Project 4138: B. Bomfleur, G. W. Grimm, S. McLoughlin. 2017. The fossil Osmundales (Royal Ferns)—a phylogenetic network analysis, revised taxonomy, and evolutionary classification of anatomically preserved trunks and rhizomes. PeerJ. 5:e3433.
Abstract
The Osmundales (Royal Fern order) originated in the late Paleozoic and is the most ancient surviving lineage of leptosporangiate ferns. In contrast to its low diversity today (less than 20 species in six genera), it has the richest fossil record of any extant group of ferns. The structurally preserved trunks and rhizomes alone are referable to more than 100 fossil species that are classified in up to 20 genera, four subfamilies, and two families. This diverse fossil record constitutes an exceptional source of information on the evolutionary history of the group from the Permian to the present. However, inconsistent terminology, varying formats of description, and the general lack of a uniform taxonomic concept renders this wealth of information poorly accessible. To this end, we provide a comprehensive review of the diversity of structural features of osmundalean axes under a standardized, descriptive terminology. A novel morphological character matrix with 45 anatomical characters scored for 15 extant species and for 114 fossil operational units (species or specimens) is analysed using networks in order to establish systematic relationships among fossil and extant Osmundales rooted in axis anatomy. The results lead us to propose an evolutionary classification for fossil Osmundales and a revised, standardized taxonomy for all taxa down to the rank of (sub)genus. We introduce several nomenclatural novelties: (1) a new subfamily Itopsidemoideae (Guaireaceae) is established to contain Itopsidema, Donwelliacaulis, and Tiania; (2) the thamnopteroid genera Zalesskya, Iegosigopteris, and Petcheropteris are all considered synonymous with Thamnopteris; (3) 12 species of Millerocaulis and Ashicaulis are assigned to modern genera (tribe Osmundeae); (4) the hitherto enigmatic Aurealcaulis is identified as an extinct subgenus of Plenasium; and (5) the poorly known Osmundites tuhajkulensis is assigned to Millerocaulis. In addition, we consider Millerocaulis stipabonettiorum a possible member of Palaeosmunda and Millerocaulis estipularis as probably constituting the earliest representative of the (Todea-)Leptopteris lineage (subtribe Todeinae) of modern Osmundoideae.Read the article »
Article DOI: 10.7717/peerj.3433
Project DOI: 10.7934/P4138, http://dx.doi.org/10.7934/P4138
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 4823 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 45 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 112 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:45/0 |
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MorphoBank Project 4138
MorphoBank Project 4138
- Creation Date:
01 December 2021 - Publication Date:
29 January 2022 - Matrix downloads: 1
Authors' Institutions
- Swedish Museum of Natural History (Naturhistoriska riksmuseet) Stockholm
- Universitaet Muenster (Westfälische Wilhelms-Universität Münster)
- University of Vienna
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| MorphoBank Curator Project Administrator | 131 | 1 | 1 | 45 | 0 | 0 | 4823 (4823, 0, 0) | 0 | 0 | 0 | ||||
| Benjamin Bomfleur Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| MorphoBank Curator Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M27717' (131 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] † Anomorrhoea fischeri Taxon name last Modified on 12/03/21 | 31 | 14 | 14 | 0 | 0 | 0 | 0 | 3 |
| [2] † Bathypteris rhomboidea Taxon name last Modified on 12/03/21 | 17 | 29 | 28 | 0 | 0 | 0 | 0 | 3 |
| [3] † Chasmatopteris principalis Taxon name last Modified on 12/03/21 | 12 | 33 | 32 | 0 | 0 | 0 | 0 | 3 |
| [4] † Grammatopteris baldaufii Taxon name last Modified on 12/03/21 | 15 | 33 | 30 | 0 | 0 | 0 | 0 | 3 |
| [5] † Grammatopteris rigollotii Taxon name last Modified on 12/03/21 | 16 | 29 | 29 | 0 | 0 | 0 | 0 | 3 |
| [6] † Grammatopteris freitasii Taxon name last Modified on 12/03/21 | 15 | 31 | 30 | 0 | 0 | 0 | 0 | 3 |
| [7] † Guairea brasiliensis Taxon name last Modified on 12/03/21 | 21 | 27 | 24 | 0 | 0 | 0 | 0 | 3 |
| [8] † Guairea carnierii Taxon name last Modified on 12/03/21 | 21 | 27 | 24 | 0 | 0 | 0 | 0 | 3 |
| [9] † Guairea millerii Taxon name last Modified on 12/03/21 | 20 | 26 | 25 | 0 | 0 | 0 | 0 | 3 |
| [10] † Palaeosmunda playfordii Taxon name last Modified on 12/03/21 | 8 | 38 | 36 | 0 | 0 | 0 | 0 | 3 |
| [11] † Palaeosmunda williamsii Taxon name last Modified on 12/03/21 | 6 | 42 | 38 | 0 | 0 | 0 | 0 | 3 |
| [12] † Rastropteris pinquanensis Taxon name last Modified on 12/03/21 | 16 | 29 | 29 | 0 | 0 | 0 | 0 | 3 |
| [13] † Shuichengella primitiva Taxon name last Modified on 12/03/21 | 19 | 26 | 26 | 0 | 0 | 0 | 0 | 3 |
| [14] † Thamnopteris diploxylon Taxon name last Modified on 12/03/21 | 25 | 22 | 20 | 0 | 0 | 0 | 0 | 3 |
| [15] † Thamnopteris gracilis Taxon name last Modified on 12/03/21 | 24 | 23 | 21 | 0 | 0 | 0 | 0 | 3 |
| [16] † Thamnopteris gwynnevaughanii Taxon name last Modified on 12/03/21 | 12 | 35 | 33 | 0 | 0 | 0 | 0 | 3 |
| [17] † Thamnopteris javorskii Taxon name last Modified on 12/03/21 | 10 | 38 | 32 | 0 | 0 | 0 | 0 | 3 |
| [18] † Thamnopteris kazanensis Taxon name last Modified on 12/03/21 | 25 | 22 | 20 | 0 | 0 | 0 | 0 | 3 |
| [19] † Thamnopteris kidstonii Taxon name last Modified on 12/03/21 | 7 | 42 | 38 | 0 | 0 | 0 | 0 | 3 |
| [20] † Thamnopteris schlechtendalii Taxon name last Modified on 12/03/21 | 9 | 38 | 36 | 0 | 0 | 0 | 0 | 3 |
| [21] † Thamnopteris splendida Taxon name last Modified on 12/03/21 | 13 | 32 | 32 | 0 | 0 | 0 | 0 | 3 |
| [22] † Thamnopteris uralica Taxon name last Modified on 12/03/21 | 24 | 23 | 21 | 0 | 0 | 0 | 0 | 3 |
| [23] † Tiania yunnanensis Taxon name last Modified on 12/03/21 | 20 | 26 | 25 | 0 | 0 | 0 | 0 | 3 |
| [24] † Zhongmingella plenasioides Taxon name last Modified on 12/03/21 | 24 | 23 | 21 | 0 | 0 | 0 | 0 | 3 |
| [25] † Osmunda beardmorensis Taxon name last Modified on 12/03/21 | 7 | 41 | 38 | 0 | 0 | 0 | 0 | 3 |
| [26] † Millerocaulis herbstii Taxon name last Modified on 12/03/21 | 17 | 29 | 28 | 0 | 0 | 0 | 0 | 3 |
| [27] † Millerocaulis lutziae Taxon name last Modified on 12/03/21 | 18 | 27 | 27 | 0 | 0 | 0 | 0 | 3 |
| [28] † Millerocaulis woolfei Taxon name last Modified on 12/03/21 | 7 | 39 | 38 | 0 | 0 | 0 | 0 | 3 |
| [29] † Donwelliacaulis chlouberii Taxon name last Modified on 12/03/21 | 19 | 26 | 26 | 0 | 0 | 0 | 0 | 3 |
| [30] † Itopsidema vancleavei Taxon name last Modified on 12/03/21 | 18 | 27 | 27 | 0 | 0 | 0 | 0 | 3 |
| [31] † Osmunda indentata Taxon name last Modified on 12/03/21 | 3 | 42 | 42 | 0 | 0 | 0 | 0 | 3 |
| [32] † Millerocaulis stipabonettiorum Taxon name last Modified on 12/03/21 | 18 | 27 | 27 | 0 | 0 | 0 | 0 | 3 |
| [33] † Millerocaulis tuhajkulensis Taxon name last Modified on 12/03/21 | 17 | 28 | 27 | 0 | 0 | 0 | 0 | 3 |
| [34] † Millerocaulis aucklandicus Taxon name last Modified on 12/03/21 | 7 | 40 | 37 | 0 | 0 | 0 | 0 | 3 |
| [35] † Millerocaulis beipiaoensis Taxon name last Modified on 12/03/21 | 7 | 38 | 38 | 0 | 0 | 0 | 0 | 3 |
| [36] † Millerocaulis broganii Taxon name last Modified on 12/03/21 | 5 | 40 | 40 | 0 | 0 | 0 | 0 | 3 |
| [37] † Millerocaulis gibbianus Taxon name last Modified on 12/03/21 | 6 | 40 | 39 | 0 | 0 | 0 | 0 | 3 |
| [38] † Millerocaulis hebeiensis Taxon name last Modified on 12/03/21 | 8 | 37 | 37 | 0 | 0 | 0 | 0 | 3 |
| [39] † Millerocaulis macromedullosus Taxon name last Modified on 12/03/21 | 9 | 36 | 36 | 0 | 0 | 0 | 0 | 3 |
| [40] † Millerocaulis patagonicus Taxon name last Modified on 12/03/21 | 7 | 39 | 38 | 0 | 0 | 0 | 0 | 3 |
| [41] † Millerocaulis richmondii Taxon name last Modified on 12/03/21 | 5 | 40 | 40 | 0 | 0 | 0 | 0 | 3 |
| [42] † Millerocaulis santaecrucis Taxon name last Modified on 12/03/21 | 14 | 31 | 31 | 0 | 0 | 0 | 0 | 3 |
| [43] † Millerocaulis spinksii Taxon name last Modified on 12/03/21 | 8 | 38 | 37 | 0 | 0 | 0 | 0 | 3 |
| [44] † Millerocaulis swanensis Taxon name last Modified on 12/03/21 | 5 | 41 | 40 | 0 | 0 | 0 | 0 | 3 |
| [45] † Millerocaulis wadei Taxon name last Modified on 12/03/21 | 6 | 42 | 39 | 0 | 0 | 0 | 0 | 3 |
| [46] † Millerocaulis websteri Taxon name last Modified on 12/03/21 | 7 | 39 | 38 | 0 | 0 | 0 | 0 | 3 |
| [47] † Millerocaulis wrightii Taxon name last Modified on 12/03/21 | 7 | 39 | 38 | 0 | 0 | 0 | 0 | 3 |
| [48] † Grammatocaulis donponii Taxon name last Modified on 12/03/21 | 12 | 33 | 32 | 0 | 0 | 0 | 0 | 3 |
| [49] † Lunea jonesii Taxon name last Modified on 12/03/21 | 10 | 35 | 35 | 0 | 0 | 0 | 0 | 3 |
| [50] † Millerocaulis chubutensis Taxon name last Modified on 12/03/21 | 10 | 35 | 35 | 0 | 0 | 0 | 0 | 3 |
| [51] † Millerocaulis donponii Taxon name last Modified on 12/03/21 | 9 | 36 | 36 | 0 | 0 | 0 | 0 | 3 |
| [52] † Millerocaulis dunlopii Taxon name last Modified on 12/03/21 | 10 | 36 | 35 | 0 | 0 | 0 | 0 | 3 |
| [53] † Millerocaulis juandahensis Taxon name last Modified on 12/03/21 | 9 | 36 | 35 | 0 | 0 | 0 | 0 | 3 |
| [54] † Millerocaulis limewoodensis Taxon name last Modified on 12/03/21 | 9 | 36 | 36 | 0 | 0 | 0 | 0 | 3 |
| [55] † Osmunda preosmunda Taxon name last Modified on 12/03/21 | 5 | 40 | 40 | 0 | 0 | 0 | 0 | 3 |
| [56] † Osmunda sinica Taxon name last Modified on 12/03/21 | 3 | 42 | 42 | 0 | 0 | 0 | 0 | 3 |
| [57] † Osmunda chengii Taxon name last Modified on 12/03/21 | 1 | 44 | 44 | 0 | 0 | 0 | 0 | 3 |
| [58] † Osmunda liaoningensis Taxon name last Modified on 12/03/21 | 1 | 47 | 44 | 0 | 0 | 0 | 0 | 3 |
| [59] † Osmunda pulchella Taxon name last Modified on 12/03/21 | 2 | 45 | 43 | 0 | 0 | 0 | 0 | 3 |
| [60] † Osmunda plumites Taxon name last Modified on 12/03/21 | 2 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [61] † Osmunda wangii Taxon name last Modified on 12/03/21 | 2 | 44 | 43 | 0 | 0 | 0 | 0 | 3 |
| [62] † Osmundacaulis andrewii Taxon name last Modified on 12/03/21 | 7 | 40 | 38 | 0 | 0 | 0 | 0 | 3 |
| [63] † Osmundacaulis griggsii Taxon name last Modified on 12/03/21 | 5 | 43 | 40 | 0 | 0 | 0 | 0 | 3 |
| [64] † Osmundacaulis hoskingii var hoskingii Taxon name last Modified on 12/03/21 | 6 | 39 | 39 | 0 | 0 | 0 | 0 | 3 |
| [65] † Osmundacaulis janae Taxon name last Modified on 12/03/21 | 30 | 16 | 15 | 0 | 0 | 0 | 0 | 3 |
| [66] † Osmundacaulis jonesii Taxon name last Modified on 12/03/21 | 5 | 41 | 40 | 0 | 0 | 0 | 0 | 3 |
| [67] † Osmundacaulis lemonii Taxon name last Modified on 12/03/21 | 8 | 38 | 37 | 0 | 0 | 0 | 0 | 3 |
| [68] † Osmundacaulis nerii Taxon name last Modified on 12/03/21 | 6 | 41 | 39 | 0 | 0 | 0 | 0 | 3 |
| [69] † Osmundacaulis pruchnickii Taxon name last Modified on 12/03/21 | 5 | 42 | 40 | 0 | 0 | 0 | 0 | 3 |
| [70] † Osmundacaulis richmondii Taxon name last Modified on 12/03/21 | 6 | 40 | 39 | 0 | 0 | 0 | 0 | 3 |
| [71] † Osmundacaulis tasmanensis Taxon name last Modified on 12/03/21 | 7 | 38 | 38 | 0 | 0 | 0 | 0 | 3 |
| [72] † Osmundacaulis tehuelchensis Taxon name last Modified on 12/03/21 | 6 | 41 | 39 | 0 | 0 | 0 | 0 | 3 |
| [73] † Osmunda johnstonii Taxon name last Modified on 12/03/21 | 1 | 44 | 44 | 0 | 0 | 0 | 0 | 3 |
| [74] † Osmundacaulis bamfordiae Taxon name last Modified on 12/03/21 | 11 | 34 | 34 | 0 | 0 | 0 | 0 | 3 |
| [75] † Osmundacaulis tidwellii Taxon name last Modified on 12/03/21 | 9 | 37 | 36 | 0 | 0 | 0 | 0 | 3 |
| [76] † Osmundacaulis zululandensis Taxon name last Modified on 12/03/21 | 8 | 38 | 37 | 0 | 0 | 0 | 0 | 3 |
| [77] † Gen indet estipularis Taxon name last Modified on 12/03/21 | 13 | 32 | 30 | 0 | 0 | 0 | 0 | 3 |
| [78] † Millerocaulis amarjolensis Taxon name last Modified on 12/03/21 | 7 | 38 | 38 | 0 | 0 | 0 | 0 | 3 |
| [79] † Millerocaulis australis Taxon name last Modified on 12/03/21 | 8 | 37 | 37 | 0 | 0 | 0 | 0 | 3 |
| [80] † Millerocaulis indica Taxon name last Modified on 12/03/21 | 11 | 34 | 33 | 0 | 0 | 0 | 0 | 3 |
| [81] † Millerocaulis guptai Taxon name last Modified on 12/03/21 | 10 | 35 | 35 | 0 | 0 | 0 | 0 | 3 |
| [82] † Millerocaulis kolbei Taxon name last Modified on 12/03/21 | 7 | 40 | 37 | 0 | 0 | 0 | 0 | 3 |
| [83] † Millerocaulis livingstonensis Taxon name last Modified on 12/03/21 | 7 | 38 | 38 | 0 | 0 | 0 | 0 | 3 |
| [84] † Millerocaulis rajmahalensis Taxon name last Modified on 12/03/21 | 5 | 40 | 40 | 0 | 0 | 0 | 0 | 3 |
| [85] † Millerocaulis sahnii Taxon name last Modified on 12/03/21 | 8 | 37 | 37 | 0 | 0 | 0 | 0 | 3 |
| [86] † Osmunda tekelili Taxon name last Modified on 12/03/21 | 6 | 40 | 39 | 0 | 0 | 0 | 0 | 3 |
| [87] † Osmunda burgii Taxon name last Modified on 12/03/21 | 5 | 40 | 38 | 0 | 0 | 0 | 0 | 3 |
| [88] † Osmunda dakotensis Taxon name last Modified on 12/03/21 | 7 | 38 | 37 | 0 | 0 | 0 | 0 | 3 |
| [89] † Osmunda nebraskensis Taxon name last Modified on 12/03/21 | 7 | 39 | 38 | 0 | 0 | 0 | 0 | 3 |
| [90] † Osmunda embreei Taxon name last Modified on 12/03/21 | 3 | 44 | 42 | 0 | 0 | 0 | 0 | 3 |
| [91] † Osmunda cinnamomea Cretaceous Canada Taxon name last Modified on 12/03/21 | 2 | 43 | 42 | 0 | 0 | 0 | 0 | 3 |
| [92] † Osmunda kidstonii Taxon name last Modified on 12/03/21 | 31 | 14 | 14 | 0 | 0 | 0 | 0 | 3 |
| [93] † Osmundacaulis atherstonei Taxon name last Modified on 12/03/21 | 10 | 37 | 35 | 0 | 0 | 0 | 0 | 3 |
| [94] † Osmundacaulis natalensis Taxon name last Modified on 12/03/21 | 9 | 38 | 36 | 0 | 0 | 0 | 0 | 3 |
| [95] † Osmundacaulis skidegatensis Taxon name last Modified on 12/03/21 | 6 | 41 | 39 | 0 | 0 | 0 | 0 | 3 |
| [96] † Osmundacaulis whittlesii Taxon name last Modified on 12/03/21 | 8 | 38 | 37 | 0 | 0 | 0 | 0 | 3 |
| [97] † Todea tidwellii Taxon name last Modified on 12/03/21 | 11 | 34 | 33 | 0 | 0 | 0 | 0 | 3 |
| [98] † Osmunda bransonii Taxon name last Modified on 12/03/21 | 8 | 38 | 37 | 0 | 0 | 0 | 0 | 3 |
| [99] † Osmunda moorei Taxon name last Modified on 12/03/21 | 6 | 39 | 37 | 0 | 0 | 0 | 0 | 3 |
| [100] † Osmunda crossii Taxon name last Modified on 12/03/21 | 8 | 38 | 33 | 0 | 0 | 0 | 0 | 3 |
| [101] † Osmunda arnoldii Taxon name last Modified on 12/03/21 | 2 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [102] † ALT1 osmunda dowkeri MILLER Taxon name last Modified on 12/03/21 | 3 | 43 | 42 | 0 | 0 | 0 | 0 | 3 |
| [103] † ALT2 osmunda dowkeri CHANDLER Taxon name last Modified on 12/03/21 | 2 | 44 | 42 | 0 | 0 | 0 | 0 | 3 |
| [104] † ALT2 osmunda chandleri Taxon name last Modified on 12/03/21 | 8 | 39 | 37 | 0 | 0 | 0 | 0 | 3 |
| [105] † Osmunda nathorstii Taxon name last Modified on 12/03/21 | 27 | 18 | 18 | 0 | 0 | 0 | 0 | 3 |
| [106] † Osmunda oregonensis Taxon name last Modified on 12/03/21 | 2 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [107] † Osmunda pluma Taxon name last Modified on 12/03/21 | 2 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [108] † Osmunda precinnamomea Taxon name last Modified on 12/03/21 | 2 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [109] † Osmunda cinnamomea Neogene USA Taxon name last Modified on 12/03/21 | 3 | 42 | 42 | 0 | 0 | 0 | 0 | 3 |
| [110] † Osmunda cinnamomea Neogene Japan Taxon name last Modified on 12/03/21 | 6 | 39 | 39 | 0 | 0 | 0 | 0 | 3 |
| [111] † Osmunda iliaensis Taxon name last Modified on 12/03/21 | 1 | 45 | 43 | 0 | 0 | 0 | 0 | 3 |
| [112] † Osmunda shimokawaensis Taxon name last Modified on 12/03/21 | 2 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [113] † Osmunda wehrii Taxon name last Modified on 12/03/21 | 1 | 44 | 44 | 0 | 0 | 0 | 0 | 3 |
| [114] Osmunda cinnamomea Taxon name last Modified on 12/05/21 | 2 | 44 | 43 | 0 | 0 | 0 | 0 | 3 |
| [115] Osmunda claytoniana Taxon name last Modified on 12/05/21 | 1 | 46 | 44 | 0 | 0 | 0 | 0 | 3 |
| [116] Osmunda banksiifolia Taxon name last Modified on 12/05/21 | 2 | 44 | 42 | 0 | 0 | 0 | 0 | 3 |
| [117] Osmunda bromeliifolia Taxon name last Modified on 12/05/21 | 1 | 44 | 44 | 0 | 0 | 0 | 0 | 3 |
| [118] Osmunda javanica Taxon name last Modified on 12/05/21 | 1 | 46 | 43 | 0 | 0 | 0 | 0 | 3 |
| [119] Osmunda vachellii Taxon name last Modified on 12/05/21 | 2 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [120] Osmunda japonica Taxon name last Modified on 12/05/21 | 1 | 44 | 44 | 0 | 0 | 0 | 0 | 3 |
| [121] Osmunda lancea Taxon name last Modified on 12/05/21 | 1 | 45 | 44 | 0 | 0 | 0 | 0 | 3 |
| [122] Osmunda regalis Taxon name last Modified on 12/05/21 | 1 | 47 | 41 | 0 | 0 | 0 | 0 | 3 |
| [123] Leptopteris fraseri Taxon name last Modified on 12/05/21 | 2 | 45 | 41 | 0 | 0 | 0 | 0 | 3 |
| [124] Leptopteris hymenophylloides Taxon name last Modified on 12/05/21 | 2 | 44 | 43 | 0 | 0 | 0 | 0 | 3 |
| [125] Leptopteris wilkesiana Taxon name last Modified on 12/05/21 | 2 | 44 | 43 | 0 | 0 | 0 | 0 | 3 |
| [126] Leptopteris superba Taxon name last Modified on 12/05/21 | 2 | 44 | 43 | 0 | 0 | 0 | 0 | 3 |
| [127] Todea barbara Taxon name last Modified on 12/05/21 | 3 | 42 | 42 | 0 | 0 | 0 | 0 | 3 |
| [128] Todea papuana Taxon name last Modified on 12/05/21 | 38 | 7 | 7 | 0 | 0 | 0 | 0 | 3 |
| [129] † Osmundacaulis hoskingii tabulatus Taxon name last Modified on 12/03/21 | 7 | 38 | 37 | 0 | 0 | 0 | 0 | 3 |
| [130] † Millerocaulis australis Vera2010 Taxon name last Modified on 12/03/21 | 11 | 36 | 33 | 0 | 0 | 0 | 0 | 3 |
| [131] † Millerocaulis zamunerae Taxon name last Modified on 12/03/21 | 5 | 41 | 39 | 0 | 0 | 0 | 0 | 3 |
Project downloads
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 15 | |
| Project downloads | 14 | |
| Matrix downloads | 1 | Morphological matrix from Bomfleur et al 2017 - (prepared by MorphoBank curators from original submission) (1 download); |