Project 386: D. A. Croft, F. Anaya. 2006. A new middle Miocene hegetotheriid (Notoungulata: Typotheria) and a phylogeny of the Hegetotheriidae. Journal of Vertebrate Paleontology. 26 (2):387-399.
Specimen: † Hemihegetotherium trilobus (Croft and Anaya, 2006) (MNHN-BOL/V:006387)

Abstract

A new species of hegetotheriid notoungulate, Hemihegetotherium trilobus, is described from the middle Miocene (Laventan SALMA) Quebrada Honda Fauna of southern Bolivia. It differs from all previously described hegetotheriines in possessing a distinctly trilobed m3, a character state previously recorded only in pachyrukhine he- getotheriids. It further differs from other species of Hemihegetotherium in having a proportionately larger p2 and in overall size (being smaller than H. achataleptum and larger than H. torresi). A phylogenetic analysis of hegetotheriid genera coded for 26 characters supports referral of this species to Hemihegetotherium, but does not support the mono- phyly of Hegetotheriinae; the monophyly of Pachyrukhinae (including Prosotherium and Propachyrucos) is strongly supported. The resulting tree suggests that the lack of tibiofibular fusion in Oligocene pachyrukhines is a secondarily derived trait, but additional postcranial data are needed to provide a stronger test of this hypothesis.
Hemihegetotherium trilobus is the most abundant ungulate at Quebrada Honda and is the only species of hegetotheriid present. This contrasts with most other South American paleofaunas in which pachyrukhines are more common than hegetotheriines (e.g., Arroyo Chasicó) and/or other small notoungulates are more common than hegetotheriids (e.g., Chucal, Nazareno). Although the absence of pachyrukhines at Quebrada Honda may be due to a lack of suitable habitat, it is also possible that pachyrukhines had not dispersed north to southern Bolivia by the middle Miocene. Further investigations of Quebrada Honda may help discriminate between these two alternatives and should also help clarify the biogeographic histories of other Miocene taxa.


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Project DOI: 10.7934/P386, http://dx.doi.org/10.7934/P386
This project containsMatrices
  • 4 Media
  • 1 Matrix
  • 1 Document
  • 10 Taxa
  • 4 Specimens
  • 21 Characters
Total size of project's media files: 857.47k

Download Project SDD File
Total scored cells: 211
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 21
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 46
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:21/0
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MorphoBank Project 386

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    Authors' Institutions

    • Case Western Reserve University

    • Universidad Autónoma Tomás Frías



    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
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    Darin Croft
    Project Administrator
    10442100211
    (211, 0, 0)
    000
    Maureen Admin
    Full membership
    0000000
    (0, 0, 0)
    000
    MorphoBank Curator
    Full membership
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    Taxonomic Overview for Matrix 'M884' (10 Taxa)

    taxon unscored
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    scored
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    "-" cellscell images labels on
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    [1] Archaeohyrax
    Last Modified in 12/05/13
    2191900003
    [2] Prohegetotherium
    Last Modified in 12/05/13
    0222100003
    [3] Prosotherium
    Last Modified in 12/05/13
    0212100003
    [4] Propachyrucos
    Last Modified in 12/05/13
    1202000003
    [5] Hegetotherium
    Last Modified in 12/05/13
    0222100003
    [6] Hemihegetotherium
    Last Modified in 12/05/13
    3181800003
    [7] Pachyrukhos
    Last Modified in 12/05/13
    0212100003
    [8] Paedotherium
    Last Modified in 12/05/13
    0242100003
    [9] Tremacyllus
    Last Modified in 12/05/13
    0222100003
    [10] † Hemihegetotherium trilobus
    Last Modified in 12/05/13
    1222000003


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