Project 3621: D. S Berman, H. C. Maddin, A. C. Henrici, S. S. Sumida, D. Scott, R. R. Reisz. 2020. New primitive caseid (Synapsida, Caseasauria) from the Early Permian of Germany. Annals of Carnegie Museum. 86 (1):43-75.
Specimen: † Martensius bromackerensis (mng:14230)
View: Dorsal view

Abstract

A new genus and species of a basal synapsid Caseidae, Martensius bromackerensis, is described based on four partial to nearly complete mostly articulated skeletons that provide a comprehensive knowledge of the skeletal morphology. All four specimens were collected from a single site, the Bromacker quarry, in the Lower Permian Artinskian Tambach Formation, Germany. The Bromacker caseid is the first to be reported from Germany and can be easily distinguished from all other caseids based on substantial lists of autapomorphic and plesiomorphic characters. Of the four caseid specimens only the smallest, a juvenile, and the largest, an adult designated as the holotype, are nearly complete, articulated, and possess skull material: in the juvenile a small partially articulated portion of the skull, and in the adult a nearly complete but dorsoventrally crushed skull. The two specimens are distinguished from one another by features attributed to different ontogenetic stages of development, which include skeletal ossification, proportional dimensions of elements, and most interestingly marginal dentitions. The last category includes a feature unique among caseids of an ontogenetic change in the dentition from insectivorous in the juvenile specimen to what is believed to be an omnivorous dentition in the adult.
A phylogenetic analysis posits the Late Pennsylvanian Eocasea martini Reisz and Fröbisch, 2014, as the basalmost member of the monophyletic Caseidae and the later occurring middle Early Permian Bromacker caseid as the sister taxon of the remaining late Early and Middle Permian members of the clade. This series of relationships parallels a proposed chronology of evolutionary changes in the dentitions and associated diets of caseids.


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Article DOI: https://doi.org/10.2992/007.086.0103

Project DOI: 10.7934/P3621, http://dx.doi.org/10.7934/P3621
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 62 Taxa
  • 1 Specimen
  • 245 Characters
Total size of project's media files: 6.03M

Download Project SDD File
Total scored cells: 7240
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 245
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 581
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:245/0
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MorphoBank Project 3621
  • Creation Date:
    22 January 2020
  • Publication Date:
    01 November 2021
  • Project views: 286

    Authors' Institutions

    • Carleton University

    • University of Toronto

    • California State University - San Bernardino

    • Carnegie Museum of Natural History



    Members

    member name taxa specimens media media
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    Amy Henrici
    Project Administrator
    6211024500007240
    (7240, 0, 0)
    00000
    David Berman
    Full membership
    0000000000
    (0, 0, 0)
    00000
    John Wible
    Observer
    0000000000
    (0, 0, 0)
    00000


    Taxonomic Overview for Matrix 'M26505' (62 Taxa)

    taxon unscored
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    [1] † Tseajaia campi
    Last Modified in 11/01/21
    4520020000003
    [2] † Limnoscelis
    Last Modified in 11/01/21
    3620920900003
    [3] † Captorhinus
    Last Modified in 11/01/21
    2222322300003
    [4] † Protorothyris archeri
    Last Modified in 11/01/21
    7816716700003
    [5] † Dimetrodon spp
    Last Modified in 11/01/21
    524324000003
    [6] † Secodontosaurus obtusidens
    Last Modified in 11/01/21
    5618918900003
    [7] † Ctenorhachis jacksoni
    Last Modified in 11/01/21
    217282800003
    [8] † Sphenacodon ferocior
    Last Modified in 11/01/21
    3720920800003
    [9] † Cryptovenator hirschbergeri
    Last Modified in 11/01/21
    233121200003
    [10] † Titanophoneus potens
    Last Modified in 11/01/21
    3121421400003
    [11] † Biarmosuchus tener
    Last Modified in 11/01/21
    8416116100003
    [12] † Raranimus dashankouensis
    Last Modified in 11/01/21
    210363500003
    [13] † Biseridens qilianicus
    Last Modified in 11/01/21
    12711811800003
    [14] † Pantelosaurus saxonicus
    Last Modified in 11/01/21
    12911611600003
    [15] † Cutleria wilmarthi
    Last Modified in 11/01/21
    146999900003
    [16] † Ianthodon schultzei
    Last Modified in 11/01/21
    12012612500003
    [17] † Haptodus garnettensis
    Last Modified in 11/01/21
    3521121000003
    [18] † Edaphosaurus boanerges
    Last Modified in 11/01/21
    2921621600003
    [19] † Edaphosaurus novomexicanus
    Last Modified in 11/01/21
    12911611600003
    [20] † Lupeosaurus kayi
    Last Modified in 11/01/21
    177686800003
    [21] † Ianthasaurus hardestii
    Last Modified in 11/01/21
    10514014000003
    [22] † Glaucosaurus megalops
    Last Modified in 11/01/21
    202454300003
    [23] † Echinerpeton intermedium
    Last Modified in 11/01/21
    197484800003
    [24] † Archaeothyris florensis
    Last Modified in 11/01/21
    149969600003
    [25] † Varanosaurus acutirostris
    Last Modified in 11/01/21
    5519019000003
    [26] † Ophiacodon spp
    Last Modified in 11/01/21
    1722922800003
    [27] † Stereophallodon ciscoensis
    Last Modified in 11/01/21
    172737300003
    [28] † Oromycter dolesorum
    Last Modified in 11/01/21
    207383800003
    [29] † Ennatosaurus tecton
    Last Modified in 11/01/21
    6518018000003
    [30] † Eothyris parkeyi
    Last Modified in 11/01/21
    12512012000003
    [31] † Oedaleops campi
    Last Modified in 11/01/21
    11413113100003
    [32] † Casea broilii
    Last Modified in 11/01/21
    5718818800003
    [33] † Cotylorhynchus romeri
    Last Modified in 11/01/21
    2921621600003
    [34] † Angelosaurus romeri
    Last Modified in 11/01/21
    13910610600003
    [35] † Angelosaurus dolani
    Last Modified in 11/01/21
    191545400003
    [36] † Caseopsis agilis
    Last Modified in 11/01/21
    214313100003
    [37] † Trichasaurus texensis
    Last Modified in 11/01/21
    210353500003
    [38] † Cotylorhynchus hancocki
    Last Modified in 11/01/21
    11013513500003
    [39] † Cotylorhynchus bransoni
    Last Modified in 11/01/21
    156898900003
    [40] † Euromycter rutenus
    Last Modified in 11/01/21
    11113413400003
    [41] † Watongia meieri
    Last Modified in 11/01/21
    187585800003
    [42] † Ruthiromia elcobriensis
    Last Modified in 11/01/21
    187585800003
    [43] † Varanodon agilis
    Last Modified in 11/01/21
    11113413400003
    [44] † Varanops brevirostris
    Last Modified in 11/01/21
    3820720700003
    [45] † Aerosaurus wellesi
    Last Modified in 11/01/21
    7117417400003
    [46] † Aerosaurus greenleorum
    Last Modified in 11/01/21
    219262600003
    [47] † Archaeovenator hamiltonensis
    Last Modified in 11/01/21
    7017517500003
    [48] † Mesenosaurus romeri
    Last Modified in 11/01/21
    7217317300003
    [49] † Mycterosaurus longiceps
    Last Modified in 11/01/21
    9814714700003
    [50] † Heleosaurus scholtzi
    Last Modified in 11/01/21
    7317317200003
    [51] † BP 1 5678 Elliotsmithia
    Last Modified in 11/01/21
    200464500003
    [52] † Pyozia mesensis
    Last Modified in 11/01/21
    193525200003
    [53] † Eocasea martini
    Last Modified in 11/01/21
    152939300003
    [54] † Casea nicholsi
    Last Modified in 11/01/21
    199464600003
    [55] † Vaughnictis smithae
    Last Modified in 11/01/21
    168777700003
    [56] † Apsisaurus witteri
    Last Modified in 11/01/21
    164818100003
    [57] † Datheosaurus macrous
    Last Modified in 11/01/21
    208373700003
    [58] † Callibrachion gaudreyi
    Last Modified in 11/01/21
    199464600003
    [59] † Phreatophasma aenigmaticum
    Last Modified in 11/01/21
    2369900003
    [60] † Milosaurus mccordi
    Last Modified in 11/01/21
    225202000003
    [61] † Alierasaurus ronchii
    Last Modified in 11/01/21
    235101000003
    [62] † Martensius bromackerensis
    Last Modified in 11/01/21
    8615915900003


    Project views

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    Total project views286
    Media views49Media search (44 views); M686907 (5 views);
    Project overview26
    Matrix views26Matrix landing page (26 views);
    Views for media list15
    Specimen list25
    Taxon list137
    Bibliography8




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    Total downloads from project6
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