Project 3770: C. De Jaime-Soguero, E. Mujal, J. Fortuny. 2020. First xiphosuran traceway in the middle Muschelkalk facies (Middle Triassic) of the Catalan Basin (NE Iberian Peninsula). Spanish Journal of Palaeontology. 35 (2):205.
Specimen: † Kouphichnium isp (Institut Català de Paleontologia Miquel Crusafont:IPS110272)
View: Traceway


In the last decade, the first ichnoassamblages from the middle Muschelkalk facies (upper Anisian–middle Ladinian) of the Catalan Basin (NE Iberian Peninsula) have been discovered. Herein, the first xiphosuran trace fossils are described from the locality of Penya Rubí, a newly discovered ichnosite from the Catalan Basin. The finding opens a window into peri-Tethys ecosystems with coastal influence. The traceway is referred to the ichnogenus Kouphichnium: a locomotion trace attributed to xiphosurans. The traceway preserves telson grooves and different imprint morphologies from the various appendages. The traceway pattern and arrangement of the different traces suggest a crawling locomotion style. The sedimentology suggests a coastal zone with areas influenced by tides (intertidal flat). The morphological variations of the ichnites are correlated to substrate rheology, the locomotion of the tracemaker and environmental conditions.

Read the article »

Article DOI: 10.7203/sjp.35.2.18483

Project DOI: 10.7934/P3770,
This project contains
  • 1 Media
  • 1 Taxon
  • 1 Specimen
Total size of project's media files: 4.22M

Download Project SDD File
Currently Viewing:
MorphoBank Project 3770
  • Creation Date:
    09 July 2020
  • Publication Date:
    24 December 2020
  • Project views: 508

    Authors' Institutions

    • Staatliches Museum fur Naturkunde Stuttgart

    • Institut Catala de Paleontologia

    • Universitat Autònoma de Barcelona


    member name taxa specimens media
    Josep Fortuny
    Project Administrator

    Project has no matrices defined.

    Project views

    type number of views Individual items viewed (where applicable)
    Total project views508
    Project overview103
    Media views176Media search (150 views); M692668 (26 views);
    Views for media list59
    Specimen list78
    Taxon list62

    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project15
    Project downloads15