Morphobank

Project 3968: A. E. Fernandes, V. Beccari, A. W. Kellner, O. Mateus. 2023. A new gnathosaurine (Pterosauria, Archaeopterodactyloidea) from the Late Jurassic of Portugal. PeerJ. 11:e16048.

Specimen: † Portuguese gnathosaurine (unvouchered)
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Abstract

An incomplete, yet remarkably-sized dentated rostrum and associated partial cervical vertebrae of a pterosaur (ML 2554) were recently discovered from the Late Jurassic (Late Kimmeridgian-Early Tithonian) Lourinhã Formation of Praia do Caniçal, of central west Portugal. This specimen exhibits features such as a spatulated anterior expansion of the rostrum, robust comb-like dentition, and pronounced rims of the tooth alveoli, indicating gnathosaurine affinities. Based on its further unique tooth and dentary morphology, a new genus and species, Lusognathus almadrava gen. et spec. nov., is proposed, making this the first named pterosaur species found within Portugal. The presence of this taxon adds yet another element to the fluvio-deltaic lagoonal environment that has been suggested as representative of the Lourinhã Formation in the Late Jurassic, further contributing to the diversity and distribution of gnathosaurines worldwide.

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Article DOI: 10.7717/peerj.16048

Project DOI: 10.7934/P3968, http://dx.doi.org/10.7934/P3968

This project contains	Matrices
1 Media 1 Matrix 181 Taxa 1 Specimen 275 Characters Total size of project's media files: 157.28k Download Project SDD File	Total scored cells: 21743 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 275 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 617 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:241/34

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MorphoBank Project 3968

Creation Date:
25 March 2021
Publication Date:
18 August 2023
Project views: 5152
Matrix downloads: 9

Authors' Institutions

Ludwig-Maximilians-Universitaet Muenchen

Museu da Lourinha

Museu Nacional do Rio de Janeiro

Bayerische Staatssammlung fuer Palaeontologie und Geologie

Universidade Nova de Lisboa

Members

Alexandra Fernandes
Project Administrator

181

275

21743
(15518, 0, 6225)

Victor Beccari
Full membership

0
(0, 0, 0)

Taxonomic Overview for Matrix 'M29183' (180 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Euparkeria capensis Taxon name last Modified on 08/18/23	14	192	192	0	69	0	0	2
[2] † Ornithosuchus longidens Taxon name last Modified on 08/18/23	47	168	167	0	61	0	0	2
[3] † Herrerasaurus ischigualastensis Taxon name last Modified on 08/18/23	37	177	177	0	61	0	0	2
[4] † Scleromochlus taylori Taxon name last Modified on 08/18/23	88	124	124	0	63	0	0	2
[5] † Preondactylus buffarinii Taxon name last Modified on 08/18/23	86	132	132	0	57	0	0	2
[6] † Austriadactylus cristatus Taxon name last Modified on 08/18/23	88	143	143	0	44	0	0	2
[7] † Peteinosaurus zambellii Taxon name last Modified on 08/18/23	213	52	52	0	10	0	0	2
[8] † Caviramus schesaplanensis Taxon name last Modified on 08/18/23	230	39	38	0	7	0	0	2
[9] † Raeticodactylus filisurensis Taxon name last Modified on 08/18/23	94	143	143	0	38	0	0	2
[10] † Eudimorphodon cromptonellus Taxon name last Modified on 08/18/23	165	91	90	0	20	0	0	2
[11] † Eudimorphodon rosenfeldi Taxon name last Modified on 08/18/23	47	169	169	0	59	0	0	2
[12] † Eudimorphodon ranzii Taxon name last Modified on 08/18/23	58	169	169	0	48	0	0	2
[13] † Parapsicephalus purdoni Taxon name last Modified on 08/18/23	176	74	73	0	26	0	0	2
[14] † Dimorphodon macronyx Taxon name last Modified on 08/18/23	37	176	176	0	62	0	0	2
[15] † Dimorphodon koi Taxon name last Modified on 08/18/23	223	47	46	0	6	0	0	2
[16] † Dimorphodon hanseni Taxon name last Modified on 08/18/23	188	76	76	0	11	0	0	2
[17] † Dimorphodon jenkinsi Taxon name last Modified on 08/18/23	248	28	27	0	0	0	0	2
[18] † Dimorphodon weintraubi Taxon name last Modified on 08/18/23	204	55	54	0	17	0	0	2
[19] † Herbstosaurus pigmaeus Taxon name last Modified on 08/18/23	258	15	15	0	2	0	0	2
[20] † Campylognathoides zitteli Taxon name last Modified on 08/18/23	49	164	164	0	62	0	0	2
[21] † Campylognathoides liasicus Taxon name last Modified on 08/18/23	13	192	192	0	70	0	0	2
[22] † Scaphognathus crassirostris Taxon name last Modified on 08/18/23	27	183	182	0	66	0	0	2
[23] † Orientognathus chaoyangensis Taxon name last Modified on 08/18/23	108	121	119	0	48	0	0	2
[24] † Dorygnathus banthensis Taxon name last Modified on 08/18/23	6	203	202	0	67	0	0	2
[25] † Klobiodon rochei Taxon name last Modified on 08/18/23	224	42	42	0	9	0	0	2
[26] † Dolicorhamphus depressirostris Taxon name last Modified on 08/18/23	233	37	37	0	5	0	0	2
[27] † Dolicorhamphus bucklandi Taxon name last Modified on 08/18/23	234	34	34	0	7	0	0	2
[28] † Fenghuangopterus lii Taxon name last Modified on 08/18/23	90	135	134	0	51	0	0	2
[29] † Rhamphorhynchus muensteri Taxon name last Modified on 08/18/23	4	202	199	0	72	0	0	2
[30] † Nesodactylus hesperius Taxon name last Modified on 08/18/23	204	55	55	0	16	0	0	2
[31] † Cacibupteryx caribensis Taxon name last Modified on 08/18/23	178	77	76	0	21	0	0	2
[32] † Qinglongopterus guoi Taxon name last Modified on 08/18/23	104	124	124	0	47	0	0	2
[33] † Harpactognathus gentryii Taxon name last Modified on 08/18/23	217	53	52	0	6	0	0	2
[34] † Angustinaripterus longicephalus Taxon name last Modified on 08/18/23	145	106	106	0	24	0	0	2
[35] † Sericipterus wucaiwanensis Taxon name last Modified on 08/18/23	123	126	126	0	26	0	0	2
[36] † Sordes pilosus Taxon name last Modified on 08/18/23	11	195	194	0	70	0	0	2
[37] † Pterorhynchus wellnhoferi Taxon name last Modified on 08/18/23	76	145	145	0	54	0	0	2
[38] † Kunpengopterus sinensis Taxon name last Modified on 08/18/23	77	144	144	0	54	0	0	2
[39] † Wukongopterus lii Taxon name last Modified on 08/18/23	112	117	117	0	46	0	0	2
[40] † Darwinopterus zhengi Taxon name last Modified on 08/18/23	207	46	46	0	22	0	0	2
[41] † Darwinopterus linglongtaensis Taxon name last Modified on 08/18/23	55	161	161	0	59	0	0	2
[42] † Darwinopterus robustodens Taxon name last Modified on 08/18/23	47	167	167	0	61	0	0	2
[43] † Archaeoistiodactylus linglongtaensis Taxon name last Modified on 08/18/23	171	80	80	0	24	0	0	2
[44] † Darwinopterus modularis Taxon name last Modified on 08/18/23	44	169	169	0	62	0	0	2
[45] † Changchengopterus pani Taxon name last Modified on 08/18/23	206	49	49	0	20	0	0	2
[46] † Dendrorhynchoides curvidentatus Taxon name last Modified on 08/18/23	117	108	108	0	50	0	0	2
[47] † Luopterus mutoudengensis Taxon name last Modified on 08/18/23	125	102	101	0	49	0	0	2
[48] † Batrachognathus volans Taxon name last Modified on 08/18/23	74	136	136	0	65	0	0	2
[49] † Jeholopterus ningchengensis Taxon name last Modified on 08/18/23	59	142	142	0	74	0	0	2
[50] † Anurognathus ammoni Taxon name last Modified on 08/18/23	51	148	148	0	76	0	0	2
[51] † Kryptodrakon progenitor Taxon name last Modified on 08/18/23	257	16	16	0	2	0	0	2
[52] † Painten pterodactyloid Taxon name last Modified on 08/18/23	39	167	167	0	69	0	0	2
[53] † Cuspicephalus scarfi Taxon name last Modified on 08/18/23	175	76	76	0	24	0	0	2
[54] † Germanodactylus rhamphastinus Taxon name last Modified on 08/18/23	61	157	157	0	57	0	0	2
[55] † Germanodactylus cristatus Taxon name last Modified on 08/18/23	43	169	169	0	63	0	0	2
[56] † Pterodactylus antiquus Taxon name last Modified on 08/18/23	19	183	183	0	73	0	0	2
[57] † Normannognathus wellnhoferi Taxon name last Modified on 08/18/23	209	57	57	0	9	0	0	2
[58] † Aurorazhdarcho primordius Taxon name last Modified on 08/18/23	206	46	46	0	23	0	0	2
[59] † Cycnorhamphus suevicus Taxon name last Modified on 08/18/23	31	182	181	0	63	0	0	2
[60] † Pterodactylus micronyx Taxon name last Modified on 08/18/23	15	185	185	0	75	0	0	2
[61] † Liaodactylus primus Taxon name last Modified on 08/18/23	137	102	102	0	36	0	0	2
[62] † Ctenochasma elegans Taxon name last Modified on 08/18/23	31	178	177	0	67	0	0	2
[63] † Pterodaustro guinazui Taxon name last Modified on 08/18/23	36	170	170	0	69	0	0	2
[64] † Beipiaopterus chenianus Taxon name last Modified on 08/18/23	209	39	39	0	27	0	0	2
[65] † Gegepterus changi Taxon name last Modified on 08/18/23	129	109	108	0	38	0	0	2
[66] † Kepodactylus insperatus Taxon name last Modified on 08/18/23	242	30	30	0	3	0	0	2
[67] † Elanodactylus prolatus Taxon name last Modified on 08/18/23	188	67	67	0	20	0	0	2
[68] † Moganopterus zhuiana Taxon name last Modified on 08/18/23	146	97	97	0	32	0	0	2
[69] † Feilongus youngi Taxon name last Modified on 08/18/23	127	113	113	0	35	0	0	2
[70] † Ardeadactylus longicollum Taxon name last Modified on 08/18/23	30	179	179	0	66	0	0	2
[71] † Huanhepterus quingyangensis Taxon name last Modified on 03/07/23	157	90	90	0	28	0	0	2
[72] † Plataleorhynchus streptophorodon Taxon name last Modified on 08/18/23	233	33	33	0	9	0	0	2
[73] † Gnathosaurus macrurus Taxon name last Modified on 08/18/23	224	42	42	0	9	0	0	2
[74] † Gnathosaurus subulatus Taxon name last Modified on 08/18/23	139	104	104	0	32	0	0	2
[75] † Haopterus gracilis Taxon name last Modified on 08/18/23	100	119	119	0	56	0	0	2
[76] † Ornithostoma sedgwicki Taxon name last Modified on 08/18/23	227	15	15	0	33	0	0	2
[77] † Volgadraco bogolubovi Taxon name last Modified on 08/18/23	218	28	28	0	29	0	0	2
[78] † Tethydraco regalis Taxon name last Modified on 08/18/23	262	12	12	0	1	0	0	2
[79] † Pteranodon sternbergi Taxon name last Modified on 08/18/23	90	105	105	0	80	0	0	2
[80] † Pteranodon longiceps Taxon name last Modified on 08/18/23	17	174	174	0	84	0	0	2
[81] † Alamodactylus byrdi Taxon name last Modified on 08/18/23	259	15	15	0	1	0	0	2
[82] † Cretornis hlavaci Taxon name last Modified on 08/18/23	257	18	18	0	0	0	0	2
[83] † Simurghia robusta Taxon name last Modified on 08/18/23	269	5	5	0	1	0	0	2
[84] † Alcione elainus Taxon name last Modified on 08/18/23	231	37	37	0	7	0	0	2
[85] † Muzquizopteryx coahuilensis Taxon name last Modified on 08/18/23	127	90	90	0	58	0	0	2
[86] † Nyctosaurus grandis Taxon name last Modified on 08/18/23	238	28	28	0	9	0	0	2
[87] † Nyctosaurus lamegoi Taxon name last Modified on 08/18/23	269	5	5	0	1	0	0	2
[88] † Nyctosaurus nanus Taxon name last Modified on 08/18/23	247	24	24	0	4	0	0	2
[89] † Nyctosaurus gracilis Taxon name last Modified on 08/18/23	19	166	166	0	90	0	0	2
[90] † Serradraco sagittirostris Taxon name last Modified on 08/18/23	253	20	20	0	2	0	0	2
[91] † Aussiedraco molnari Taxon name last Modified on 08/18/23	237	32	32	0	6	0	0	2
[92] † Hongshanopterus lacustris Taxon name last Modified on 08/18/23	180	75	74	0	21	0	0	2
[93] † Targaryendraco wiedenrothi Taxon name last Modified on 08/18/23	231	37	37	0	7	0	0	2
[94] † Lonchodectes compressirostris Taxon name last Modified on 08/18/23	214	46	46	0	15	0	0	2
[95] † Ikrandraco avatar Taxon name last Modified on 08/18/23	88	137	137	0	50	0	0	2
[96] † Ikrandraco machaerorhynchus Taxon name last Modified on 08/18/23	249	24	24	0	2	0	0	2
[97] † Lonchodraco giganteus Taxon name last Modified on 08/18/23	188	72	72	0	15	0	0	2
[98] † Lonchodraco microdon Taxon name last Modified on 08/18/23	246	20	20	0	9	0	0	2
[99] † Lonchodraco denticulatus Taxon name last Modified on 08/18/23	249	24	24	0	2	0	0	2
[100] † Nurhachius ignaciobritoi Taxon name last Modified on 08/18/23	55	155	154	0	66	0	0	2
[101] † Liaoxipterus brachyognathus Taxon name last Modified on 08/18/23	234	31	31	0	10	0	0	2
[102] † Istiodactylus sinensis Taxon name last Modified on 08/18/23	88	125	125	0	62	0	0	2
[103] † Istiodactylus latidens Taxon name last Modified on 08/18/23	76	144	144	0	55	0	0	2
[104] † Pterodactylus polyodon Taxon name last Modified on 08/18/23	238	27	27	0	10	0	0	2
[105] † Zhenyuanopterus longirostris Taxon name last Modified on 08/18/23	83	137	137	0	55	0	0	2
[106] † Boreopterus giganticus Taxon name last Modified on 08/18/23	142	90	90	0	43	0	0	2
[107] † Boreopterus cuiae Taxon name last Modified on 08/18/23	112	107	106	0	57	0	0	2
[108] † Hamipterus tianshanensis Taxon name last Modified on 08/18/23	47	176	176	0	52	0	0	2
[109] † Brasileodactylus araripensis Taxon name last Modified on 08/18/23	232	37	37	0	6	0	0	2
[110] † Barbosania gracilirostris Taxon name last Modified on 08/18/23	103	123	123	0	49	0	0	2
[111] † Cearadactylus atrox Taxon name last Modified on 08/18/23	142	97	97	0	36	0	0	2
[112] † Guidraco venator Taxon name last Modified on 08/18/23	140	98	98	0	37	0	0	2
[113] † Ludodactylus colorhinus Taxon name last Modified on 08/18/23	251	23	23	0	1	0	0	2
[114] † Ludodactylus sibbicki Taxon name last Modified on 08/18/23	128	110	110	0	37	0	0	2
[115] † Aetodactylus halli Taxon name last Modified on 08/18/23	204	51	51	0	20	0	0	2
[116] † Camposipterus nasutus Taxon name last Modified on 08/18/23	237	29	29	0	9	0	0	2
[117] † Cimoliopterus dunni Taxon name last Modified on 08/18/23	231	40	40	0	4	0	0	2
[118] † Cimoliopterus cuvieri Taxon name last Modified on 08/18/23	231	40	40	0	4	0	0	2
[119] † Aerodraco sedgwickii Taxon name last Modified on 08/18/23	243	28	28	0	4	0	0	2
[120] † Liaoningopterus gui Taxon name last Modified on 08/18/23	160	96	95	0	20	0	0	2
[121] † Anhanguera robustus Taxon name last Modified on 08/18/23	112	135	135	0	28	0	0	2
[122] † Anhanguera piscator Taxon name last Modified on 08/18/23	35	189	189	0	51	0	0	2
[123] † Anhanguera blittersdorffi Taxon name last Modified on 08/18/23	129	114	114	0	32	0	0	2
[124] † Tropeognathus mesembrinus Taxon name last Modified on 08/18/23	123	117	117	0	35	0	0	2
[125] † Ornithocheirus platystomus Taxon name last Modified on 08/18/23	251	24	24	0	0	0	0	2
[126] † Ornithocheirus simus Taxon name last Modified on 08/18/23	246	28	28	0	1	0	0	2
[127] † Siroccopteryx moroccensis Taxon name last Modified on 08/18/23	238	34	34	0	3	0	0	2
[128] † Coloborhynchus capito Taxon name last Modified on 08/18/23	246	26	26	0	3	0	0	2
[129] † Coloborhynchus wadleighi Taxon name last Modified on 08/18/23	238	34	34	0	3	0	0	2
[130] † Coloborhynchus clavirostris Taxon name last Modified on 08/18/23	245	29	29	0	1	0	0	2
[131] † Eopteranodon lii Taxon name last Modified on 08/18/23	122	92	92	0	61	0	0	2
[132] † Huaxiapterus jii Taxon name last Modified on 08/18/23	107	101	101	0	67	0	0	2
[133] † Sinopterus dongi Taxon name last Modified on 08/18/23	64	137	137	0	74	0	0	2
[134] † Sinopterus benxiensis Taxon name last Modified on 08/18/23	116	99	99	0	60	0	0	2
[135] † Sinopterus corollatus Taxon name last Modified on 08/18/23	109	97	97	0	69	0	0	2
[136] † Bakonydraco galaczi Taxon name last Modified on 08/18/23	195	50	50	0	30	0	0	2
[137] † Keresdrakon vilsoni Taxon name last Modified on 08/18/23	132	98	97	0	46	0	0	2
[138] † Caiuajara dobruskii Taxon name last Modified on 08/18/23	75	139	138	0	62	0	0	2
[139] † Tupandactylus navigans Taxon name last Modified on 08/18/23	172	61	61	0	42	0	0	2
[140] † Tupandactylus imperator Taxon name last Modified on 08/18/23	151	75	75	0	49	0	0	2
[141] † Vectidraco daisymorrisae Taxon name last Modified on 08/18/23	262	10	10	0	3	0	0	2
[142] † Europejara olcadesorum Taxon name last Modified on 08/18/23	196	44	44	0	35	0	0	2
[143] † Tapejara wellnhoferi Taxon name last Modified on 08/18/23	24	175	175	0	76	0	0	2
[144] † Microtuban altivolans Taxon name last Modified on 08/18/23	235	28	28	0	12	0	0	2
[145] † Noripterus complicidens Taxon name last Modified on 08/18/23	147	95	95	0	33	0	0	2
[146] † Noripterus parvus Taxon name last Modified on 08/18/23	51	172	172	0	52	0	0	2
[147] † Domeykodactylus ceciliae Taxon name last Modified on 08/18/23	240	31	31	0	4	0	0	2
[148] † Dsungaripterus weii Taxon name last Modified on 08/18/23	33	185	185	0	57	0	0	2
[149] † Tupuxuara leonardii Taxon name last Modified on 08/18/23	47	152	152	0	76	0	0	2
[150] † Tupuxuara longicristatus Taxon name last Modified on 08/18/23	205	36	36	0	34	0	0	2
[151] † Thalassodromeus sethi Taxon name last Modified on 08/18/23	131	96	96	0	48	0	0	2
[152] † Lacusovagus magnificens Taxon name last Modified on 08/18/23	214	26	26	0	35	0	0	2
[153] † Xericeps curvirostris Taxon name last Modified on 08/18/23	232	18	18	0	25	0	0	2
[154] † Alanqa saharica Taxon name last Modified on 08/18/23	229	23	23	0	23	0	0	2
[155] † Argentinadraco barrealensis Taxon name last Modified on 08/18/23	230	22	22	0	23	0	0	2
[156] † Aerotitan sudamericanus Taxon name last Modified on 08/18/23	235	15	15	0	25	0	0	2
[157] † Eoazhdarcho liaoxiensis Taxon name last Modified on 08/18/23	168	59	59	0	48	0	0	2
[158] † Shenzhoupterus chaoyangensis Taxon name last Modified on 08/18/23	117	88	88	0	70	0	0	2
[159] † Jidapterus edentus Taxon name last Modified on 08/18/23	66	127	127	0	82	0	0	2
[160] † Chaoyangopterus zhangi Taxon name last Modified on 08/18/23	108	96	96	0	71	0	0	2
[161] † Radiodactylus langstoni Taxon name last Modified on 08/18/23	262	13	13	0	0	0	0	2
[162] † Montanazhdarcho minor Taxon name last Modified on 08/18/23	188	55	55	0	32	0	0	2
[163] † Azhdarcho lancicollis Taxon name last Modified on 08/18/23	138	90	90	0	47	0	0	2
[164] † Albadraco tharmisensis Taxon name last Modified on 08/18/23	219	27	27	0	29	0	0	2
[165] † Mistralazhdarcho maggii Taxon name last Modified on 08/18/23	200	38	38	0	37	0	0	2
[166] † Leptostomia begaaensis Taxon name last Modified on 08/18/23	221	26	26	0	28	0	0	2
[167] † Apatorhamphus gyrostega Taxon name last Modified on 08/18/23	207	30	30	0	38	0	0	2
[168] † Aralazhdarcho bostobensis Taxon name last Modified on 08/18/23	233	20	20	0	22	0	0	2
[169] † Phosphatodraco mauritanicus Taxon name last Modified on 08/18/23	263	11	11	0	1	0	0	2
[170] † Eurazhdarcho langendorfensis Taxon name last Modified on 08/18/23	261	12	12	0	2	0	0	2
[171] † Zhejiangopterus linhaiensis Taxon name last Modified on 08/18/23	74	116	116	0	85	0	0	2
[172] † Cryodrakon boreas Taxon name last Modified on 08/18/23	237	34	34	0	4	0	0	2
[173] † Wellnhopterus brevirostris Taxon name last Modified on 08/18/23	178	59	59	0	38	0	0	2
[174] † Hatzegopteryx thambema Taxon name last Modified on 08/18/23	210	39	39	0	26	0	0	2
[175] † Arambourgiania philadelphiae Taxon name last Modified on 08/18/23	264	10	10	0	1	0	0	2
[176] † Quetzalcoatlus lawsoni Taxon name last Modified on 08/18/23	38	160	158	0	79	0	0	2
[177] † Quetzalcoatlus northropi Taxon name last Modified on 08/18/23	236	34	34	0	5	0	0	2
[178] † Dearc sgiathanach Taxon name last Modified on 08/18/23	252	16	16	0	7	0	0	2
[179] † Tacuadactylus luciae Taxon name last Modified on 03/07/23	265	10	10	0	0	0	0	2
[180] † Lusognathus almadrava Taxon name last Modified on 03/07/23	248	25	25	0	2	0	0	2

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Project overview	426
Media views	721	Media search (608 views); M823739 (113 views);
Matrix views	250	Matrix landing page (234 views); A new gnathosaurine from the Late Jurassic of Portugal (16 views);
Specimen list	515
Taxon list	2854
Bibliography	152
Views for media list	234

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	9
Matrix downloads	9	A new gnathosaurine from the Late Jurassic of Portugal (9 downloads);