Morphobank

Project 3968: A. E. Fernandes, V. Beccari, A. W. Kellner, O. Mateus. 2023. A new gnathosaurine (Pterosauria, Archaeopterodactyloidea) from the Late Jurassic of Portugal. PeerJ. 11:e16048.

Specimen: † Portuguese gnathosaurine (unvouchered)
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Abstract

An incomplete, yet remarkably-sized dentated rostrum and associated partial cervical vertebrae of a pterosaur (ML 2554) were recently discovered from the Late Jurassic (Late Kimmeridgian-Early Tithonian) Lourinhã Formation of Praia do Caniçal, of central west Portugal. This specimen exhibits features such as a spatulated anterior expansion of the rostrum, robust comb-like dentition, and pronounced rims of the tooth alveoli, indicating gnathosaurine affinities. Based on its further unique tooth and dentary morphology, a new genus and species, Lusognathus almadrava gen. et spec. nov., is proposed, making this the first named pterosaur species found within Portugal. The presence of this taxon adds yet another element to the fluvio-deltaic lagoonal environment that has been suggested as representative of the Lourinhã Formation in the Late Jurassic, further contributing to the diversity and distribution of gnathosaurines worldwide.

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Article DOI: 10.7717/peerj.16048

Project DOI: 10.7934/P3968, http://dx.doi.org/10.7934/P3968

This project contains	Matrices
1 Media 1 Matrix 181 Taxa 1 Specimen 275 Characters Total size of project's media files: 157.28k Download Project SDD File	Total scored cells: 21743 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 275 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 617 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:241/34

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MorphoBank Project 3968

Creation Date:
25 March 2021
Publication Date:
18 August 2023
Project views: 3090
Matrix downloads: 3

Authors' Institutions

Ludwig-Maximilians-Universitaet Muenchen

Museu da Lourinha

Museu Nacional do Rio de Janeiro

Bayerische Staatssammlung fuer Palaeontologie und Geologie

Universidade Nova de Lisboa

Members

Alexandra Fernandes
Project Administrator

181

275

21743
(15518, 0, 6225)

Victor Beccari
Full membership

0
(0, 0, 0)

Taxonomic Overview for Matrix 'M29183' (180 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Euparkeria capensis Last Modified in 08/18/23	14	192	192	0	69	0	0	2
[2] † Ornithosuchus longidens Last Modified in 08/18/23	47	168	167	0	61	0	0	2
[3] † Herrerasaurus ischigualastensis Last Modified in 08/18/23	37	177	177	0	61	0	0	2
[4] † Scleromochlus taylori Last Modified in 08/18/23	88	124	124	0	63	0	0	2
[5] † Preondactylus buffarinii Last Modified in 08/18/23	86	132	132	0	57	0	0	2
[6] † Austriadactylus cristatus Last Modified in 08/18/23	88	143	143	0	44	0	0	2
[7] † Peteinosaurus zambellii Last Modified in 08/18/23	213	52	52	0	10	0	0	2
[8] † Caviramus schesaplanensis Last Modified in 08/18/23	230	39	38	0	7	0	0	2
[9] † Raeticodactylus filisurensis Last Modified in 08/18/23	94	143	143	0	38	0	0	2
[10] † Eudimorphodon cromptonellus Last Modified in 08/18/23	165	91	90	0	20	0	0	2
[11] † Eudimorphodon rosenfeldi Last Modified in 08/18/23	47	169	169	0	59	0	0	2
[12] † Eudimorphodon ranzii Last Modified in 08/18/23	58	169	169	0	48	0	0	2
[13] † Parapsicephalus purdoni Last Modified in 08/18/23	176	74	73	0	26	0	0	2
[14] † Dimorphodon macronyx Last Modified in 08/18/23	37	176	176	0	62	0	0	2
[15] † Dimorphodon koi Last Modified in 08/18/23	223	47	46	0	6	0	0	2
[16] † Dimorphodon hanseni Last Modified in 08/18/23	188	76	76	0	11	0	0	2
[17] † Dimorphodon jenkinsi Last Modified in 08/18/23	248	28	27	0	0	0	0	2
[18] † Dimorphodon weintraubi Last Modified in 08/18/23	204	55	54	0	17	0	0	2
[19] † Herbstosaurus pigmaeus Last Modified in 08/18/23	258	15	15	0	2	0	0	2
[20] † Campylognathoides zitteli Last Modified in 08/18/23	49	164	164	0	62	0	0	2
[21] † Campylognathoides liasicus Last Modified in 08/18/23	13	192	192	0	70	0	0	2
[22] † Scaphognathus crassirostris Last Modified in 08/18/23	27	183	182	0	66	0	0	2
[23] † Orientognathus chaoyangensis Last Modified in 08/18/23	108	121	119	0	48	0	0	2
[24] † Dorygnathus banthensis Last Modified in 08/18/23	6	203	202	0	67	0	0	2
[25] † Klobiodon rochei Last Modified in 08/18/23	224	42	42	0	9	0	0	2
[26] † Dolicorhamphus depressirostris Last Modified in 08/18/23	233	37	37	0	5	0	0	2
[27] † Dolicorhamphus bucklandi Last Modified in 08/18/23	234	34	34	0	7	0	0	2
[28] † Fenghuangopterus lii Last Modified in 08/18/23	90	135	134	0	51	0	0	2
[29] † Rhamphorhynchus muensteri Last Modified in 08/18/23	4	202	199	0	72	0	0	2
[30] † Nesodactylus hesperius Last Modified in 08/18/23	204	55	55	0	16	0	0	2
[31] † Cacibupteryx caribensis Last Modified in 08/18/23	178	77	76	0	21	0	0	2
[32] † Qinglongopterus guoi Last Modified in 08/18/23	104	124	124	0	47	0	0	2
[33] † Harpactognathus gentryii Last Modified in 08/18/23	217	53	52	0	6	0	0	2
[34] † Angustinaripterus longicephalus Last Modified in 08/18/23	145	106	106	0	24	0	0	2
[35] † Sericipterus wucaiwanensis Last Modified in 08/18/23	123	126	126	0	26	0	0	2
[36] † Sordes pilosus Last Modified in 08/18/23	11	195	194	0	70	0	0	2
[37] † Pterorhynchus wellnhoferi Last Modified in 08/18/23	76	145	145	0	54	0	0	2
[38] † Kunpengopterus sinensis Last Modified in 08/18/23	77	144	144	0	54	0	0	2
[39] † Wukongopterus lii Last Modified in 08/18/23	112	117	117	0	46	0	0	2
[40] † Darwinopterus zhengi Last Modified in 08/18/23	207	46	46	0	22	0	0	2
[41] † Darwinopterus linglongtaensis Last Modified in 08/18/23	55	161	161	0	59	0	0	2
[42] † Darwinopterus robustodens Last Modified in 08/18/23	47	167	167	0	61	0	0	2
[43] † Archaeoistiodactylus linglongtaensis Last Modified in 08/18/23	171	80	80	0	24	0	0	2
[44] † Darwinopterus modularis Last Modified in 08/18/23	44	169	169	0	62	0	0	2
[45] † Changchengopterus pani Last Modified in 08/18/23	206	49	49	0	20	0	0	2
[46] † Dendrorhynchoides curvidentatus Last Modified in 08/18/23	117	108	108	0	50	0	0	2
[47] † Luopterus mutoudengensis Last Modified in 08/18/23	125	102	101	0	49	0	0	2
[48] † Batrachognathus volans Last Modified in 08/18/23	74	136	136	0	65	0	0	2
[49] † Jeholopterus ningchengensis Last Modified in 08/18/23	59	142	142	0	74	0	0	2
[50] † Anurognathus ammoni Last Modified in 08/18/23	51	148	148	0	76	0	0	2
[51] † Kryptodrakon progenitor Last Modified in 08/18/23	257	16	16	0	2	0	0	2
[52] † Painten pterodactyloid Last Modified in 08/18/23	39	167	167	0	69	0	0	2
[53] † Cuspicephalus scarfi Last Modified in 08/18/23	175	76	76	0	24	0	0	2
[54] † Germanodactylus rhamphastinus Last Modified in 08/18/23	61	157	157	0	57	0	0	2
[55] † Germanodactylus cristatus Last Modified in 08/18/23	43	169	169	0	63	0	0	2
[56] † Pterodactylus antiquus Last Modified in 08/18/23	19	183	183	0	73	0	0	2
[57] † Normannognathus wellnhoferi Last Modified in 08/18/23	209	57	57	0	9	0	0	2
[58] † Aurorazhdarcho primordius Last Modified in 08/18/23	206	46	46	0	23	0	0	2
[59] † Cycnorhamphus suevicus Last Modified in 08/18/23	31	182	181	0	63	0	0	2
[60] † Pterodactylus micronyx Last Modified in 08/18/23	15	185	185	0	75	0	0	2
[61] † Liaodactylus primus Last Modified in 08/18/23	137	102	102	0	36	0	0	2
[62] † Ctenochasma elegans Last Modified in 08/18/23	31	178	177	0	67	0	0	2
[63] † Pterodaustro guinazui Last Modified in 08/18/23	36	170	170	0	69	0	0	2
[64] † Beipiaopterus chenianus Last Modified in 08/18/23	209	39	39	0	27	0	0	2
[65] † Gegepterus changi Last Modified in 08/18/23	129	109	108	0	38	0	0	2
[66] † Kepodactylus insperatus Last Modified in 08/18/23	242	30	30	0	3	0	0	2
[67] † Elanodactylus prolatus Last Modified in 08/18/23	188	67	67	0	20	0	0	2
[68] † Moganopterus zhuiana Last Modified in 08/18/23	146	97	97	0	32	0	0	2
[69] † Feilongus youngi Last Modified in 08/18/23	127	113	113	0	35	0	0	2
[70] † Ardeadactylus longicollum Last Modified in 08/18/23	30	179	179	0	66	0	0	2
[71] † Huanhepterus quingyangensis Last Modified in 03/07/23	157	90	90	0	28	0	0	2
[72] † Plataleorhynchus streptophorodon Last Modified in 08/18/23	233	33	33	0	9	0	0	2
[73] † Gnathosaurus macrurus Last Modified in 08/18/23	224	42	42	0	9	0	0	2
[74] † Gnathosaurus subulatus Last Modified in 08/18/23	139	104	104	0	32	0	0	2
[75] † Haopterus gracilis Last Modified in 08/18/23	100	119	119	0	56	0	0	2
[76] † Ornithostoma sedgwicki Last Modified in 08/18/23	227	15	15	0	33	0	0	2
[77] † Volgadraco bogolubovi Last Modified in 08/18/23	218	28	28	0	29	0	0	2
[78] † Tethydraco regalis Last Modified in 08/18/23	262	12	12	0	1	0	0	2
[79] † Pteranodon sternbergi Last Modified in 08/18/23	90	105	105	0	80	0	0	2
[80] † Pteranodon longiceps Last Modified in 08/18/23	17	174	174	0	84	0	0	2
[81] † Alamodactylus byrdi Last Modified in 08/18/23	259	15	15	0	1	0	0	2
[82] † Cretornis hlavaci Last Modified in 08/18/23	257	18	18	0	0	0	0	2
[83] † Simurghia robusta Last Modified in 08/18/23	269	5	5	0	1	0	0	2
[84] † Alcione elainus Last Modified in 08/18/23	231	37	37	0	7	0	0	2
[85] † Muzquizopteryx coahuilensis Last Modified in 08/18/23	127	90	90	0	58	0	0	2
[86] † Nyctosaurus grandis Last Modified in 08/18/23	238	28	28	0	9	0	0	2
[87] † Nyctosaurus lamegoi Last Modified in 08/18/23	269	5	5	0	1	0	0	2
[88] † Nyctosaurus nanus Last Modified in 08/18/23	247	24	24	0	4	0	0	2
[89] † Nyctosaurus gracilis Last Modified in 08/18/23	19	166	166	0	90	0	0	2
[90] † Serradraco sagittirostris Last Modified in 08/18/23	253	20	20	0	2	0	0	2
[91] † Aussiedraco molnari Last Modified in 08/18/23	237	32	32	0	6	0	0	2
[92] † Hongshanopterus lacustris Last Modified in 08/18/23	180	75	74	0	21	0	0	2
[93] † Targaryendraco wiedenrothi Last Modified in 08/18/23	231	37	37	0	7	0	0	2
[94] † Lonchodectes compressirostris Last Modified in 08/18/23	214	46	46	0	15	0	0	2
[95] † Ikrandraco avatar Last Modified in 08/18/23	88	137	137	0	50	0	0	2
[96] † Ikrandraco machaerorhynchus Last Modified in 08/18/23	249	24	24	0	2	0	0	2
[97] † Lonchodraco giganteus Last Modified in 08/18/23	188	72	72	0	15	0	0	2
[98] † Lonchodraco microdon Last Modified in 08/18/23	246	20	20	0	9	0	0	2
[99] † Lonchodraco denticulatus Last Modified in 08/18/23	249	24	24	0	2	0	0	2
[100] † Nurhachius ignaciobritoi Last Modified in 08/18/23	55	155	154	0	66	0	0	2
[101] † Liaoxipterus brachyognathus Last Modified in 08/18/23	234	31	31	0	10	0	0	2
[102] † Istiodactylus sinensis Last Modified in 08/18/23	88	125	125	0	62	0	0	2
[103] † Istiodactylus latidens Last Modified in 08/18/23	76	144	144	0	55	0	0	2
[104] † Pterodactylus polyodon Last Modified in 08/18/23	238	27	27	0	10	0	0	2
[105] † Zhenyuanopterus longirostris Last Modified in 08/18/23	83	137	137	0	55	0	0	2
[106] † Boreopterus giganticus Last Modified in 08/18/23	142	90	90	0	43	0	0	2
[107] † Boreopterus cuiae Last Modified in 08/18/23	112	107	106	0	57	0	0	2
[108] † Hamipterus tianshanensis Last Modified in 08/18/23	47	176	176	0	52	0	0	2
[109] † Brasileodactylus araripensis Last Modified in 08/18/23	232	37	37	0	6	0	0	2
[110] † Barbosania gracilirostris Last Modified in 08/18/23	103	123	123	0	49	0	0	2
[111] † Cearadactylus atrox Last Modified in 08/18/23	142	97	97	0	36	0	0	2
[112] † Guidraco venator Last Modified in 08/18/23	140	98	98	0	37	0	0	2
[113] † Ludodactylus colorhinus Last Modified in 08/18/23	251	23	23	0	1	0	0	2
[114] † Ludodactylus sibbicki Last Modified in 08/18/23	128	110	110	0	37	0	0	2
[115] † Aetodactylus halli Last Modified in 08/18/23	204	51	51	0	20	0	0	2
[116] † Camposipterus nasutus Last Modified in 08/18/23	237	29	29	0	9	0	0	2
[117] † Cimoliopterus dunni Last Modified in 08/18/23	231	40	40	0	4	0	0	2
[118] † Cimoliopterus cuvieri Last Modified in 08/18/23	231	40	40	0	4	0	0	2
[119] † Aerodraco sedgwickii Last Modified in 08/18/23	243	28	28	0	4	0	0	2
[120] † Liaoningopterus gui Last Modified in 08/18/23	160	96	95	0	20	0	0	2
[121] † Anhanguera robustus Last Modified in 08/18/23	112	135	135	0	28	0	0	2
[122] † Anhanguera piscator Last Modified in 08/18/23	35	189	189	0	51	0	0	2
[123] † Anhanguera blittersdorffi Last Modified in 08/18/23	129	114	114	0	32	0	0	2
[124] † Tropeognathus mesembrinus Last Modified in 08/18/23	123	117	117	0	35	0	0	2
[125] † Ornithocheirus platystomus Last Modified in 08/18/23	251	24	24	0	0	0	0	2
[126] † Ornithocheirus simus Last Modified in 08/18/23	246	28	28	0	1	0	0	2
[127] † Siroccopteryx moroccensis Last Modified in 08/18/23	238	34	34	0	3	0	0	2
[128] † Coloborhynchus capito Last Modified in 08/18/23	246	26	26	0	3	0	0	2
[129] † Coloborhynchus wadleighi Last Modified in 08/18/23	238	34	34	0	3	0	0	2
[130] † Coloborhynchus clavirostris Last Modified in 08/18/23	245	29	29	0	1	0	0	2
[131] † Eopteranodon lii Last Modified in 08/18/23	122	92	92	0	61	0	0	2
[132] † Huaxiapterus jii Last Modified in 08/18/23	107	101	101	0	67	0	0	2
[133] † Sinopterus dongi Last Modified in 08/18/23	64	137	137	0	74	0	0	2
[134] † Sinopterus benxiensis Last Modified in 08/18/23	116	99	99	0	60	0	0	2
[135] † Sinopterus corollatus Last Modified in 08/18/23	109	97	97	0	69	0	0	2
[136] † Bakonydraco galaczi Last Modified in 08/18/23	195	50	50	0	30	0	0	2
[137] † Keresdrakon vilsoni Last Modified in 08/18/23	132	98	97	0	46	0	0	2
[138] † Caiuajara dobruskii Last Modified in 08/18/23	75	139	138	0	62	0	0	2
[139] † Tupandactylus navigans Last Modified in 08/18/23	172	61	61	0	42	0	0	2
[140] † Tupandactylus imperator Last Modified in 08/18/23	151	75	75	0	49	0	0	2
[141] † Vectidraco daisymorrisae Last Modified in 08/18/23	262	10	10	0	3	0	0	2
[142] † Europejara olcadesorum Last Modified in 08/18/23	196	44	44	0	35	0	0	2
[143] † Tapejara wellnhoferi Last Modified in 08/18/23	24	175	175	0	76	0	0	2
[144] † Microtuban altivolans Last Modified in 08/18/23	235	28	28	0	12	0	0	2
[145] † Noripterus complicidens Last Modified in 08/18/23	147	95	95	0	33	0	0	2
[146] † Noripterus parvus Last Modified in 08/18/23	51	172	172	0	52	0	0	2
[147] † Domeykodactylus ceciliae Last Modified in 08/18/23	240	31	31	0	4	0	0	2
[148] † Dsungaripterus weii Last Modified in 08/18/23	33	185	185	0	57	0	0	2
[149] † Tupuxuara leonardii Last Modified in 08/18/23	47	152	152	0	76	0	0	2
[150] † Tupuxuara longicristatus Last Modified in 08/18/23	205	36	36	0	34	0	0	2
[151] † Thalassodromeus sethi Last Modified in 08/18/23	131	96	96	0	48	0	0	2
[152] † Lacusovagus magnificens Last Modified in 08/18/23	214	26	26	0	35	0	0	2
[153] † Xericeps curvirostris Last Modified in 08/18/23	232	18	18	0	25	0	0	2
[154] † Alanqa saharica Last Modified in 08/18/23	229	23	23	0	23	0	0	2
[155] † Argentinadraco barrealensis Last Modified in 08/18/23	230	22	22	0	23	0	0	2
[156] † Aerotitan sudamericanus Last Modified in 08/18/23	235	15	15	0	25	0	0	2
[157] † Eoazhdarcho liaoxiensis Last Modified in 08/18/23	168	59	59	0	48	0	0	2
[158] † Shenzhoupterus chaoyangensis Last Modified in 08/18/23	117	88	88	0	70	0	0	2
[159] † Jidapterus edentus Last Modified in 08/18/23	66	127	127	0	82	0	0	2
[160] † Chaoyangopterus zhangi Last Modified in 08/18/23	108	96	96	0	71	0	0	2
[161] † Radiodactylus langstoni Last Modified in 08/18/23	262	13	13	0	0	0	0	2
[162] † Montanazhdarcho minor Last Modified in 08/18/23	188	55	55	0	32	0	0	2
[163] † Azhdarcho lancicollis Last Modified in 08/18/23	138	90	90	0	47	0	0	2
[164] † Albadraco tharmisensis Last Modified in 08/18/23	219	27	27	0	29	0	0	2
[165] † Mistralazhdarcho maggii Last Modified in 08/18/23	200	38	38	0	37	0	0	2
[166] † Leptostomia begaaensis Last Modified in 08/18/23	221	26	26	0	28	0	0	2
[167] † Apatorhamphus gyrostega Last Modified in 08/18/23	207	30	30	0	38	0	0	2
[168] † Aralazhdarcho bostobensis Last Modified in 08/18/23	233	20	20	0	22	0	0	2
[169] † Phosphatodraco mauritanicus Last Modified in 08/18/23	263	11	11	0	1	0	0	2
[170] † Eurazhdarcho langendorfensis Last Modified in 08/18/23	261	12	12	0	2	0	0	2
[171] † Zhejiangopterus linhaiensis Last Modified in 08/18/23	74	116	116	0	85	0	0	2
[172] † Cryodrakon boreas Last Modified in 08/18/23	237	34	34	0	4	0	0	2
[173] † Wellnhopterus brevirostris Last Modified in 08/18/23	178	59	59	0	38	0	0	2
[174] † Hatzegopteryx thambema Last Modified in 08/18/23	210	39	39	0	26	0	0	2
[175] † Arambourgiania philadelphiae Last Modified in 08/18/23	264	10	10	0	1	0	0	2
[176] † Quetzalcoatlus lawsoni Last Modified in 08/18/23	38	160	158	0	79	0	0	2
[177] † Quetzalcoatlus northropi Last Modified in 08/18/23	236	34	34	0	5	0	0	2
[178] † Dearc sgiathanach Last Modified in 08/18/23	252	16	16	0	7	0	0	2
[179] † Tacuadactylus luciae Last Modified in 03/07/23	265	10	10	0	0	0	0	2
[180] † Lusognathus almadrava Last Modified in 03/07/23	248	25	25	0	2	0	0	2

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type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	3
Matrix downloads	3	A new gnathosaurine from the Late Jurassic of Portugal (3 downloads);