Morphobank

Project 3966: A. E. Fernandes, O. Mateus, B. Andres, M. J. Polcyn, A. S. Schulp, A. O. Gonçalves, L. L. Jacobs. 2022. Pterosaurs from the Late Cretaceous of Angola. Diversity. 14 (9):741.

Specimen: † Pterosauria
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Abstract

Here, we describe the first pterosaur remains from Angola, an assemblage of fourteen bones from the Lower Maastrichtian marine deposits of Bentiaba, Namibe Province. One new species is introduced, Epapatelo otyikokolo, gen. et sp. nov., which comprises an articulated partial left humerus and ulna as well as an articulated left ulna and radius (from a second individual). Phylogenetic analysis confirms a non-nyctosaurid pteranodontian attribution for this new taxon and supports a new apomorphy-based clade, Aponyctosauria, which is here defined. Late Cretaceous pteranodontians are rare in Sub-Saharan Africa and throughout the Southern Hemisphere. Preliminary histological analysis also reveals a likely sub-adult age for one of the specimens. This fossil assemblage provides a first glimpse of Angolan pterosaur paleobiodiversity providing further insight into the Gondwanan ecosystems of the Upper Cretaceous.

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Article DOI: 10.3390/d14090741

Project DOI: 10.7934/P3966, http://dx.doi.org/10.7934/P3966

This project contains	Matrices
16 Media 1 Matrix 139 Taxa 16 Specimens 271 Characters Total size of project's media files: 3.06G Download Project SDD File	Total scored cells: 14584 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 271 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 548 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:245/26

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MorphoBank Project 3966

Creation Date:
25 March 2021
Publication Date:
03 March 2023
Project views: 7004
Media downloads: 2
Matrix downloads: 5

Authors' Institutions

University of Sheffield

Universiteit Utrecht

Museu da Lourinha

Bayerische Staatssammlung fuer Palaeontologie und Geologie

Naturalis Biodiversity Center

Universidade Nova de Lisboa

Southern Methodist University

Universidade Agostinho Neto

Members

Alexandra Fernandes
Project Administrator

139

271

14584
(12177, 0, 2407)

Taxonomic Overview for Matrix 'M29172' (132 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] Euparkeria capensis Last Modified in 03/06/23	47	189	189	0	35	0	0	1
[2] Ornithosuchus longidens Last Modified in 03/06/23	73	165	164	0	34	0	0	1
[3] Herrerasaurus ischigualastensis Last Modified in 03/06/23	64	173	173	0	34	0	0	1
[4] Scleromochlus taylori Last Modified in 03/06/23	114	122	122	0	35	0	0	1
[5] Eudimorphodon ranzii Last Modified in 03/06/23	78	166	166	0	27	0	0	1
[6] Eudimorphodon rosenfeldi Last Modified in 03/06/23	62	167	167	0	42	0	0	1
[7] Eudimorphodon cromptonellus Last Modified in 03/06/23	166	91	90	0	15	0	0	1
[8] Peteinosaurus zambellii Last Modified in 03/06/23	212	52	52	0	7	0	0	1
[9] Caviramus schesaplanensis Last Modified in 03/06/23	229	40	39	0	3	0	0	1
[10] Raeticodactylus filisurensis Last Modified in 03/06/23	102	141	141	0	28	0	0	1
[11] Austriadactylus cristatus Last Modified in 03/06/23	101	141	140	0	30	0	0	1
[12] Preondactylus buffarinii Last Modified in 03/06/23	100	131	131	0	40	0	0	1
[13] Dimorphodon macronyx Last Modified in 03/06/23	55	173	173	0	43	0	0	1
[14] Parapsicephalus purdoni Last Modified in 03/06/23	192	74	73	0	6	0	0	1
[15] Campylognathoides liasicus Last Modified in 03/06/23	37	188	188	0	46	0	0	1
[16] Campylognathoides zitteli Last Modified in 03/06/23	71	160	160	0	40	0	0	1
[17] Sericipterus wucaiwanensis Last Modified in 03/06/23	134	123	123	0	14	0	0	1
[18] Angustinaripterus longicephalus Last Modified in 03/06/23	155	104	104	0	12	0	0	1
[19] Harpactognathus gentryii Last Modified in 03/06/23	218	52	51	0	2	0	0	1
[20] Cacibupteryx caribensis Last Modified in 03/06/23	188	77	76	0	7	0	0	1
[21] Rhamphorhynchus muensteri Last Modified in 03/06/23	30	198	195	0	46	0	0	1
[22] Qinglongopterus guoi Last Modified in 03/06/23	113	120	120	0	38	0	0	1
[23] Nesodactylus hesperius Last Modified in 03/06/23	202	53	53	0	16	0	0	1
[24] Dorygnathus banthensis Last Modified in 03/06/23	29	197	196	0	46	0	0	1
[25] Scaphognathus crassirostris Last Modified in 03/06/23	48	179	178	0	45	0	0	1
[26] Sordes pilosus Last Modified in 03/06/23	36	191	190	0	45	0	0	1
[27] Darwinopterus modularis Last Modified in 03/06/23	64	166	166	0	41	0	0	1
[28] Wukongopterus lii Last Modified in 03/06/23	131	111	111	0	29	0	0	1
[29] Pterorhynchus wellnhoferi Last Modified in 03/06/23	96	141	141	0	34	0	0	1
[30] Changchengopterus pani Last Modified in 03/06/23	203	48	48	0	20	0	0	1
[31] Batrachognathus volans Last Modified in 03/06/23	114	132	132	0	25	0	0	1
[32] Jeholopterus ningchengensis Last Modified in 03/06/23	93	138	138	0	40	0	0	1
[33] Anurognathus ammoni Last Modified in 03/06/23	87	144	144	0	40	0	0	1
[34] Dendrorhynchoides curvidentatus Last Modified in 03/06/23	141	104	104	0	26	0	0	1
[35] Kryptodrakon progenitor Last Modified in 03/06/23	253	16	16	0	2	0	0	1
[36] Gnathosaurus subulatus Last Modified in 03/06/23	159	102	102	0	10	0	0	1
[37] Gnathosaurus macrurus Last Modified in 03/06/23	226	40	40	0	5	0	0	1
[38] Plataleorhynchus streptophorodon Last Modified in 03/06/23	239	31	31	0	1	0	0	1
[39] Huanhepterus quingyangensis Last Modified in 03/06/23	167	85	85	0	19	0	0	1
[40] Moganopterus zhuiana Last Modified in 03/06/23	163	95	95	0	13	0	0	1
[41] Elanodactylus prolatus Last Modified in 03/06/23	186	65	65	0	20	0	0	1
[42] Kepodactylus insperatus Last Modified in 03/06/23	241	28	28	0	2	0	0	1
[43] Ctenochasma elegans Last Modified in 03/06/23	56	174	173	0	42	0	0	1
[44] Pterodaustro guinazui Last Modified in 03/06/23	66	166	166	0	39	0	0	1
[45] Beipiaopterus chenianus Last Modified in 03/06/23	208	37	37	0	26	0	0	1
[46] Gegepterus changae Last Modified in 03/06/23	147	107	106	0	18	0	0	1
[47] Feilongus youngi Last Modified in 03/06/23	149	109	109	0	13	0	0	1
[48] Cycnorhamphus suevicus Last Modified in 03/06/23	61	175	174	0	36	0	0	1
[49] Ardeadactylus longicollum Last Modified in 03/06/23	58	174	174	0	39	0	0	1
[50] Pterodactylus antiquus Last Modified in 03/06/23	50	179	179	0	42	0	0	1
[51] Normannognathus wellnhoferi Last Modified in 03/06/23	217	53	53	0	1	0	0	1
[52] Germanodactylus cristatus Last Modified in 03/06/23	65	165	165	0	41	0	0	1
[53] Germanodactylus rhamphastinus Last Modified in 03/06/23	83	153	153	0	35	0	0	1
[54] Haopterus gracilis Last Modified in 03/06/23	133	112	112	0	26	0	0	1
[55] Aetodactylus halli Last Modified in 03/06/23	216	51	51	0	4	0	0	1
[56] Anhanguera santanae Last Modified in 03/06/23	67	179	179	0	25	0	0	1
[57] Anhanguera piscator Last Modified in 03/06/23	52	186	186	0	33	0	0	1
[58] Anhanguera blittersdorffi Last Modified in 03/06/23	144	113	113	0	14	0	0	1
[59] Anhanguera araripensis Last Modified in 03/06/23	136	123	123	0	12	0	0	1
[60] Liaoningopterus gui Last Modified in 03/06/23	173	94	93	0	5	0	0	1
[61] Siroccopteryx moroccensis Last Modified in 03/06/23	236	35	35	0	0	0	0	1
[62] Tropeognathus mesembrinus Last Modified in 03/06/23	139	117	117	0	15	0	0	1
[63] Coloborhynchus clavirostris Last Modified in 03/06/23	241	30	30	0	0	0	0	1
[64] Coloborhynchus wadleighi Last Modified in 03/06/23	236	35	35	0	0	0	0	1
[65] Ornithocheirus simus Last Modified in 03/06/23	244	27	27	0	0	0	0	1
[66] Brasileodactylus araripensis Last Modified in 03/06/23	233	38	38	0	0	0	0	1
[67] Ludodactylus sibbicki Last Modified in 03/06/23	147	111	111	0	13	0	0	1
[68] Boreopterus cuiae Last Modified in 03/06/23	144	104	103	0	24	0	0	1
[69] Guidraco venator Last Modified in 03/06/23	161	97	97	0	13	0	0	1
[70] Zhenyuanopterus longiristris Last Modified in 03/06/23	99	140	139	0	33	0	0	1
[71] Cearadactylus atrox Last Modified in 03/06/23	157	103	103	0	11	0	0	1
[72] Hongshanopterus lacustris Last Modified in 03/06/23	191	73	72	0	8	0	0	1
[73] Lonchodectes compressirostris Last Modified in 03/06/23	235	35	35	0	1	0	0	1
[74] Lonchodraco giganteus Last Modified in 03/06/23	199	70	70	0	2	0	0	1
[75] Cimoliopterus cuvieri Last Modified in 03/06/23	229	41	41	0	1	0	0	1
[76] Cimoliopterus dunni Last Modified in 03/06/23	231	39	39	0	1	0	0	1
[77] Nurhachius ignaciobritoi Last Modified in 03/06/23	86	152	151	0	34	0	0	1
[78] Liaoxipterus brachyognathus Last Modified in 03/06/23	235	32	32	0	4	0	0	1
[79] Istiodactylus sinensis Last Modified in 03/06/23	120	123	123	0	28	0	0	1
[80] Istiodactylus latidens Last Modified in 03/06/23	109	140	140	0	22	0	0	1
[81] Pteranodon longiceps Last Modified in 03/06/23	66	171	171	0	34	0	0	1
[82] Pteranodon sternbergi Last Modified in 03/06/23	136	100	100	0	35	0	0	1
[83] Tethydraco regalis Last Modified in 03/06/23	258	12	12	0	1	0	0	1
[84] Alamodactylus byrdi Last Modified in 03/06/23	255	15	15	0	1	0	0	1
[85] Nyctosaurus grandis Last Modified in 03/06/23	238	26	26	0	7	0	0	1
[86] Alcione elainus Last Modified in 03/06/23	227	37	37	0	7	0	0	1
[87] Simurghia robusta Last Modified in 03/06/23	265	5	5	0	1	0	0	1
[88] Cretornis hlavaci Last Modified in 03/06/23	253	18	18	0	0	0	0	1
[89] Nyctosaurus gracilis Last Modified in 03/06/23	70	163	163	0	38	0	0	1
[90] Nyctosaurus nanus Last Modified in 03/06/23	245	24	24	0	2	0	0	1
[91] Nyctosaurus lamegoi Last Modified in 03/06/23	265	5	5	0	1	0	0	1
[92] Muzquizopteryx coahuilensis Last Modified in 03/06/23	167	91	91	0	13	0	0	1
[93] Tupandactylus navigans Last Modified in 03/06/23	202	61	61	0	8	0	0	1
[94] Tupandactylus imperator Last Modified in 03/06/23	189	74	74	0	8	0	0	1
[95] Bakonydraco galaczi Last Modified in 03/06/23	237	31	31	0	3	0	0	1
[96] Tapejara wellnhoferi Last Modified in 03/06/23	63	172	172	0	36	0	0	1
[97] Europejara olcadesorum Last Modified in 03/06/23	223	45	45	0	3	0	0	1
[98] Vectidraco daisymorrisae Last Modified in 03/06/23	257	11	11	0	3	0	0	1
[99] Caiuajara dobruskii Last Modified in 03/06/23	114	136	135	0	22	0	0	1
[100] Huaxiapterus benxiensis Last Modified in 03/06/23	153	96	96	0	22	0	0	1
[101] Huaxiapterus corallatus Last Modified in 03/06/23	144	94	94	0	33	0	0	1
[102] Eopteranodon lii Last Modified in 03/06/23	157	90	90	0	24	0	0	1
[103] Huaxiapterus jii Last Modified in 03/06/23	142	100	100	0	29	0	0	1
[104] Sinopterus dongi Last Modified in 03/06/23	100	134	134	0	37	0	0	1
[105] Bennettazhia oregonensis Last Modified in 03/06/23	253	16	16	0	2	0	0	1
[106] Dsungaripterus weii Last Modified in 03/06/23	54	181	181	0	36	0	0	1
[107] Domeykodactylus ceciliae Last Modified in 03/06/23	240	30	30	0	1	0	0	1
[108] Noripterus parvus Last Modified in 03/06/23	72	168	168	0	31	0	0	1
[109] Noripterus complicidens Last Modified in 03/06/23	153	92	92	0	26	0	0	1
[110] Tupuxuara longicristatus Last Modified in 03/06/23	234	36	36	0	1	0	0	1
[111] Tupuxuara leonardii Last Modified in 03/06/23	89	149	149	0	33	0	0	1
[112] Alanqa saharica Last Modified in 03/06/23	244	26	26	0	1	0	0	1
[113] Aerotitan sudamericanus Last Modified in 03/06/23	255	14	14	0	2	0	0	1
[114] Thalassodromeus sethi Last Modified in 03/06/23	167	96	96	0	8	0	0	1
[115] Chaoyangopterus zhangi Last Modified in 03/06/23	163	84	84	0	24	0	0	1
[116] Jidapterus edentus Last Modified in 03/06/23	151	88	88	0	32	0	0	1
[117] Eoazhdarcho liaoxiensis Last Modified in 03/06/23	199	53	53	0	19	0	0	1
[118] Shenzhoupterus chaoyangensis Last Modified in 03/06/23	169	84	84	0	18	0	0	1
[119] Radiodactylus langstoni Last Modified in 03/06/23	258	13	13	0	0	0	0	1
[120] Microtuban altivolans Last Modified in 03/06/23	232	28	28	0	11	0	0	1
[121] Aralazhdarcho bestobensis Last Modified in 03/06/23	254	17	17	0	0	0	0	1
[122] Volgadraco bogolubovi Last Modified in 03/06/23	242	26	26	0	3	0	0	1
[123] Eurazhdarcho langendorfensis Last Modified in 03/06/23	259	10	10	0	2	0	0	1
[124] Montanazhdarcho minor Last Modified in 03/06/23	211	53	53	0	7	0	0	1
[125] Phosphatodraco mauritanicus Last Modified in 03/06/23	261	9	9	0	1	0	0	1
[126] Zhejiangopterus linhaiensis Last Modified in 03/06/23	126	114	114	0	31	0	0	1
[127] Azhdarcho lancicollis Last Modified in 03/06/23	184	84	84	0	3	0	0	1
[128] OCP dek GE 716 Last Modified in 03/06/23	263	7	7	0	1	0	0	1
[129] Hatzegopteryx thambema Last Modified in 03/06/23	251	20	20	0	0	0	0	1
[130] Arambourgiania philadelphiae Last Modified in 03/06/23	262	8	8	0	1	0	0	1
[131] Quetzalcoatlus spp Last Modified in 03/06/23	106	146	145	0	20	0	0	1
[132] Epapatelo otyikokolo Last Modified in 03/06/23	261	9	9	0	1	0	0	1

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	7004
Project overview	304
Matrix views	197	Matrix landing page (181 views); (3 views); Pterosaurs from the Late Cretaceous of Angola (13 views);
Media views	2253	Media search (598 views); M838532 (115 views); M838540 (108 views); M838538 (110 views); M740037 (100 views); M838537 (107 views); M838539 (99 views); M838541 (100 views); M838543 (101 views); M838544 (104 views); M838545 (104 views); M838546 (105 views); M838547 (100 views); M838548 (101 views); M838551 (100 views); M838552 (101 views); M838555 (100 views);
Specimen list	1535
Taxon list	2212
Bibliography	140
Views for media list	358
Documents list	5

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	8
Media downloads	2	M838532 (2 downloads);
Matrix downloads	5	Pterosaurs from the Late Cretaceous of Angola (5 downloads);
Project downloads	1