46913 (1 download); M46914 (1 download); M46915 (1 download); M46955 (1 download); M46956 (1 download); M46974 (1 download); M46995 (1 download); M46996 (1 download); M46997 (1 download); M46998 (1 download); M47794 (1 download); M47795 (1 download); M47885 (1 download); M47986 (1 download); M48029 (1 download); M48049 (1 download); M48058 (1 download); M48059 (1 download); M48060 (1 download); M48061 (1 download); M48062 (1 download); M107818 (1 download); M107820 (1 download); M107821 (1 download); M107822 (1 download); M107823 (1 download); M142314 (1 download); M142315 (1 download); M142316 (1 download); M142317 (1 download); M142318 (1 download); M142319 (1 download); M142320 (1 download); M142321 (1 download); M142322 (1 download); M149197 (1 download); M149208 (1 download); M149209 (1 download); M149210 (1 download); M149211 (1 download); M149212 (1 download); M149213 (1 download); M149214 (1 download); M286139 (1 download); M286140 (1 download); M286141 (1 download); M286142 (1 download); M286143 (1 download); M286144 (1 download); M286145 (1 download); M286146 (1 download); M286147 (1 download); M286148 (1 download); M286149 (1 download); M286150 (1 download); M286151 (1 download); M286152 (1 download); M286153 (1 download); M286154 (1 download); M286155 (1 download); M286156 (1 download); M46191 (1 download); M46361 (1 download); M46479 (1 download); M46480 (1 download); M46849 (1 download); M47774 (1 download); M47791 (1 download); M47797 (1 download); M47914 (1 download); M47915 (1 download); M48047 (1 download); M48056 (1 download); M48516 (1 download); M46192 (1 download); M46193 (1 download); M46194 (1 download); M46195 (1 download); M46197 (1 download); M46198 (1 download); M46199 (1 download); M46200 (1 download); M46202 (1 download); M46362 (1 download); M46412 (1 download); M46415 (1 download); M46430 (1 download); M46431 (1 download); M46450 (1 download); M46828 (1 download); M46829 (1 download); M46917 (1 download); M46946 (1 download); M46958 (1 download); M46959 (1 download); M46960 (1 download); M46983 (1 download); M47002 (1 download); M47003 (1 download); M47004 (1 download); M47005 (1 download); M47031 (1 download); M47775 (1 download); M47792 (1 download); M47858 (1 download); M47887 (1 download); M47888 (1 download); M47965 (1 download); M47974 (1 download); M48497 (1 download); M48498 (1 download); M48499 (1 download); M48500 (1 download); M48501 (1 download); M48502 (1 download); M48503 (1 download); M48504 (1 download); M48505 (1 download); M48506 (1 download); M48507 (1 download); M48508 (1 download); M48509 (1 download); M48510 (1 download); M48511 (1 download); M48512 (1 download); M48513 (1 download); M46203 (1 download); M46204 (1 download); M46205 (1 download); M46206 (1 download); M46207 (1 download); M46208 (1 download); M46209 (1 download); M46210 (1 download); M46211 (1 download); M46212 (1 download); M46403 (1 download); M46407 (1 download); M46424 (1 download); M46429 (1 download); M46484 (1 download); M46485 (1 download); M46830 (1 download); M46850 (1 download); M46851 (1 download); M46852 (1 download); M46918 (1 download); M46919 (1 download); M46920 (1 download); M46921 (1 download); M46961 (1 download); M47006 (1 download); M47007 (1 download); M47009 (1 download); M47010 (1 download); M47029 (1 download); M47889 (1 download); M47890 (1 download); M47954 (1 download); 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