Project 352: E. V. Bärmann. Towards a comprehensive phylogeny of Bovidae, Chapter 2: A total evidence analysis of bovid phylogeny - Dissertation by Eva Bärmann, University of Cambridge UK.
Specimen: Raphicerus melanotis (ZMB/Mammals:2125)

Abstract

Combined-data analyses, also termed total evidence analyses, have proven to be beneficial for phylogeny reconstructions for a range of taxa. For Bovidae, the most diverse clade of ruminants (Artiodactyla, Mammalia), such an analysis was last performed a decade ago. Since then, the range of possible analysis techniques has expanded, now including model-based Bayesian inference that can also be used to incorporate non-molecular data. Furthermore, the possibility of making morphological data widely accessible online, providing detailed information on character states via photographs, facilitates the use and re-use of this type of data. This study presents a data set of 88 cranial, 19 soft-tissue, and 27 behavioural characters, sampled from 58 bovid species. These data are analysed in a simultaneous analysis with molecular data from nine genes (four mitochondrial and five nuclear markers), sampled for 57 bovid species. A few morphological and behavioural characters are identified as apomorphies of bovid taxa. Despite the fact that the majority of characters show homoplasies and cannot unambiguously be optimized on the phylogenetic tree, they allow for inferring the phylogenetic position of two fossil bovid species: †Turcoceros grangeri and †Gazella lydekkeri. The study is designed as a starting point for further combined data analyses of bovids or ruminants in general.


Read the article »

Project DOI: 10.7934/P352, http://dx.doi.org/10.7934/P352
This project containsMatrices
  • 806 Media
  • 3 Matrices
  • 86 Taxa
  • 212 Specimens
  • 134 Characters
Total size of project's media files: 1468 MB

Download Project SDD File
Total scored cells: 8111
Total media associated with cells: 6
Total labels associated with cell media: 0
Characters
Total characters: 134
Total characters with associated media: 10
Total characters with media with labels: 0
Total character states: 332
Total character states with associated media: 170
Total character states with media with labels:122
Total unordered/ordered characters:134/0
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MorphoBank Project 352


    Members

    member name taxa specimens media chars character
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    notes  medialabels
    rules
    Eva Baermann
    Project Administrator
    Last logged in 05/08/13
    86212806134112801588111
    (7554, 429, 128)
    443600
    Robert Asher
    Full membership
    Last logged in 05/18/10
    00000000
    (0, 0, 0)
    0000


    Taxonomic Overview for Matrix 'M817' (61 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    0878620002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    0898500002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    0847931002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    0878511002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    0878710002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    0888700002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0838032002
    [8] Capra falconeri
    Last Modified in 12/05/13
    1848231002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0868111002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    0847731002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    0817752002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    0878601002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    0878501002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    0878311002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    0878401002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    0878601002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1828151002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    0878501002
    [19] Gazella gazella
    Last Modified in 12/05/13
    1838131002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0878401002
    [21] Gazella marica
    Last Modified in 12/05/13
    0808071002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0848131002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    0878301002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    0868411002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    0898501002
    [26] Kobus kob
    Last Modified in 12/05/13
    0827761002
    [27] Kobus leche
    Last Modified in 12/05/13
    0837741002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    0797881002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    0838241002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    0888500002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    0858521002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    0868511002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    0868611002
    [34] Nanger dama
    Last Modified in 12/05/13
    0878301002
    [35] Nanger granti
    Last Modified in 12/05/13
    0878201002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    0888301002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0858412002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    0878701002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    0878302002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    0898601002
    [41] Oryx beisa
    Last Modified in 12/05/13
    1868221002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    0878511202
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0848331002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    0878601002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    0908101002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0878301002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    0837841002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    0818061002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    0878402002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0868602002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0828152002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    0868221002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    0878321202
    [54] Redunca redunca
    Last Modified in 12/05/13
    0817861002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    0888112002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0878301002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    0898301202
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0847432002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    0878701002
    [60] † Gazella lydekkeri
    Last Modified in 12/05/13
    0680191002
    [61] † Turcoceros grangeri
    Last Modified in 12/05/13
    1691180002


    Taxonomic Overview for Matrix 'M848' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    022523002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    027100002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    026401002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    025020002
    [5] Antidorcas marsupialis
    Last Modified in 12/05/13
    027000002
    [6] Antilope cervicapra
    Last Modified in 12/05/13
    026010002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    026711002
    [8] Capra falconeri
    Last Modified in 12/05/13
    022023002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    0221241002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    024921002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    029401002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    024221002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    017082002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018090002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    027000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    081190002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    1161100002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    023140002
    [19] Gazella gazella
    Last Modified in 12/05/13
    027100002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    0161110002
    [21] Gazella marica
    Last Modified in 12/05/13
    025720002
    [22] Gazella spekei
    Last Modified in 12/05/13
    0131140002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    025120002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    026101002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    023004002
    [26] Kobus kob
    Last Modified in 12/05/13
    027402002
    [27] Kobus leche
    Last Modified in 12/05/13
    026304002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    018564002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    617504002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    025020002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    025020002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    027100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    026110002
    [34] Nanger dama
    Last Modified in 12/05/13
    019080002
    [35] Nanger granti
    Last Modified in 12/05/13
    026010002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    019080002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    0100161002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    025220002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    080190002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    025211002
    [41] Oryx beisa
    Last Modified in 12/05/13
    026603002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    025120002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    0140112002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    024803002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    026701002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    0100170002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    020561002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    018581002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    025020002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    0120150002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    0100170002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    024203002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    024003002
    [54] Redunca redunca
    Last Modified in 12/05/13
    024303002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019153002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    0152111002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    026101002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0143121002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    022123002


    Taxonomic Overview for Matrix 'M849' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Addax nasomaculatus
    Last Modified in 12/05/13
    019800002
    [2] Aepyceros melampus
    Last Modified in 12/05/13
    019000002
    [3] Alcelaphus buselaphus
    Last Modified in 12/05/13
    019900002
    [4] Ammodorcas clarkei
    Last Modified in 12/05/13
    018510002
    [5] Antilope cervicapra
    Last Modified in 12/05/13
    017520002
    [6] Antidorcas marsupialis
    Last Modified in 12/05/13
    019000002
    [7] Boselaphus tragocamelus
    Last Modified in 12/05/13
    0191200002
    [8] Capra falconeri
    Last Modified in 12/05/13
    018010002
    [9] Cephalophus rufilatus
    Last Modified in 12/05/13
    118600002
    [10] Cephalophus silvicultor
    Last Modified in 12/05/13
    019600002
    [11] Connochaetes taurinus
    Last Modified in 12/05/13
    019700002
    [12] Damaliscus pygargus
    Last Modified in 12/05/13
    019000002
    [13] Dorcatragus megalotis
    Last Modified in 12/05/13
    019000002
    [14] Eudorcas rufifrons
    Last Modified in 12/05/13
    018610002
    [15] Eudorcas thomsoni
    Last Modified in 12/05/13
    019000002
    [16] Gazella bennettii
    Last Modified in 12/05/13
    017520002
    [17] Gazella cuvieri
    Last Modified in 12/05/13
    018510002
    [18] Gazella dorcas
    Last Modified in 12/05/13
    018510002
    [19] Gazella gazella
    Last Modified in 12/05/13
    019600002
    [20] Gazella leptoceros
    Last Modified in 12/05/13
    017520002
    [21] Gazella marica
    Last Modified in 12/05/13
    019800002
    [22] Gazella spekei
    Last Modified in 12/05/13
    018510002
    [23] Gazella subgutturosa
    Last Modified in 12/05/13
    019600002
    [24] Hippotragus niger
    Last Modified in 12/05/13
    019000002
    [25] Kobus ellipsiprymnus
    Last Modified in 12/05/13
    019000002
    [26] Kobus kob
    Last Modified in 12/05/13
    019500002
    [27] Kobus leche
    Last Modified in 12/05/13
    019500002
    [28] Kobus megaceros
    Last Modified in 12/05/13
    019500002
    [29] Kobus vardoni
    Last Modified in 12/05/13
    019600002
    [30] Litocranius walleri
    Last Modified in 12/05/13
    019000002
    [31] Madoqua guentheri
    Last Modified in 12/05/13
    019000002
    [32] Madoqua kirki
    Last Modified in 12/05/13
    019100002
    [33] Madoqua saltiana
    Last Modified in 12/05/13
    019000002
    [34] Nanger dama
    Last Modified in 12/05/13
    018510002
    [35] Nanger granti
    Last Modified in 12/05/13
    019000002
    [36] Nanger soemmerringi
    Last Modified in 12/05/13
    018510002
    [37] Neotragus batesi
    Last Modified in 12/05/13
    018210002
    [38] Neotragus moschatus
    Last Modified in 12/05/13
    018010002
    [39] Neotragus pygmaeus
    Last Modified in 12/05/13
    018410002
    [40] Oreotragus oreotragus
    Last Modified in 12/05/13
    019000002
    [41] Oryx beisa
    Last Modified in 12/05/13
    019900002
    [42] Ourebia ourebi
    Last Modified in 12/05/13
    019000002
    [43] Pantholops hodgsoni
    Last Modified in 12/05/13
    019000002
    [44] Pelea capreolus
    Last Modified in 12/05/13
    019400002
    [45] Philantomba monticola
    Last Modified in 12/05/13
    019600002
    [46] Procapra gutturosa
    Last Modified in 12/05/13
    019400002
    [47] Procapra picticaudata
    Last Modified in 12/05/13
    019300002
    [48] Procapra przewalskii
    Last Modified in 12/05/13
    019400002
    [49] Raphicerus campestris
    Last Modified in 12/05/13
    019000002
    [50] Raphicerus melanotis
    Last Modified in 12/05/13
    018010002
    [51] Raphicerus sharpei
    Last Modified in 12/05/13
    019000002
    [52] Redunca arundinum
    Last Modified in 12/05/13
    019500002
    [53] Redunca fulvorufula
    Last Modified in 12/05/13
    019000002
    [54] Redunca redunca
    Last Modified in 12/05/13
    019500002
    [55] Rupicapra rupicapra
    Last Modified in 12/05/13
    019700002
    [56] Saiga tatarica
    Last Modified in 12/05/13
    018010002
    [57] Sylvicapra grimmia
    Last Modified in 12/05/13
    019000002
    [58] Tetracerus quadricornis
    Last Modified in 12/05/13
    0191300002
    [59] Tragelaphus imberbis
    Last Modified in 12/05/13
    019000002


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    Total project views187939
    Project overview3719
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M107807 (151 views); M107817 (218 views); M107818 (209 views); M107821 (156 views); M142302 (127 views); M142304 (168 views); M142305 (173 views); M142307 (223 views); M142308 (141 views); M142309 (202 views); M148781 (208 views); M149147 (48 views); M148170 (131 views); M148782 (133 views); M148795 (144 views); M284576 (5 views); M149111 (52 views); M148171 (123 views); M46500 (196 views); M46842 (144 views); M148784 (140 views); M46495 (112 views); M46825 (174 views); M148785 (135 views); M148780 (141 views); M149225 (50 views); M98198 (4 views); M148799 (142 views); M148794 (139 views); M148152 (177 views); M148778 (184 views); M46343 (344 views); M46187 (335 views); M46167 (341 views); M46172 (393 views); M148173 (179 views); M46179 (372 views); M148820 (49 views); M148175 (158 views); M148788 (136 views); M149219 (48 views); M148814 (55 views); M149230 (37 views); M149193 (57 views); M148800 (135 views); M46335 (395 views); M148818 (44 views); M46148 (344 views); M46155 (394 views); M149170 (50 views); M107804 (141 views); M148169 (128 views); M98201 (7 views); M149208 (68 views); M148790 (180 views); M149145 (50 views); M148166 (134 views); M149141 (81 views); M149292 (27 views); M149259 (6 views); M148813 (50 views); M149150 (49 views); M148826 (46 views); M149122 (48 views); M148823 (64 views); M148798 (210 views); M149211 (51 views); M148827 (48 views); M149174 (41 views); M149160 (42 views); M148824 (45 views); M149135 (63 views); M148779 (170 views); M47883 (5 views); M149149 (43 views); M148812 (41 views); M149168 (47 views); M149192 (54 views); M149137 (41 views); M149189 (63 views); M149180 (54 views); M46348 (364 views); M149213 (46 views); M148803 (43 views); M149275 (5 views); M46852 (169 views); M47020 (5 views); M149253 (6 views); M149118 (49 views); M148805 (45 views); M149194 (48 views); M149143 (52 views); M148811 (67 views); M149223 (39 views); M149222 (48 views); M149159 (44 views); M149172 (44 views); M148801 (44 views); M148802 (46 views); M148804 (46 views); M148806 (41 views); M148807 (65 views); M148808 (88 views); M148809 (48 views); M148810 (41 views); M148815 (50 views); M148816 (69 views); M148817 (53 views); M148819 (41 views); M148821 (43 views); M148825 (52 views); M149110 (64 views); M149112 (41 views); M149113 (41 views); M149114 (50 views); M149115 (47 views); M149116 (45 views); M149117 (53 views); M149119 (51 views); M149123 (52 views); M149134 (86 views); M149136 (43 views); M149138 (42 views); M149139 (44 views); M149140 (49 views); M149142 (55 views); M149144 (48 views); M149146 (44 views); M149148 (40 views); M149151 (43 views); M149152 (54 views); M149153 (61 views); M149154 (52 views); M149155 (41 views); M149156 (43 views); M149157 (44 views); M149158 (53 views); M149161 (51 views); M149162 (41 views); M149163 (49 views); M149164 (45 views); M149165 (51 views); M149166 (49 views); M149167 (43 views); M149169 (41 views); M149173 (47 views); M149175 (49 views); M149176 (62 views); M149177 (64 views); M149179 (43 views); M149181 (60 views); M149182 (47 views); M149183 (53 views); M149184 (50 views); M149185 (44 views); M149187 (57 views); M149188 (69 views); M149190 (57 views); M149191 (95 views); M149196 (50 views); M149197 (60 views); M149198 (45 views); M149209 (49 views); M149210 (50 views); M149212 (44 views); M149214 (45 views); M149218 (46 views); M149220 (45 views); M149221 (38 views); M149224 (40 views); M149226 (41 views); M149227 (38 views); M149228 (45 views); M149229 (39 views); M149242 (8 views); M46411 (170 views); M46193 (345 views); M46333 (335 views); M46449 (131 views); M286147 (5 views); M46405 (149 views); M149233 (5 views); M48516 (6 views); M46824 (170 views); M47787 (4 views); M46919 (5 views); M46431 (135 views); M46905 (6 views); M47950 (5 views); M47788 (6 views); M47955 (6 views); M46188 (338 views); M46202 (344 views); M149289 (7 views); M47887 (4 views); M149282 (4 views); M46326 (330 views); M46421 (128 views); M46415 (169 views); M46453 (121 views); M46153 (366 views); M46904 (5 views); M46843 (174 views); M149286 (5 views); M151747 (4 views); M46851 (144 views); M46118 (333 views); M46127 (353 views); M46128 (347 views); M46134 (346 views); M46135 (352 views); M46144 (350 views); M46145 (364 views); M46146 (378 views); M46147 (367 views); M46151 (336 views); M46152 (347 views); M46154 (352 views); M46166 (346 views); M46169 (328 views); M46171 (404 views); M46176 (366 views); M46177 (349 views); M46178 (375 views); M46180 (344 views); M46184 (331 views); M46185 (335 views); M46186 (336 views); M46189 (355 views); M46190 (379 views); M46191 (342 views); M46192 (364 views); M46194 (321 views); M46195 (351 views); M46197 (340 views); M46198 (328 views); M46199 (352 views); M46200 (344 views); M46204 (328 views); M46205 (314 views); M46206 (333 views); M46207 (365 views); M46208 (318 views); M46210 (339 views); M46211 (335 views); M46212 (389 views); M46230 (388 views); M46327 (353 views); M46328 (420 views); M46329 (349 views); M46330 (354 views); M46331 (363 views); M46334 (382 views); M46337 (355 views); M46340 (350 views); M46341 (342 views); M46342 (346 views); M46344 (366 views); M46345 (333 views); M46350 (347 views); M46351 (315 views); M46352 (358 views); M46353 (353 views); M46354 (362 views); M46355 (368 views); M46356 (324 views); M46357 (367 views); M46358 (346 views); M46359 (356 views); M46361 (348 views); M46362 (353 views); M46403 (317 views); M46406 (172 views); M46407 (181 views); M46412 (161 views); M46413 (133 views); M46414 (135 views); M46416 (152 views); M46417 (123 views); M46418 (174 views); M46419 (176 views); M46420 (134 views); M46422 (122 views); M46423 (122 views); M46424 (161 views); M46425 (170 views); M46426 (114 views); M46427 (155 views); M46428 (136 views); M46429 (121 views); M46430 (151 views); M46432 (126 views); M46434 (152 views); M46436 (128 views); M46437 (136 views); M46438 (155 views); M46440 (111 views); M46441 (138 views); M46442 (162 views); M46443 (165 views); M46444 (174 views); M46446 (135 views); M46447 (103 views); M46448 (129 views); M46450 (123 views); M46451 (150 views); M46452 (168 views); M46454 (174 views); M46455 (154 views); M46456 (124 views); M46457 (124 views); M46458 (151 views); M46459 (129 views); M46471 (155 views); M46472 (129 views); M46475 (125 views); M46476 (173 views); M46479 (168 views); M46480 (149 views); M46481 (185 views); M46482 (113 views); M46483 (144 views); M46484 (150 views); M46485 (128 views); M46486 (122 views); M46487 (160 views); M46488 (139 views); M46489 (137 views); M46490 (129 views); M46492 (142 views); M46493 (129 views); M46494 (153 views); M46496 (126 views); M46497 (130 views); M46498 (153 views); M46499 (155 views); M46809 (116 views); M46810 (123 views); M46812 (119 views); M46814 (138 views); M46815 (122 views); M46817 (129 views); M46818 (155 views); M46819 (169 views); M46820 (141 views); M46822 (156 views); M46823 (161 views); M46827 (120 views); M46828 (151 views); M46829 (121 views); M46830 (138 views); M46831 (116 views); M46832 (140 views); M46833 (120 views); M46834 (131 views); M46835 (171 views); M46837 (140 views); M46838 (136 views); M46839 (108 views); M46840 (136 views); M46841 (116 views); M46844 (152 views); M46845 (117 views); M46846 (110 views); M46847 (118 views); M46848 (106 views); M46849 (130 views); M46850 (144 views); M46853 (149 views); M46854 (104 views); M46855 (136 views); M46856 (109 views); M46857 (134 views); M46858 (120 views); M46859 (122 views); M46860 (163 views); M46861 (152 views); M46862 (148 views); M46863 (116 views); M46864 (115 views); M284578 (8 views); M149294 (5 views); M149243 (4 views); M149268 (4 views); M149271 (6 views); M46946 (5 views); M47791 (8 views); M98218 (4 views); M149285 (4 views); M284582 (4 views); M46950 (5 views); M98192 (5 views); M284577 (7 views); M46986 (7 views); M286153 (6 views); M149231 (7 views); M286145 (4 views); M286141 (4 views); M149248 (6 views); M149291 (5 views); M47018 (6 views); M48485 (6 views); M47022 (5 views); M46995 (5 views); M47794 (4 views); M47030 (4 views); M47009 (4 views); M47948 (4 views); M47786 (4 views); M46998 (4 views); M47007 (4 views); M47779 (4 views); M47954 (4 views); M47785 (4 views); M47965 (7 views); M47899 (6 views); M47986 (4 views); M47778 (4 views); M47790 (4 views); M48056 (6 views); M47976 (4 views); M48488 (5 views); M48058 (4 views); M48029 (4 views); M47970 (5 views); M46988 (4 views); M98208 (5 views); M47892 (4 views); M47781 (5 views); M47953 (7 views); M48509 (4 views); M98204 (4 views); M47011 (4 views); M47915 (25 views); M47014 (4 views); M47010 (5 views); M47797 (5 views); M48512 (5 views); M47016 (4 views); M47905 (5 views); M47906 (4 views); M47886 (5 views); M98212 (5 views); M98213 (5 views); M47795 (4 views); M98196 (4 views); M47951 (4 views); M47978 (6 views); M48047 (45 views); M48048 (4 views); M48487 (8 views); M48059 (4 views); M98195 (6 views); M47003 (5 views); M48503 (5 views); M47964 (5 views); M48514 (4 views); M46921 (6 views); M51943 (7 views); M47917 (5 views); M286140 (4 views); M48062 (5 views); M98209 (5 views); M149260 (8 views); M149297 (7 views); M46957 (10 views); M47017 (8 views); M149283 (4 views); M149281 (3 views); M149264 (4 views); M149288 (5 views); M46966 (7 views); M149279 (4 views); M47796 (5 views); M47789 (4 views); M98215 (4 views); M47884 (5 views); M149290 (4 views); M48053 (8 views); M48511 (4 views); M46923 (6 views); M47903 (4 views); M46939 (6 views); M47890 (4 views); M46926 (7 views); M47977 (6 views); M46952 (8 views); M47031 (5 views); M46924 (6 views); M46960 (7 views); M47775 (4 views); M48508 (4 views); M48499 (5 views); M46969 (7 views); M46929 (6 views); M46976 (20 views); M46964 (6 views); M98203 (4 views); M46967 (7 views); M46937 (7 views); M46972 (8 views); M47782 (4 views); M46971 (6 views); M46981 (7 views); M98217 (4 views); M46982 (7 views); M46955 (6 views); M98214 (4 views); M46959 (9 views); M47005 (4 views); M47858 (4 views); M48507 (4 views); M46951 (7 views); M46910 (6 views); M48497 (7 views); M46961 (6 views); M47981 (5 views); M46956 (8 views); M46979 (6 views); M47885 (4 views); M46968 (8 views); M46965 (8 views); M48057 (5 views); M46940 (6 views); M46934 (6 views); M46945 (6 views); M46980 (11 views); M46973 (6 views); M46978 (6 views); M46928 (8 views); M46970 (6 views); M46920 (7 views); M46975 (9 views); M48502 (4 views); M46953 (6 views); M98197 (5 views); M46962 (6 views); M46984 (6 views); M98210 (5 views); M46911 (7 views); M46925 (6 views); M48049 (5 views); M46917 (7 views); M47975 (5 views); M46932 (6 views); M46909 (6 views); M48515 (5 views); M47877 (5 views); M98194 (4 views); M48500 (4 views); M47949 (6 views); M149246 (5 views); M98207 (5 views); M47006 (4 views); M149235 (4 views); M47904 (4 views); M47889 (4 views); M98219 (4 views); M286142 (6 views); M46913 (6 views); M48505 (5 views); M47888 (4 views); M47776 (4 views); M46915 (5 views); M47974 (4 views); M149295 (4 views); M47780 (4 views); M284579 (5 views); M149269 (4 views); M149239 (6 views); M47792 (4 views); M47777 (4 views); M47021 (4 views); M46914 (5 views); M47019 (4 views); M47004 (4 views); M47012 (4 views); M47916 (5 views); M47871 (4 views); M98211 (5 views); M48061 (4 views); M98202 (5 views); M48510 (7 views); M48506 (5 views); M48055 (4 views); M151745 (4 views); M149267 (4 views); M149252 (4 views); M149255 (5 views); M149298 (4 views); M46943 (5 views); M286149 (4 views); M286154 (6 views); M46927 (5 views); M149238 (4 views); M149266 (4 views); M149244 (5 views); M149280 (4 views); M286143 (4 views); M286139 (4 views); M286146 (4 views); M286152 (7 views); M149250 (4 views); M149274 (4 views); M149299 (4 views); M149277 (4 views); M286155 (6 views); M286156 (6 views); M149287 (5 views); M149241 (4 views); M151743 (7 views); M149232 (4 views); M286150 (4 views); M46941 (6 views); M47882 (4 views); M47963 (4 views); M46907 (5 views); M46963 (7 views); M46912 (6 views); M48486 (5 views); M48050 (6 views); M47952 (3 views); M46935 (6 views); M46918 (4 views); M48054 (3 views); M48498 (3 views); M46938 (4 views); M48051 (3 views); M284581 (4 views); M149237 (3 views); M149296 (3 views); M149293 (3 views); M284584 (3 views); M149234 (4 views); M284580 (4 views); M149258 (8 views); M149256 (3 views); M149263 (3 views); M149240 (3 views); M149257 (4 views); M149236 (5 views); M46991 (3 views); M46989 (3 views); M47783 (3 views); M46922 (7 views); M47784 (3 views); M149278 (3 views); M46865 (6 views); M47034 (3 views); M46993 (3 views); M47774 (3 views); M46983 (5 views); M47874 (3 views); M46908 (5 views); M46996 (3 views); M286148 (3 views); M47896 (3 views); M149245 (3 views); M47914 (29 views); M286144 (3 views); M286151 (4 views); M48501 (3 views); M47956 (6 views); M149247 (3 views); M149254 (3 views); M149249 (3 views); M46997 (5 views); M151744 (3 views); M151746 (3 views); M149270 (3 views); M98193 (33 views); M46994 (3 views); M51944 (5 views); M47013 (4 views); M48052 (4 views); M149273 (3 views); M47895 (3 views); M47033 (3 views); M149265 (3 views); M46977 (8 views); M47015 (3 views); M46992 (4 views); M149261 (4 views); M47793 (3 views); M48513 (3 views); M149276 (3 views); M149284 (3 views); M98206 (5 views); M46958 (6 views); M47875 (4 views); M47897 (3 views); M149272 (3 views); M46936 (6 views); M46974 (5 views); M46947 (8 views); M47001 (3 views); M149251 (3 views); M47032 (4 views); M47876 (3 views); M47002 (3 views); M47029 (3 views); M47859 (3 views); M48060 (3 views); M47880 (3 views); M47973 (3 views); M46930 (6 views); M47891 (3 views); M48504 (3 views); M284583 (6 views); M98216 (4 views); M46933 (5 views);
    Bibliography9930
    Taxon list9832
    Specimen list78881




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project1095
    Project downloads213
    Matrix downloads61Skull characters Bovidae (44 downloads); Behavior (8 downloads); Glands and Colouration (9 downloads);
    Media downloads821M142313 (2 downloads); M142296 (1 download); M148795 (4 downloads); M148798 (3 downloads); M46337 (2 downloads); M46421 (2 downloads); M46492 (1 download); M47899 (2 downloads); M48485 (2 downloads); M48486 (2 downloads); M48487 (2 downloads); M48488 (2 downloads); M46418 (2 downloads); M46440 (2 downloads); M46820 (2 downloads); M149138 (2 downloads); M149143 (2 downloads); M107819 (2 downloads); M46187 (1 download); M46438 (1 download); M46907 (1 download); M46953 (1 download); M47789 (1 download); M47884 (1 download); M47978 (1 download); M149134 (1 download); M149135 (1 download); M149136 (1 download); M149137 (1 download); M149139 (2 downloads); M149140 (1 download); M149141 (2 downloads); M149142 (1 download); M149144 (1 download); M46117 (1 download); M46178 (1 download); M46179 (1 download); M46180 (1 download); M46446 (1 download); M46471 (1 download); M46472 (1 download); M46812 (1 download); M46815 (1 download); M46950 (1 download); M46986 (1 download); M47034 (1 download); M47875 (1 download); M48048 (1 download); M148176 (1 download); M151746 (1 download); M151747 (1 download); M148792 (1 download); M148793 (1 download); M148794 (1 download); M148796 (1 download); M148797 (1 download); M148799 (1 download); M148800 (1 download); M148801 (2 downloads); M148802 (1 download); M148803 (1 download); M148804 (1 download); M148805 (1 download); M148806 (1 download); M148807 (1 download); M148808 (1 download); M148809 (1 download); M148810 (1 download); M148811 (1 download); M46139 (1 download); M46943 (1 download); M46952 (1 download); M46992 (1 download); M47033 (1 download); M47793 (1 download); M47880 (1 download); M47963 (1 download); M46134 (1 download); M46135 (1 download); M46176 (1 download); M46177 (1 download); M46184 (1 download); M46444 (1 download); M46809 (1 download); M46810 (1 download); M46817 (1 download); M46843 (1 download); M46844 (1 download); M46976 (1 download); M46989 (1 download); M47032 (1 download); M47876 (1 download); M47975 (1 download); M98192 (1 download); M98193 (1 download); M98194 (1 download); M98195 (1 download); M98196 (1 download); M98197 (1 download); M98198 (1 download); M98201 (1 download); M98202 (1 download); M98203 (1 download); M98204 (1 download); M98206 (1 download); M98207 (1 download); M98208 (1 download); M98209 (1 download); M98210 (1 download); M98211 (1 download); M98212 (1 download); M98213 (1 download); M98214 (1 download); M98215 (1 download); M98216 (1 download); M98217 (1 download); M98218 (1 download); M98219 (1 download); M149298 (1 download); M46185 (1 download); M46186 (1 download); M46443 (1 download); M46449 (1 download); M46475 (1 download); M46476 (1 download); M46814 (1 download); M46842 (1 download); M46904 (1 download); M46988 (1 download); M47882 (1 download); M47883 (1 download); M47950 (1 download); M48055 (1 download); M46148 (1 download); M46151 (1 download); M46188 (1 download); M46451 (1 download); M46452 (1 download); M46823 (1 download); M46845 (1 download); M46910 (1 download); M46994 (1 download); M149299 (1 download); M46144 (1 download); M46145 (1 download); M46146 (1 download); M46147 (1 download); M46437 (1 download); M46822 (1 download); M46908 (1 download); M46909 (1 download); M46978 (1 download); M46993 (1 download); M46152 (2 downloads); M46153 (1 download); M46154 (1 download); M46155 (1 download); M46166 (1 download); M46167 (1 download); M46189 (1 download); M46414 (1 download); M46434 (1 download); M46435 (1 download); M46436 (1 download); M46824 (1 download); M46825 (1 download); M46846 (1 download); M46847 (1 download); M46848 (1 download); M46911 (2 downloads); M46912 (1 download); M46913 (1 download); M46914 (1 download); M46915 (1 download); M46955 (1 download); M46956 (1 download); M46974 (1 download); M46995 (1 download); M46996 (1 download); M46997 (1 download); M46998 (1 download); M47794 (1 download); M47795 (1 download); M47885 (1 download); M47986 (1 download); M48029 (1 download); M48049 (1 download); M48058 (1 download); M48059 (1 download); M48060 (1 download); M48061 (1 download); M48062 (1 download); M107818 (1 download); M107820 (1 download); M107821 (1 download); M107822 (1 download); M107823 (1 download); M142314 (1 download); M142315 (1 download); M142316 (1 download); M142317 (1 download); M142318 (1 download); M142319 (1 download); M142320 (1 download); M142321 (1 download); M142322 (1 download); M149197 (1 download); M149208 (1 download); M149209 (1 download); M149210 (1 download); M149211 (1 download); M149212 (1 download); M149213 (1 download); M149214 (1 download); M286139 (1 download); M286140 (1 download); M286141 (1 download); M286142 (1 download); M286143 (1 download); M286144 (1 download); M286145 (1 download); M286146 (1 download); M286147 (1 download); M286148 (1 download); M286149 (1 download); M286150 (1 download); M286151 (1 download); M286152 (1 download); M286153 (1 download); M286154 (1 download); M286155 (1 download); M286156 (1 download); M46191 (1 download); M46361 (1 download); M46479 (1 download); M46480 (1 download); M46849 (1 download); M47774 (1 download); M47791 (1 download); M47797 (1 download); M47914 (1 download); M47915 (1 download); M48047 (1 download); M48056 (1 download); M48516 (1 download); M46192 (1 download); M46193 (1 download); M46194 (1 download); M46195 (1 download); M46197 (1 download); M46198 (1 download); M46199 (1 download); M46200 (1 download); M46202 (1 download); M46362 (1 download); M46412 (1 download); M46415 (1 download); M46430 (1 download); M46431 (1 download); M46450 (1 download); M46828 (1 download); M46829 (1 download); M46917 (1 download); M46946 (1 download); M46958 (1 download); M46959 (1 download); M46960 (1 download); M46983 (1 download); M47002 (1 download); M47003 (1 download); M47004 (1 download); M47005 (1 download); M47031 (1 download); M47775 (1 download); M47792 (1 download); M47858 (1 download); M47887 (1 download); M47888 (1 download); M47965 (1 download); M47974 (1 download); M48497 (1 download); M48498 (1 download); M48499 (1 download); M48500 (1 download); M48501 (1 download); M48502 (1 download); M48503 (1 download); M48504 (1 download); M48505 (1 download); M48506 (1 download); M48507 (1 download); M48508 (1 download); M48509 (1 download); M48510 (1 download); M48511 (1 download); M48512 (1 download); M48513 (1 download); M46203 (1 download); M46204 (1 download); M46205 (1 download); M46206 (1 download); M46207 (1 download); M46208 (1 download); M46209 (1 download); M46210 (1 download); M46211 (1 download); M46212 (1 download); M46403 (1 download); M46407 (1 download); M46424 (1 download); M46429 (1 download); M46484 (1 download); M46485 (1 download); M46830 (1 download); M46850 (1 download); M46851 (1 download); M46852 (1 download); M46918 (1 download); M46919 (1 download); M46920 (1 download); M46921 (1 download); M46961 (1 download); M47006 (1 download); M47007 (1 download); M47009 (1 download); M47010 (1 download); M47029 (1 download); M47889 (1 download); M47890 (1 download); M47954 (1 download); 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