Project 3733: A. H. Turner, S. Montanari, M. A. Norell. 2021. A New Dromaeosaurid from the Late Cretaceous Khulsan Locality of Mongolia. American Museum Novitates. 2020 (3965):1-48.
Specimen: † Shri devi (IGM:100/980)
View: ventral

Abstract

A new dromaeosaurid dinosaur, Shri devi, from the Late Cretaceous deposit of the Barun Goyot Formation at Khulsan, Mongolia, is described here. The Barun Goyot Formation (herein referred to as the Barun Goyot) is stratigraphically intermediate between the overlying Nemegt Formation and the underlying Djadokhta Formation, where much of the dromaeosaurid diversity has been reported to date. Sediments of the Barun Goyot are typically considered Upper Campanian in age. Although dromaeosaurid remains have been noted to occur in the Barun Goyot for decades, descriptive and taxonomic work has never been completed for the material. The holotype specimen (IGM 100/980) consists of a partial articulated individual preserving the right hindlimb, left tibiotarsus, pelvis, and adjacent cervical, dorsal and caudal vertebrae. IGM 100/980 is referable to a clade with Velociraptor based on the presence of a distinct ambiens tubercle located proximally on the anterior face of the pubis, a well-developed anterior tuberosity proximally located on the ischium, and a rounded longitudinal ischial ridge. It is distinguishable from V. mongoliensis based on a weak fourth trochanter (shared with all other dromaeosaurids), deep anterior pedicular fossae in the cervical vertebrae, epipophyses in the last four cervicals are not raised but instead represented by rugose circular scars. A suite of axial and appendicular characters diagnoses the new species. New discoveries including Shri devi allow for an improved understanding of dromaeosaurid anatomy, as well as the temporal and regional variation of the dromaeosaurid fauna of Mongolia and Inner Mongolia (Nei Mongol), China during the Late Cretaceous.


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Article DOI: 10.1206/3965.1

Project DOI: 10.7934/P3733, http://dx.doi.org/10.7934/P3733
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 1 Document
  • 157 Taxa
  • 1 Specimen
  • 853 Characters
Total size of project's media files: 34.51M

Download Project SDD File
Total scored cells: 45751
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 853
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 1947
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:713/140
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MorphoBank Project 3733

    This research
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    Authors' Institutions

    • American Museum of Natural History (AMNH)

    • Stony Brook University



    Members

    member name taxa specimens media chars character
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    (scored, NPA, "-")
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    Alan Turner
    Project Administrator
    157118530045751
    (45749, 0, 2)
    000
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    Full membership
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    Taxonomic Overview for Matrix 'M26635' (157 Taxa)

    taxon unscored
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    scored
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    [1] Allosaurus
    Taxon name last Modified on 05/22/20
    8477276900002
    [2] Sinraptor
    Taxon name last Modified on 05/22/20
    21164264200002
    [3] Dromaeosaurus albertensis
    Taxon name last Modified on 05/22/20
    61224124100002
    [4] Deinonychus antirrhopus
    Taxon name last Modified on 05/22/20
    26558958800002
    [5] Velociraptor mongoliensis
    Taxon name last Modified on 05/22/20
    12673172700002
    [6] Velociraptor osmolskae
    Taxon name last Modified on 05/22/20
    821323200002
    [7] Balaur bondoc
    Taxon name last Modified on 05/22/20
    59825725500002
    [8] Tsaagan mangas
    Taxon name last Modified on 05/22/20
    50235135100002
    [9] Linheraptor exquisitus
    Taxon name last Modified on 05/22/20
    46239339100002
    [10] Bambiraptor feinbergorum
    Taxon name last Modified on 05/22/20
    23861761500002
    [11] Graciliraptor lujiatunensis
    Taxon name last Modified on 05/22/20
    75110310200002
    [12] Atrociraptor marshalli
    Taxon name last Modified on 05/22/20
    789656400002
    [13] Utahraptor ostrommaysorum
    Taxon name last Modified on 05/22/20
    72612812700002
    [14] Adasaurus mongoliensis
    Taxon name last Modified on 05/22/20
    56529028800002
    [15] Saurornitholestes langstoni
    Taxon name last Modified on 05/22/20
    68317017000002
    [16] Achillobator giganticus
    Taxon name last Modified on 05/22/20
    65919419400002
    [17] Shanag ashile
    Taxon name last Modified on 05/22/20
    795585800002
    [18] Rahonavis ostromi
    Taxon name last Modified on 05/22/20
    63122322200002
    [19] Mahakala omnogovae
    Taxon name last Modified on 05/22/20
    57128528200002
    [20] Buitreraptor gonzalozorum
    Taxon name last Modified on 05/22/20
    46039739300002
    [21] Austroraptor
    Taxon name last Modified on 05/22/20
    68217317100002
    [22] Neuquenraptor unenlagia
    Taxon name last Modified on 05/22/20
    61923423400002
    [23] Microraptor zhaoianus
    Taxon name last Modified on 05/22/20
    30455254900002
    [24] Sinornithosaurus millenii
    Taxon name last Modified on 05/22/20
    40644844700002
    [25] Tianyuraptor ostromi
    Taxon name last Modified on 05/22/20
    52233133100002
    [26] Zhenyuanlong suni
    Taxon name last Modified on 05/22/20
    39546045800002
    [27] Changyuraptor yangi
    Taxon name last Modified on 05/22/20
    64520920800002
    [28] Hesperonychus elizabethae
    Taxon name last Modified on 05/22/20
    817363600002
    [29] Pyroraptor olympius
    Taxon name last Modified on 05/22/20
    832212100002
    [30] Luanchuanraptor henanensis
    Taxon name last Modified on 05/22/20
    71114214200002
    [31] Yurgovuchia doellingi
    Taxon name last Modified on 05/22/20
    788656500002
    [32] Dakotaraptor steini
    Taxon name last Modified on 05/22/20
    75010310300002
    [33] Acheroraptor temertyorum
    Taxon name last Modified on 05/22/20
    802515100002
    [34] Anchiornis huxleyi
    Taxon name last Modified on 05/22/20
    23262862100002
    [35] Xiaotingia zhengi
    Taxon name last Modified on 05/22/20
    52632832700002
    [36] Eosinopteryx brevipenna
    Taxon name last Modified on 05/22/20
    53931431400002
    [37] Aurornis xui
    Taxon name last Modified on 05/22/20
    51134234200002
    [38] Troodon formosus
    Taxon name last Modified on 05/22/20
    59525825800002
    [39] Jinfengopteryx elegans
    Taxon name last Modified on 05/22/20
    69016416300002
    [40] Mei long
    Taxon name last Modified on 05/22/20
    43941641400002
    [41] Sinovenator changii
    Taxon name last Modified on 05/22/20
    44241441100002
    [42] Sinusonasus magnodens
    Taxon name last Modified on 05/22/20
    66818518500002
    [43] Sinornithoides youngi
    Taxon name last Modified on 05/22/20
    55529929800002
    [44] EK troodontid IGM 10044
    Taxon name last Modified on 05/22/20
    807474600002
    [45] Byronosaurus jaffei
    Taxon name last Modified on 05/22/20
    66418918900002
    [46] Xixiasaurus henanensis
    Taxon name last Modified on 05/22/20
    72512812800002
    [47] Saurornithoides mongoliensis
    Taxon name last Modified on 05/22/20
    60824624500002
    [48] Zanabazar junior
    Taxon name last Modified on 05/22/20
    59925525400002
    [49] Archaeopteryx lithographica
    Taxon name last Modified on 05/22/20
    65020320300002
    [50] Confuciusornis sanctus
    Taxon name last Modified on 05/22/20
    39246546100002
    [51] Sapeornis chaoyangensis
    Taxon name last Modified on 05/22/20
    43542041800002
    [52] Jeholornis prima
    Taxon name last Modified on 05/22/20
    48936436400002
    [53] Jixiangornis orientalis
    Taxon name last Modified on 05/22/20
    52632832700002
    [54] Yanornis martini
    Taxon name last Modified on 05/22/20
    57328528000002
    [55] Apsaravis ukhaana
    Taxon name last Modified on 05/22/20
    56229629100002
    [56] Yixianornis grabaui
    Taxon name last Modified on 05/22/20
    51134634200002
    [57] Neuquenornis volans
    Taxon name last Modified on 05/22/20
    71514113800002
    [58] Patagopteryx deferrariisi
    Taxon name last Modified on 05/22/20
    60025825300002
    [59] Cathayornis yandica
    Taxon name last Modified on 05/22/20
    57628327700002
    [60] Concornis lacustris
    Taxon name last Modified on 05/22/20
    69816115500002
    [61] Gobipteryx minuta
    Taxon name last Modified on 05/22/20
    72513112800002
    [62] Vorona berivotrensis
    Taxon name last Modified on 05/22/20
    777767600002
    [63] Songlingornis linghensis
    Taxon name last Modified on 05/22/20
    790656300002
    [64] Pengornis houi
    Taxon name last Modified on 05/22/20
    58227827100002
    [65] Hesperornis
    Taxon name last Modified on 05/22/20
    50335035000002
    [66] Baptornis advenus
    Taxon name last Modified on 05/22/20
    72912612400002
    [67] Ichthyornis
    Taxon name last Modified on 05/22/20
    43542441800002
    [68] Iaceornis marshii
    Taxon name last Modified on 05/22/20
    73312112000002
    [69] Limenavis patagonica
    Taxon name last Modified on 05/22/20
    814413900002
    [70] Lithornis
    Taxon name last Modified on 05/22/20
    54431730900002
    [71] Hongshanornis longicresta
    Taxon name last Modified on 05/22/20
    62323423000002
    [72] Liaoningornis longidigitris
    Taxon name last Modified on 05/22/20
    784696900002
    [73] Crypturellus undulatus
    Taxon name last Modified on 05/22/20
    45639839700002
    [74] Gallus gallus
    Taxon name last Modified on 05/22/20
    45440139900002
    [75] Crax pauxi
    Taxon name last Modified on 05/22/20
    44840540500002
    [76] Anas platyrhynchus
    Taxon name last Modified on 05/22/20
    46838638500002
    [77] Chauna torquata
    Taxon name last Modified on 05/22/20
    44840640500002
    [78] Epidexipteryx hui
    Taxon name last Modified on 05/22/20
    68017517300002
    [79] Incisivosaurus gauthieri
    Taxon name last Modified on 05/22/20
    57727627600002
    [80] Citipati osmolskae
    Taxon name last Modified on 05/22/20
    30655054700002
    [81] Oviraptor philoceratops
    Taxon name last Modified on 05/22/20
    64520820800002
    [82] Microvenator celer
    Taxon name last Modified on 05/22/20
    61523823800002
    [83] Caudipteryx zoui
    Taxon name last Modified on 05/22/20
    52732732600002
    [84] Ingenia yanshani
    Taxon name last Modified on 05/22/20
    53332132000002
    [85] Rinchenia mongoliensis
    Taxon name last Modified on 05/22/20
    68616716700002
    [86] Conchoraptor gracilis
    Taxon name last Modified on 05/22/20
    53332132000002
    [87] Chirostenotes pergracilis
    Taxon name last Modified on 05/22/20
    56129429200002
    [88] Avimimus portentosus
    Taxon name last Modified on 05/22/20
    52832632500002
    [89] Falcarius utahensis
    Taxon name last Modified on 05/22/20
    31553953800002
    [90] Beipiaosaurus inexpectus
    Taxon name last Modified on 05/22/20
    65519819800002
    [91] Segnosaurus galbinensis
    Taxon name last Modified on 05/22/20
    64321021000002
    [92] Erlikosaurus andrewsi
    Taxon name last Modified on 05/22/20
    52832532500002
    [93] Alxasaurus elesitaiensis
    Taxon name last Modified on 05/22/20
    65020420300002
    [94] Neimongosaurus yangi
    Taxon name last Modified on 05/22/20
    62722722600002
    [95] Erliansaurus bellamanus
    Taxon name last Modified on 05/22/20
    72213113100002
    [96] Suzhousaurus megatherioides
    Taxon name last Modified on 05/22/20
    64920520400002
    [97] Nothronychus
    Taxon name last Modified on 05/22/20
    50235335100002
    [98] Enigmosaurus mongoliensis
    Taxon name last Modified on 05/22/20
    788656500002
    [99] Nanshiungosaurus bohlini
    Taxon name last Modified on 05/22/20
    814393900002
    [100] Therizinosaurus cheloniformis
    Taxon name last Modified on 05/22/20
    73411911900002
    [101] Haplocheirus sollers
    Taxon name last Modified on 05/22/20
    35449949900002
    [102] Alvarezsaurus calvoi
    Taxon name last Modified on 05/22/20
    70215115100002
    [103] Patagonykus puertai
    Taxon name last Modified on 05/22/20
    65020320300002
    [104] Achillesaurus
    Taxon name last Modified on 05/22/20
    794595900002
    [105] Mononykus olecranus
    Taxon name last Modified on 05/22/20
    53531831800002
    [106] Shuvuuia deserti
    Taxon name last Modified on 05/22/20
    25659959700002
    [107] Parvicursor remotus
    Taxon name last Modified on 05/22/20
    72013313300002
    [108] Albertonykus borealis
    Taxon name last Modified on 05/22/20
    829242400002
    [109] Albinykus baatar
    Taxon name last Modified on 05/22/20
    785686800002
    [110] Bonapartenykus ultimus
    Taxon name last Modified on 05/22/20
    780737300002
    [111] Ceratonykus oculatus
    Taxon name last Modified on 05/22/20
    71314014000002
    [112] Linhenykus monodactylus
    Taxon name last Modified on 05/22/20
    69116216200002
    [113] Xixianykus zhangi
    Taxon name last Modified on 05/22/20
    68916416400002
    [114] Nqwebasaurus thwazi
    Taxon name last Modified on 05/22/20
    65619719700002
    [115] Shenzhousaurus orientalis
    Taxon name last Modified on 05/22/20
    59625725700002
    [116] Ornithomimus edmonticus
    Taxon name last Modified on 05/22/20
    33951551400002
    [117] Archaeornithomimus asiaticus
    Taxon name last Modified on 05/22/20
    51334334000002
    [118] Anserimimus planinychus
    Taxon name last Modified on 05/22/20
    72812512500002
    [119] Struthiomimus altus
    Taxon name last Modified on 05/22/20
    23861661500002
    [120] Gallimimus bullatus
    Taxon name last Modified on 05/22/20
    16169469200002
    [121] Garudimimus brevipes
    Taxon name last Modified on 05/22/20
    35150250200002
    [122] Pelecanimimus polydon
    Taxon name last Modified on 05/22/20
    61024324300002
    [123] Harpymimus okladnikovi
    Taxon name last Modified on 05/22/20
    48037437300002
    [124] Beishanlong
    Taxon name last Modified on 05/22/20
    66119219200002
    [125] Sinornithomimus dongi
    Taxon name last Modified on 05/22/20
    30155255200002
    [126] Qiupalong henanensis
    Taxon name last Modified on 05/22/20
    74311011000002
    [127] Huaxiagnathus orientalis
    Taxon name last Modified on 05/22/20
    52632832700002
    [128] Sinosauropteryx prima
    Taxon name last Modified on 05/22/20
    49835635500002
    [129] Compsognathus longipes
    Taxon name last Modified on 05/22/20
    42043443300002
    [130] Juravenator starki
    Taxon name last Modified on 05/22/20
    48936536400002
    [131] Sinocalliopteryx gigas
    Taxon name last Modified on 05/22/20
    45839639500002
    [132] Mirischia asymmetrica
    Taxon name last Modified on 05/22/20
    777767600002
    [133] Ornitholestes hermanni
    Taxon name last Modified on 05/22/20
    33851651500002
    [134] Coelurus fragilis
    Taxon name last Modified on 05/22/20
    62722622600002
    [135] Tanycolagreus topwilsoni
    Taxon name last Modified on 05/22/20
    49735735600002
    [136] Tugulusaurus faciles
    Taxon name last Modified on 05/22/20
    786676700002
    [137] Zuolong salleei
    Taxon name last Modified on 05/22/20
    49435935900002
    [138] Bicentenaria argentina
    Taxon name last Modified on 05/22/20
    67517817800002
    [139] Kileskus aristotocus
    Taxon name last Modified on 05/22/20
    783707000002
    [140] Guanlong wucaii
    Taxon name last Modified on 05/22/20
    17867667500002
    [141] Sinotyrannus kazuoensis
    Taxon name last Modified on 05/22/20
    772818100002
    [142] Proceratosaurus bradleyi
    Taxon name last Modified on 05/22/20
    65419919900002
    [143] Dilong paradoxus
    Taxon name last Modified on 05/22/20
    28556856800002
    [144] Eotyrannus lengi
    Taxon name last Modified on 05/22/20
    58626726700002
    [145] Juratyrant langhami
    Taxon name last Modified on 05/22/20
    69515815800002
    [146] Xiongguanlong baimoensis
    Taxon name last Modified on 05/22/20
    54331031000002
    [147] Dryptosaurus acquilunguis
    Taxon name last Modified on 05/22/20
    71014414300002
    [148] Appalachiosaurus montgomeriensis
    Taxon name last Modified on 05/22/20
    61124224200002
    [149] Bistahieversor sealeyi
    Taxon name last Modified on 05/22/20
    54730630600002
    [150] Albertosaurus sarcophagus
    Taxon name last Modified on 05/22/20
    16769368600002
    [151] Gorgosaurus libratus
    Taxon name last Modified on 05/22/20
    8876876500002
    [152] Alioramus
    Taxon name last Modified on 05/22/20
    31054454300002
    [153] Teratophoneus curriei
    Taxon name last Modified on 05/22/20
    71114214200002
    [154] Daspletosaurus torosus
    Taxon name last Modified on 05/22/20
    18966766400002
    [155] Tyrannosaurus rex
    Taxon name last Modified on 05/22/20
    6978978400002
    [156] Tarbosaurus baatar
    Taxon name last Modified on 05/22/20
    9475975900002
    [157] † Shri devi
    Taxon name last Modified on 05/22/20
    62922322202002


    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project167
    Matrix downloads56A new dromaeosaurid from Khulsan (56 downloads);
    Project downloads103
    Document downloads8TNT file with analysis script (8 downloads);