Morphobank

Project 3966: A. E. Fernandes, O. Mateus, B. Andres, M. J. Polcyn, A. S. Schulp, A. O. Gonçalves, L. L. Jacobs. 2022. Pterosaurs from the Late Cretaceous of Angola. Diversity. 14 (9):741.

Specimen: † Pterosauria
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Abstract

Here, we describe the first pterosaur remains from Angola, an assemblage of fourteen bones from the Lower Maastrichtian marine deposits of Bentiaba, Namibe Province. One new species is introduced, Epapatelo otyikokolo, gen. et sp. nov., which comprises an articulated partial left humerus and ulna as well as an articulated left ulna and radius (from a second individual). Phylogenetic analysis confirms a non-nyctosaurid pteranodontian attribution for this new taxon and supports a new apomorphy-based clade, Aponyctosauria, which is here defined. Late Cretaceous pteranodontians are rare in Sub-Saharan Africa and throughout the Southern Hemisphere. Preliminary histological analysis also reveals a likely sub-adult age for one of the specimens. This fossil assemblage provides a first glimpse of Angolan pterosaur paleobiodiversity providing further insight into the Gondwanan ecosystems of the Upper Cretaceous.

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Article DOI: 10.3390/d14090741

Project DOI: 10.7934/P3966, http://dx.doi.org/10.7934/P3966

This project contains	Matrices
16 Media 1 Matrix 139 Taxa 16 Specimens 271 Characters Total size of project's media files: 3.06G Download Project SDD File	Total scored cells: 14584 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 271 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 548 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:245/26

Currently Viewing:
MorphoBank Project 3966

Creation Date:
25 March 2021
Publication Date:
03 March 2023
Media downloads: 2
Matrix downloads: 5

Authors' Institutions

University of Sheffield

Universiteit Utrecht

Museu da Lourinha

Bayerische Staatssammlung fuer Palaeontologie und Geologie

Naturalis Biodiversity Center

Universidade Nova de Lisboa

Southern Methodist University

Universidade Agostinho Neto

Members

Alexandra Fernandes
Project Administrator

139

271

14584
(12177, 0, 2407)

Taxonomic Overview for Matrix 'M29172' (132 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] Euparkeria capensis Taxon name last Modified on 03/06/23	47	189	189	0	35	0	0	1
[2] Ornithosuchus longidens Taxon name last Modified on 03/06/23	73	165	164	0	34	0	0	1
[3] Herrerasaurus ischigualastensis Taxon name last Modified on 03/06/23	64	173	173	0	34	0	0	1
[4] Scleromochlus taylori Taxon name last Modified on 03/06/23	114	122	122	0	35	0	0	1
[5] Eudimorphodon ranzii Taxon name last Modified on 03/06/23	78	166	166	0	27	0	0	1
[6] Eudimorphodon rosenfeldi Taxon name last Modified on 03/06/23	62	167	167	0	42	0	0	1
[7] Eudimorphodon cromptonellus Taxon name last Modified on 03/06/23	166	91	90	0	15	0	0	1
[8] Peteinosaurus zambellii Taxon name last Modified on 03/06/23	212	52	52	0	7	0	0	1
[9] Caviramus schesaplanensis Taxon name last Modified on 03/06/23	229	40	39	0	3	0	0	1
[10] Raeticodactylus filisurensis Taxon name last Modified on 03/06/23	102	141	141	0	28	0	0	1
[11] Austriadactylus cristatus Taxon name last Modified on 03/06/23	101	141	140	0	30	0	0	1
[12] Preondactylus buffarinii Taxon name last Modified on 03/06/23	100	131	131	0	40	0	0	1
[13] Dimorphodon macronyx Taxon name last Modified on 03/06/23	55	173	173	0	43	0	0	1
[14] Parapsicephalus purdoni Taxon name last Modified on 03/06/23	192	74	73	0	6	0	0	1
[15] Campylognathoides liasicus Taxon name last Modified on 03/06/23	37	188	188	0	46	0	0	1
[16] Campylognathoides zitteli Taxon name last Modified on 03/06/23	71	160	160	0	40	0	0	1
[17] Sericipterus wucaiwanensis Taxon name last Modified on 03/06/23	134	123	123	0	14	0	0	1
[18] Angustinaripterus longicephalus Taxon name last Modified on 03/06/23	155	104	104	0	12	0	0	1
[19] Harpactognathus gentryii Taxon name last Modified on 03/06/23	218	52	51	0	2	0	0	1
[20] Cacibupteryx caribensis Taxon name last Modified on 03/06/23	188	77	76	0	7	0	0	1
[21] Rhamphorhynchus muensteri Taxon name last Modified on 03/06/23	30	198	195	0	46	0	0	1
[22] Qinglongopterus guoi Taxon name last Modified on 03/06/23	113	120	120	0	38	0	0	1
[23] Nesodactylus hesperius Taxon name last Modified on 03/06/23	202	53	53	0	16	0	0	1
[24] Dorygnathus banthensis Taxon name last Modified on 03/06/23	29	197	196	0	46	0	0	1
[25] Scaphognathus crassirostris Taxon name last Modified on 03/06/23	48	179	178	0	45	0	0	1
[26] Sordes pilosus Taxon name last Modified on 03/06/23	36	191	190	0	45	0	0	1
[27] Darwinopterus modularis Taxon name last Modified on 03/06/23	64	166	166	0	41	0	0	1
[28] Wukongopterus lii Taxon name last Modified on 03/06/23	131	111	111	0	29	0	0	1
[29] Pterorhynchus wellnhoferi Taxon name last Modified on 03/06/23	96	141	141	0	34	0	0	1
[30] Changchengopterus pani Taxon name last Modified on 03/06/23	203	48	48	0	20	0	0	1
[31] Batrachognathus volans Taxon name last Modified on 03/06/23	114	132	132	0	25	0	0	1
[32] Jeholopterus ningchengensis Taxon name last Modified on 03/06/23	93	138	138	0	40	0	0	1
[33] Anurognathus ammoni Taxon name last Modified on 03/06/23	87	144	144	0	40	0	0	1
[34] Dendrorhynchoides curvidentatus Taxon name last Modified on 03/06/23	141	104	104	0	26	0	0	1
[35] Kryptodrakon progenitor Taxon name last Modified on 03/06/23	253	16	16	0	2	0	0	1
[36] Gnathosaurus subulatus Taxon name last Modified on 03/06/23	159	102	102	0	10	0	0	1
[37] Gnathosaurus macrurus Taxon name last Modified on 03/06/23	226	40	40	0	5	0	0	1
[38] Plataleorhynchus streptophorodon Taxon name last Modified on 03/06/23	239	31	31	0	1	0	0	1
[39] Huanhepterus quingyangensis Taxon name last Modified on 03/06/23	167	85	85	0	19	0	0	1
[40] Moganopterus zhuiana Taxon name last Modified on 03/06/23	163	95	95	0	13	0	0	1
[41] Elanodactylus prolatus Taxon name last Modified on 03/06/23	186	65	65	0	20	0	0	1
[42] Kepodactylus insperatus Taxon name last Modified on 03/06/23	241	28	28	0	2	0	0	1
[43] Ctenochasma elegans Taxon name last Modified on 03/06/23	56	174	173	0	42	0	0	1
[44] Pterodaustro guinazui Taxon name last Modified on 03/06/23	66	166	166	0	39	0	0	1
[45] Beipiaopterus chenianus Taxon name last Modified on 03/06/23	208	37	37	0	26	0	0	1
[46] Gegepterus changae Taxon name last Modified on 03/06/23	147	107	106	0	18	0	0	1
[47] Feilongus youngi Taxon name last Modified on 03/06/23	149	109	109	0	13	0	0	1
[48] Cycnorhamphus suevicus Taxon name last Modified on 03/06/23	61	175	174	0	36	0	0	1
[49] Ardeadactylus longicollum Taxon name last Modified on 03/06/23	58	174	174	0	39	0	0	1
[50] Pterodactylus antiquus Taxon name last Modified on 03/06/23	50	179	179	0	42	0	0	1
[51] Normannognathus wellnhoferi Taxon name last Modified on 03/06/23	217	53	53	0	1	0	0	1
[52] Germanodactylus cristatus Taxon name last Modified on 03/06/23	65	165	165	0	41	0	0	1
[53] Germanodactylus rhamphastinus Taxon name last Modified on 03/06/23	83	153	153	0	35	0	0	1
[54] Haopterus gracilis Taxon name last Modified on 03/06/23	133	112	112	0	26	0	0	1
[55] Aetodactylus halli Taxon name last Modified on 03/06/23	216	51	51	0	4	0	0	1
[56] Anhanguera santanae Taxon name last Modified on 03/06/23	67	179	179	0	25	0	0	1
[57] Anhanguera piscator Taxon name last Modified on 03/06/23	52	186	186	0	33	0	0	1
[58] Anhanguera blittersdorffi Taxon name last Modified on 03/06/23	144	113	113	0	14	0	0	1
[59] Anhanguera araripensis Taxon name last Modified on 03/06/23	136	123	123	0	12	0	0	1
[60] Liaoningopterus gui Taxon name last Modified on 03/06/23	173	94	93	0	5	0	0	1
[61] Siroccopteryx moroccensis Taxon name last Modified on 03/06/23	236	35	35	0	0	0	0	1
[62] Tropeognathus mesembrinus Taxon name last Modified on 03/06/23	139	117	117	0	15	0	0	1
[63] Coloborhynchus clavirostris Taxon name last Modified on 03/06/23	241	30	30	0	0	0	0	1
[64] Coloborhynchus wadleighi Taxon name last Modified on 03/06/23	236	35	35	0	0	0	0	1
[65] Ornithocheirus simus Taxon name last Modified on 03/06/23	244	27	27	0	0	0	0	1
[66] Brasileodactylus araripensis Taxon name last Modified on 03/06/23	233	38	38	0	0	0	0	1
[67] Ludodactylus sibbicki Taxon name last Modified on 03/06/23	147	111	111	0	13	0	0	1
[68] Boreopterus cuiae Taxon name last Modified on 03/06/23	144	104	103	0	24	0	0	1
[69] Guidraco venator Taxon name last Modified on 03/06/23	161	97	97	0	13	0	0	1
[70] Zhenyuanopterus longiristris Taxon name last Modified on 03/06/23	99	140	139	0	33	0	0	1
[71] Cearadactylus atrox Taxon name last Modified on 03/06/23	157	103	103	0	11	0	0	1
[72] Hongshanopterus lacustris Taxon name last Modified on 03/06/23	191	73	72	0	8	0	0	1
[73] Lonchodectes compressirostris Taxon name last Modified on 03/06/23	235	35	35	0	1	0	0	1
[74] Lonchodraco giganteus Taxon name last Modified on 03/06/23	199	70	70	0	2	0	0	1
[75] Cimoliopterus cuvieri Taxon name last Modified on 03/06/23	229	41	41	0	1	0	0	1
[76] Cimoliopterus dunni Taxon name last Modified on 03/06/23	231	39	39	0	1	0	0	1
[77] Nurhachius ignaciobritoi Taxon name last Modified on 03/06/23	86	152	151	0	34	0	0	1
[78] Liaoxipterus brachyognathus Taxon name last Modified on 03/06/23	235	32	32	0	4	0	0	1
[79] Istiodactylus sinensis Taxon name last Modified on 03/06/23	120	123	123	0	28	0	0	1
[80] Istiodactylus latidens Taxon name last Modified on 03/06/23	109	140	140	0	22	0	0	1
[81] Pteranodon longiceps Taxon name last Modified on 03/06/23	66	171	171	0	34	0	0	1
[82] Pteranodon sternbergi Taxon name last Modified on 03/06/23	136	100	100	0	35	0	0	1
[83] Tethydraco regalis Taxon name last Modified on 03/06/23	258	12	12	0	1	0	0	1
[84] Alamodactylus byrdi Taxon name last Modified on 03/06/23	255	15	15	0	1	0	0	1
[85] Nyctosaurus grandis Taxon name last Modified on 03/06/23	238	26	26	0	7	0	0	1
[86] Alcione elainus Taxon name last Modified on 03/06/23	227	37	37	0	7	0	0	1
[87] Simurghia robusta Taxon name last Modified on 03/06/23	265	5	5	0	1	0	0	1
[88] Cretornis hlavaci Taxon name last Modified on 03/06/23	253	18	18	0	0	0	0	1
[89] Nyctosaurus gracilis Taxon name last Modified on 03/06/23	70	163	163	0	38	0	0	1
[90] Nyctosaurus nanus Taxon name last Modified on 03/06/23	245	24	24	0	2	0	0	1
[91] Nyctosaurus lamegoi Taxon name last Modified on 03/06/23	265	5	5	0	1	0	0	1
[92] Muzquizopteryx coahuilensis Taxon name last Modified on 03/06/23	167	91	91	0	13	0	0	1
[93] Tupandactylus navigans Taxon name last Modified on 03/06/23	202	61	61	0	8	0	0	1
[94] Tupandactylus imperator Taxon name last Modified on 03/06/23	189	74	74	0	8	0	0	1
[95] Bakonydraco galaczi Taxon name last Modified on 03/06/23	237	31	31	0	3	0	0	1
[96] Tapejara wellnhoferi Taxon name last Modified on 03/06/23	63	172	172	0	36	0	0	1
[97] Europejara olcadesorum Taxon name last Modified on 03/06/23	223	45	45	0	3	0	0	1
[98] Vectidraco daisymorrisae Taxon name last Modified on 03/06/23	257	11	11	0	3	0	0	1
[99] Caiuajara dobruskii Taxon name last Modified on 03/06/23	114	136	135	0	22	0	0	1
[100] Huaxiapterus benxiensis Taxon name last Modified on 03/06/23	153	96	96	0	22	0	0	1
[101] Huaxiapterus corallatus Taxon name last Modified on 03/06/23	144	94	94	0	33	0	0	1
[102] Eopteranodon lii Taxon name last Modified on 03/06/23	157	90	90	0	24	0	0	1
[103] Huaxiapterus jii Taxon name last Modified on 03/06/23	142	100	100	0	29	0	0	1
[104] Sinopterus dongi Taxon name last Modified on 03/06/23	100	134	134	0	37	0	0	1
[105] Bennettazhia oregonensis Taxon name last Modified on 03/06/23	253	16	16	0	2	0	0	1
[106] Dsungaripterus weii Taxon name last Modified on 03/06/23	54	181	181	0	36	0	0	1
[107] Domeykodactylus ceciliae Taxon name last Modified on 03/06/23	240	30	30	0	1	0	0	1
[108] Noripterus parvus Taxon name last Modified on 03/06/23	72	168	168	0	31	0	0	1
[109] Noripterus complicidens Taxon name last Modified on 03/06/23	153	92	92	0	26	0	0	1
[110] Tupuxuara longicristatus Taxon name last Modified on 03/06/23	234	36	36	0	1	0	0	1
[111] Tupuxuara leonardii Taxon name last Modified on 03/06/23	89	149	149	0	33	0	0	1
[112] Alanqa saharica Taxon name last Modified on 03/06/23	244	26	26	0	1	0	0	1
[113] Aerotitan sudamericanus Taxon name last Modified on 03/06/23	255	14	14	0	2	0	0	1
[114] Thalassodromeus sethi Taxon name last Modified on 03/06/23	167	96	96	0	8	0	0	1
[115] Chaoyangopterus zhangi Taxon name last Modified on 03/06/23	163	84	84	0	24	0	0	1
[116] Jidapterus edentus Taxon name last Modified on 03/06/23	151	88	88	0	32	0	0	1
[117] Eoazhdarcho liaoxiensis Taxon name last Modified on 03/06/23	199	53	53	0	19	0	0	1
[118] Shenzhoupterus chaoyangensis Taxon name last Modified on 03/06/23	169	84	84	0	18	0	0	1
[119] Radiodactylus langstoni Taxon name last Modified on 03/06/23	258	13	13	0	0	0	0	1
[120] Microtuban altivolans Taxon name last Modified on 03/06/23	232	28	28	0	11	0	0	1
[121] Aralazhdarcho bestobensis Taxon name last Modified on 03/06/23	254	17	17	0	0	0	0	1
[122] Volgadraco bogolubovi Taxon name last Modified on 03/06/23	242	26	26	0	3	0	0	1
[123] Eurazhdarcho langendorfensis Taxon name last Modified on 03/06/23	259	10	10	0	2	0	0	1
[124] Montanazhdarcho minor Taxon name last Modified on 03/06/23	211	53	53	0	7	0	0	1
[125] Phosphatodraco mauritanicus Taxon name last Modified on 03/06/23	261	9	9	0	1	0	0	1
[126] Zhejiangopterus linhaiensis Taxon name last Modified on 03/06/23	126	114	114	0	31	0	0	1
[127] Azhdarcho lancicollis Taxon name last Modified on 03/06/23	184	84	84	0	3	0	0	1
[128] OCP dek GE 716 Taxon name last Modified on 03/06/23	263	7	7	0	1	0	0	1
[129] Hatzegopteryx thambema Taxon name last Modified on 03/06/23	251	20	20	0	0	0	0	1
[130] Arambourgiania philadelphiae Taxon name last Modified on 03/06/23	262	8	8	0	1	0	0	1
[131] Quetzalcoatlus spp Taxon name last Modified on 03/06/23	106	146	145	0	20	0	0	1
[132] Epapatelo otyikokolo Taxon name last Modified on 03/06/23	261	9	9	0	1	0	0	1

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	8
Media downloads	2	M838532 (2 downloads);
Matrix downloads	5	Pterosaurs from the Late Cretaceous of Angola (5 downloads);
Project downloads	1