Morphobank

Project 3587: C. Olivier, B. Battail, S. Bourquin, C. Rossignol, J. Steyer, N. Jalil. 2019. New dicynodonts (Therapsida, Anomodontia) from near the Permo-Triassic boundary of Laos: implications for dicynodont survivorship across the Permo-Triassic mass extinction and the paleobiogeography of Southeast Asian blocks. Journal of Vertebrate Paleontology. 39 (2):e1584745.

Specimen: † Dicynodon lacerticeps (unvouchered)
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Abstract

The dicynodonts are an emblematic group of herbivorous therapsids that survived the Permo-Triassic (P-Tr) crisis. Laotian dicynodonts from stratigraphically constrained beds, recently dated using the U-Pb zircon method, yield new insights into terrestrial faunas of Southeast Asia during the latest Permian and earliest Triassic. Summarily described, they were originally attributed to the genus Dicynodon. We provide a new phylogenetic analysis for Laotian dicynodonts, based on three well-preserved skulls, indicating that they belong to two new taxa: Counillonia superoculis, gen. et sp. nov., and Repelinosaurus robustus, gen. et sp. nov. Our phylogenetic analysis of Dicynodontia indicates that (1) Counillonia is closely related to some ‘Dicynodon'-grade taxa and (2) Repelinosaurus is a kannemeyeriiform. The phylogenetic affinities of these new Laotian dicynodonts allow discussion of the survivorship of multiple lineages (Kannemeyeriiformes and ‘Dicynodon'grade dicynodontoids) across the P-Tr crisis. The Laotian dicynodonts also shed new light on the paleobiogeography of Southeast Asia from the late Paleozoic to the early Mesozoic, particularly the timing of collisions between the Indochina, the South China, and the North China blocks. The presence of dicynodonts in Laos, most likely in the Early Triassic, thus implies that the connection between the Indochina Block and the South China Block occurred no later than the latest Permian or earliest Triassic (i.e., when the dicynodonts provide direct evidence for a connection).

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Article DOI: 10.1080/02724634.2019.1584745

Project DOI: 10.7934/P3587, http://dx.doi.org/10.7934/P3587

This project contains	Matrices
1 Media 1 Matrix 2 Documents 106 Taxa 1 Specimen 194 Characters Total size of project's media files: 346.47k Download Project SDD File	Total scored cells: 14436 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 194 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 424 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:1/193

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MorphoBank Project 3587

Creation Date:
15 November 2019
Publication Date:
15 January 2020
Matrix downloads: 2

Authors' Institutions

Muséum national d'Histoire naturelle, Paris (Natural History Museum, Paris)

Centre National de la Recherche Scientifique (CNRS)

University of Rennes

Universidade de Sao Paulo (USP, University of Sao Paulo)

Sorbonne Université

Université Cadi Ayyad

Members

Ana Andruchow-Colombo
Project Administrator

106

194

14436
(12941, 0, 1495)

Chloe Olivier
Full membership

0
(0, 0, 0)

Taxonomic Overview for Matrix 'M26359' (106 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Biarmosuchus Taxon name last Modified on 01/15/20	33	150	150	0	11	0	0	2
[2] † Hipposaurus boonstrai Taxon name last Modified on 01/15/20	33	148	148	0	13	0	0	2
[3] † Archaeosyodon praeventor Taxon name last Modified on 01/15/20	85	106	106	0	3	0	0	2
[4] † Titanophoneus potens Taxon name last Modified on 01/15/20	22	159	158	0	14	0	0	2
[5] † Gorgonops torvus Taxon name last Modified on 01/15/20	25	161	161	0	8	0	0	2
[6] † Lycosuchus vanderrieti Taxon name last Modified on 01/15/20	27	153	153	0	14	0	0	2
[7] † Glanosuchus macrops Taxon name last Modified on 01/15/20	18	161	161	0	15	0	0	2
[8] † Biseridens qilianicus Taxon name last Modified on 01/15/20	75	115	115	0	4	0	0	2
[9] † Anomocephalus africanus Taxon name last Modified on 01/15/20	166	28	28	0	0	0	0	2
[10] † Tiarajudens eccentricus Taxon name last Modified on 01/15/20	139	53	53	0	2	0	0	2
[11] † Otsheria netzvetajevi Taxon name last Modified on 01/15/20	97	86	86	0	11	0	0	2
[12] † Ulemica Taxon name last Modified on 01/15/20	55	126	125	0	14	0	0	2
[13] † Suminia getmanovi Taxon name last Modified on 01/15/20	22	152	152	0	20	0	0	2
[14] † Patranomodon nyaphulii Taxon name last Modified on 01/15/20	51	129	129	0	14	0	0	2
[15] † Galeops whaitsi Taxon name last Modified on 01/15/20	63	117	117	0	14	0	0	2
[16] † Galepus jouberti Taxon name last Modified on 01/15/20	151	40	40	0	3	0	0	2
[17] † Galechirus scholtzi Taxon name last Modified on 01/15/20	132	57	57	0	5	0	0	2
[18] † Eodicynodon oelofseni Taxon name last Modified on 01/15/20	94	93	93	0	7	0	0	2
[19] † Eodicynodon oosthuizeni Taxon name last Modified on 01/15/20	6	166	166	0	22	0	0	2
[20] † Colobodectes cluveri Taxon name last Modified on 01/15/20	79	104	104	0	11	0	0	2
[21] † Lanthanostegus mohoii Taxon name last Modified on 01/15/20	133	59	59	0	2	0	0	2
[22] † Eosimops newtoni Taxon name last Modified on 01/15/20	18	158	156	0	20	0	0	2
[23] † Prosictodon dubei Taxon name last Modified on 01/15/20	53	126	126	0	15	0	0	2
[24] † Diictodon feliceps Taxon name last Modified on 01/15/20	8	167	164	0	22	0	0	2
[25] † Robertia broomiana Taxon name last Modified on 01/15/20	14	160	160	0	20	0	0	2
[26] † Pristerodon mackayi Taxon name last Modified on 01/15/20	7	165	165	0	22	0	0	2
[27] † Brachyprosopus broomi Taxon name last Modified on 01/15/20	23	152	152	0	19	0	0	2
[28] † Endothiodon bathystoma Taxon name last Modified on 01/15/20	15	160	157	0	22	0	0	2
[29] † Endothiodon tolani Taxon name last Modified on 01/15/20	61	124	123	0	10	0	0	2
[30] † Niassodon mfumukasi Taxon name last Modified on 01/15/20	54	130	130	0	10	0	0	2
[31] † Digalodon rubidgei Taxon name last Modified on 01/15/20	63	120	120	0	11	0	0	2
[32] † Emydops Taxon name last Modified on 01/15/20	8	165	164	0	22	0	0	2
[33] † Dicynodontoides Taxon name last Modified on 01/15/20	10	163	163	0	21	0	0	2
[34] † Kombuisia frerensis Taxon name last Modified on 01/15/20	65	119	119	0	10	0	0	2
[35] † Myosaurus gracilis Taxon name last Modified on 01/15/20	30	146	146	0	18	0	0	2
[36] † Sauroscaptor tharavati Taxon name last Modified on 01/15/20	91	94	94	0	9	0	0	2
[37] † Cistecephalus microrhinus Taxon name last Modified on 01/15/20	13	162	160	0	21	0	0	2
[38] † Cistecephaloides boonstrai Taxon name last Modified on 01/15/20	60	121	121	0	13	0	0	2
[39] † Kawingasaurus fossilis Taxon name last Modified on 01/15/20	42	136	136	0	16	0	0	2
[40] † Rhachiocephalus magnus Taxon name last Modified on 01/15/20	22	154	153	0	19	0	0	2
[41] † Kitchinganomodon crassus Taxon name last Modified on 01/15/20	43	134	134	0	17	0	0	2
[42] † Oudenodon bainii Taxon name last Modified on 01/15/20	6	166	165	0	23	0	0	2
[43] † Tropidostoma dubium Taxon name last Modified on 01/15/20	17	155	155	0	22	0	0	2
[44] † Australobarbarus Taxon name last Modified on 01/15/20	41	131	131	0	22	0	0	2
[45] † Odontocyclops whaitsi Taxon name last Modified on 01/15/20	21	150	150	0	23	0	0	2
[46] † Idelesaurus tataricus Taxon name last Modified on 01/15/20	46	132	132	0	16	0	0	2
[47] † Aulacephalodon bainii Taxon name last Modified on 01/15/20	6	170	165	0	23	0	0	2
[48] † Pelanomodon moschops Taxon name last Modified on 01/15/20	69	111	110	0	15	0	0	2
[49] † Geikia locusticeps Taxon name last Modified on 01/15/20	40	138	138	0	16	0	0	2
[50] † Geikia elginensis Taxon name last Modified on 01/15/20	88	96	96	0	10	0	0	2
[51] † Elph borealis Taxon name last Modified on 01/15/20	69	113	113	0	12	0	0	2
[52] † Interpresosaurus blomi Taxon name last Modified on 01/15/20	150	44	44	0	0	0	0	2
[53] † Katumbia parringtoni Taxon name last Modified on 01/15/20	60	120	120	0	14	0	0	2
[54] † Delectosaurus arefjevi Taxon name last Modified on 01/15/20	74	108	107	0	13	0	0	2
[55] † Dicynodon lacerticeps Taxon name last Modified on 01/15/20	42	137	136	0	16	0	0	2
[56] † Dicynodon huenei Taxon name last Modified on 01/15/20	15	157	156	0	23	0	0	2
[57] † Daptocephalus leoniceps Taxon name last Modified on 01/15/20	32	145	145	0	17	0	0	2
[58] † Daqingshanodon limbus Taxon name last Modified on 01/15/20	58	121	121	0	15	0	0	2
[59] † Dinanomodon gilli Taxon name last Modified on 01/15/20	56	123	123	0	15	0	0	2
[60] † Peramodon amalitzkii Taxon name last Modified on 01/15/20	87	95	95	0	12	0	0	2
[61] † Vivaxosaurus trautscholdi Taxon name last Modified on 01/15/20	35	142	140	0	19	0	0	2
[62] † Jimusaria sinkiangensis Taxon name last Modified on 01/15/20	61	120	120	0	13	0	0	2
[63] † Sintocephalus alticeps Taxon name last Modified on 01/15/20	101	80	80	0	13	0	0	2
[64] † Basilodon woodwardi Taxon name last Modified on 01/15/20	70	111	111	0	13	0	0	2
[65] † Turfanodon bogdaensis Taxon name last Modified on 01/15/20	103	86	86	0	5	0	0	2
[66] † Keyseria benjamini Taxon name last Modified on 01/15/20	83	100	100	0	11	0	0	2
[67] † Gordonia traquairi Taxon name last Modified on 01/15/20	81	101	101	0	12	0	0	2
[68] † Syops vanhoepeni Taxon name last Modified on 01/15/20	105	87	87	0	2	0	0	2
[69] † Euptychognathus bathyrhynchus Taxon name last Modified on 01/15/20	66	113	113	0	15	0	0	2
[70] † Lystrosaurus hedini Taxon name last Modified on 01/15/20	46	132	132	0	16	0	0	2
[71] † Lystrosaurus maccaigi Taxon name last Modified on 01/15/20	37	138	138	0	19	0	0	2
[72] † Lystrosaurus curvatus Taxon name last Modified on 01/15/20	33	138	138	0	23	0	0	2
[73] † Lystrosaurus declivis Taxon name last Modified on 01/15/20	12	159	159	0	23	0	0	2
[74] † Lystrosaurus murrayi Taxon name last Modified on 01/15/20	12	159	159	0	23	0	0	2
[75] † Angonisaurus cruickshanki Taxon name last Modified on 01/15/20	49	126	126	0	19	0	0	2
[76] † Tetragonias njalilus Taxon name last Modified on 01/15/20	10	163	162	0	22	0	0	2
[77] † Shansiodon Taxon name last Modified on 01/15/20	34	138	138	0	22	0	0	2
[78] † Vinceria andina Taxon name last Modified on 01/15/20	86	96	96	0	12	0	0	2
[79] † Rhinodicynodon gracile Taxon name last Modified on 01/15/20	45	130	130	0	19	0	0	2
[80] † Dinodontosaurus Taxon name last Modified on 01/15/20	15	156	156	0	23	0	0	2
[81] † Kannemeyeria simocephalus Taxon name last Modified on 01/15/20	11	161	161	0	22	0	0	2
[82] † Kannemeyeria lophorhinus Taxon name last Modified on 01/15/20	53	129	129	0	12	0	0	2
[83] † Dolichuranus primaevus Taxon name last Modified on 01/15/20	35	141	141	0	18	0	0	2
[84] † Wadiasaurus indicus Taxon name last Modified on 01/15/20	68	108	107	0	19	0	0	2
[85] † Rabidosaurus cristatus Taxon name last Modified on 01/15/20	118	73	73	0	3	0	0	2
[86] † Rhadiodromus Taxon name last Modified on 01/15/20	87	95	94	0	13	0	0	2
[87] † Rechnisaurus cristarhynchus Taxon name last Modified on 01/15/20	112	75	75	0	7	0	0	2
[88] † Uralokannemeyeria vjuschkovi Taxon name last Modified on 01/15/20	104	79	79	0	11	0	0	2
[89] † Shaanbeikannemeyeria Taxon name last Modified on 01/15/20	58	121	121	0	15	0	0	2
[90] † Xiyukannemeyeria brevirostris Taxon name last Modified on 01/15/20	55	125	125	0	14	0	0	2
[91] † Parakannemeyeria Taxon name last Modified on 01/15/20	22	150	149	0	23	0	0	2
[92] † Sinokannemeyeria Taxon name last Modified on 01/15/20	27	145	145	0	22	0	0	2
[93] † Stahleckeria potens Taxon name last Modified on 01/15/20	12	161	161	0	21	0	0	2
[94] † Eubrachiosaurus browni Taxon name last Modified on 01/15/20	183	9	9	0	2	0	0	2
[95] † Sangusaurus parringtonii Taxon name last Modified on 01/15/20	46	134	134	0	14	0	0	2
[96] † Ischigualastia jenseni Taxon name last Modified on 01/15/20	24	150	150	0	20	0	0	2
[97] † Jachaleria Taxon name last Modified on 01/15/20	45	130	130	0	19	0	0	2
[98] † Zambiasaurus submersus Taxon name last Modified on 01/15/20	132	60	60	0	2	0	0	2
[99] † Placerias hesternus Taxon name last Modified on 01/15/20	72	120	120	0	2	0	0	2
[100] † Moghreberia nmachouensis Taxon name last Modified on 01/15/20	97	97	97	0	0	0	0	2
[101] † Rastodon procurvidens Taxon name last Modified on 01/15/20	80	101	101	0	13	0	0	2
[102] † Bulbasaurus phylloxyron Taxon name last Modified on 01/15/20	59	125	121	0	14	0	0	2
[103] † Compsodon helmoedi Taxon name last Modified on 01/15/20	70	111	111	0	13	0	0	2
[104] † Counillonia superoculis Taxon name last Modified on 01/15/20	107	78	78	0	9	0	0	2
[105] † Repelinosaurus robustus (1993-2) Taxon name last Modified on 01/15/20	113	75	75	0	6	0	0	2
[106] † Repelinosaurus robustus (1995-9) Taxon name last Modified on 01/15/20	106	82	82	0	6	0	0	2

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	125
Project downloads	122
Document downloads	1	Character list and original tnt matrix (without character names) (1 download);
Matrix downloads	2	Laos (2 downloads);