Project 4146: S. Pates, J. P. Botting, L. A. Muir, J. M. Wolfe. 2022. Ordovician opabiniid-like animals and the role of the proboscis in euarthropod head evolution. Nature Communications. 13 (1):null.
Specimen: † Mieridduryn bonniae (NMW:NMW.2021.3G.7:)
View: Lateral 2d compression

Abstract

A crucial step in the evolution of Euarthropoda (chelicerates, myriapods, pancrustaceans) was the transition between fossil groups that possessed frontal appendages innervated by the first segment of the brain (protocerebrum), and living groups with a protocerebral labrum and paired appendages innervated by the second brain segment (deutocerebrum). Appendage homologies between the groups are controversial. Here we describe two specimens of opabiniid-like euarthropods, each bearing an anterior proboscis (a fused protocerebral appendage), from the Middle Ordovician Castle Bank Biota, Wales, UK. Phylogenetic analyses support a paraphyletic grade of stem group euarthropods with fused protocerebral appendages and a posterior-facing mouth, as in the iconic Cambrian panarthropod Opabinia. These results suggest that the labrum may have reduced from an already-fused proboscis, rather than a pair of arthropodized appendages. If some shared features between the Castle Bank specimens and radiodonts are considered convergent rather than homologous, phylogenetic analyses retrieve them as opabiniids, substantially extending the geographic and temporal range of Opabiniidae.


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Article DOI: 10.1038/s41467-022-34204-w

Project DOI: 10.7934/P4146, http://dx.doi.org/10.7934/P4146
This project containsMatrices
  • 1 Media
  • 8 Matrices
  • 62 Taxa
  • 1 Specimen
  • 133 Characters
Total size of project's media files: 407.96k

Download Project SDD File
Total scored cells: 49056
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 1038
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 307
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:1038/0
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MorphoBank Project 4146

    This research
    supported by

    Authors' Institutions

    • Harvard University

    • University of Cambridge

    • Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences

    • Amgueddfa Cymru (National Museum Wales)



    Members

    member name taxa specimens media chars character
    comms
       
    notes  media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    comms
        
    notes  medialabels
    rules
    Stephen Pates
    Project Administrator
    62111038010360049056
    (21862, 0, 27194)
    065000
    Jo Wolfe
    Full membership
    000000000
    (0, 0, 0)
    00000


    Taxonomic Overview for Matrix 'M27722' (57 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    023230106002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122424093002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273434068002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253535069002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552323051002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116363055002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244747058002
    [8] † nmw.2021.3g.8
    Last Modified in 12/09/21
    562626047002
    [9] † Mieridduryn bonniae
    Last Modified in 12/09/21
    414343045002
    [10] † Utaurora comosa
    Last Modified in 05/31/22
    473030052002
    [11] † Opabinia regalis
    Last Modified in 05/31/22
    235353053002
    [12] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    326767030002
    [13] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495656024002
    [14] † Cambroraster falcatus
    Last Modified in 05/31/22
    178282030002
    [15] † Hurdia triangulata
    Last Modified in 05/31/22
    178585027002
    [16] † Hurdia victoria
    Last Modified in 05/31/22
    178585027002
    [17] † Peytoia nathorsti
    Last Modified in 05/31/22
    237575031002
    [18] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    78464605002
    [19] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    247777028002
    [20] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178181031002
    [21] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148484031002
    [22] † Houcaris saron
    Last Modified in 12/09/21
    178181031002
    [23] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104242077002
    [24] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104242077002
    [25] † Leanchoilia superlata
    Last Modified in 05/31/22
    153433081002
    [26] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74040082002
    [27] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144040075002
    [28] † Kuamaia lata
    Last Modified in 05/31/22
    144040075002
    [29] Limulus polyphemus
    Last Modified in 05/31/22
    14646082002
    [30] Triops cancriformis
    Last Modified in 05/31/22
    15050078002
    [31] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144343072002
    [32] † Onychodictyon ferox
    Last Modified in 05/31/22
    145252063002
    [33] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244040065002
    [34] † Diania cactiformis
    Last Modified in 05/31/22
    204141068002
    [35] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313232066002
    [36] † Paucipodia inermis
    Last Modified in 05/31/22
    163636077002
    [37] † Microdictyon sinicum
    Last Modified in 05/31/22
    144545070002
    [38] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224343064002
    [39] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194141069002
    [40] † Hallucigenia fortis
    Last Modified in 05/31/22
    174646066002
    [41] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463030053002
    [42] † Luolishania longicruris
    Last Modified in 05/31/22
    175151061002
    [43] † Collinsium ciliosum
    Last Modified in 05/31/22
    145252063002
    [44] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542626049002
    [45] † Collins monster emu bay
    Last Modified in 05/31/22
    483535046002
    [46] † Acinocricus stichus
    Last Modified in 05/31/22
    552828046002
    [47] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661616047002
    [48] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292828072002
    [49] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283030071002
    [50] Euperipatoides sp
    Last Modified in 05/31/22
    05454075002
    [51] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15454074002
    [52] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15454074002
    [53] Batillipes pennaki
    Last Modified in 05/31/22
    35555071002
    [54] Echiniscus testudo
    Last Modified in 05/31/22
    26060067002
    [55] Actinarctus doryphorus
    Last Modified in 05/31/22
    45858067002
    [56] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05656073002
    [57] Hypsibius dujardini
    Last Modified in 05/31/22
    05656073002


    Taxonomic Overview for Matrix 'M27723' (56 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    023230106002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122424093002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273434068002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253535069002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552323051002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116363055002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244747058002
    [8] † Mieridduryn bonniae
    Last Modified in 12/09/21
    315352046002
    [9] † Utaurora comosa
    Last Modified in 05/31/22
    473030052002
    [10] † Opabinia regalis
    Last Modified in 05/31/22
    235353053002
    [11] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    326767030002
    [12] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495656024002
    [13] † Cambroraster falcatus
    Last Modified in 05/31/22
    178282030002
    [14] † Hurdia triangulata
    Last Modified in 05/31/22
    178585027002
    [15] † Hurdia victoria
    Last Modified in 05/31/22
    178585027002
    [16] † Peytoia nathorsti
    Last Modified in 05/31/22
    237575031002
    [17] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    78464605002
    [18] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    247777028002
    [19] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178181031002
    [20] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148484031002
    [21] † Houcaris saron
    Last Modified in 12/09/21
    178181031002
    [22] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104242077002
    [23] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104242077002
    [24] † Leanchoilia superlata
    Last Modified in 05/31/22
    153433081002
    [25] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74040082002
    [26] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144040075002
    [27] † Kuamaia lata
    Last Modified in 05/31/22
    144040075002
    [28] Limulus polyphemus
    Last Modified in 05/31/22
    14646082002
    [29] Triops cancriformis
    Last Modified in 05/31/22
    15050078002
    [30] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144343072002
    [31] † Onychodictyon ferox
    Last Modified in 05/31/22
    145252063002
    [32] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244040065002
    [33] † Diania cactiformis
    Last Modified in 05/31/22
    204141068002
    [34] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313232066002
    [35] † Paucipodia inermis
    Last Modified in 05/31/22
    163636077002
    [36] † Microdictyon sinicum
    Last Modified in 05/31/22
    144545070002
    [37] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224343064002
    [38] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194141069002
    [39] † Hallucigenia fortis
    Last Modified in 05/31/22
    174646066002
    [40] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463030053002
    [41] † Luolishania longicruris
    Last Modified in 05/31/22
    175151061002
    [42] † Collinsium ciliosum
    Last Modified in 05/31/22
    145252063002
    [43] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542626049002
    [44] † Collins monster emu bay
    Last Modified in 05/31/22
    483535046002
    [45] † Acinocricus stichus
    Last Modified in 05/31/22
    552828046002
    [46] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661616047002
    [47] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292828072002
    [48] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283030071002
    [49] Euperipatoides sp
    Last Modified in 05/31/22
    05454075002
    [50] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15454074002
    [51] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15454074002
    [52] Batillipes pennaki
    Last Modified in 05/31/22
    35555071002
    [53] Echiniscus testudo
    Last Modified in 05/31/22
    26060067002
    [54] Actinarctus doryphorus
    Last Modified in 05/31/22
    45858067002
    [55] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05656073002
    [56] Hypsibius dujardini
    Last Modified in 05/31/22
    05656073002


    Taxonomic Overview for Matrix 'M27724' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    023230107002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122424094002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273434069002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253535070002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552323052002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116363056002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244848058002
    [8] † nmw.2021.3g.8
    Last Modified in 12/09/21
    572626047002
    [9] † Mieridduryn bonniae
    Last Modified in 12/09/21
    414444045002
    [10] † Utaurora comosa
    Last Modified in 05/31/22
    483030052002
    [11] † Opabinia regalis
    Last Modified in 05/31/22
    245353053002
    [12] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    336767030002
    [13] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495757024002
    [14] † Cambroraster falcatus
    Last Modified in 05/31/22
    178383030002
    [15] † Hurdia triangulata
    Last Modified in 05/31/22
    178686027002
    [16] † Hurdia victoria
    Last Modified in 05/31/22
    178686027002
    [17] † Peytoia nathorsti
    Last Modified in 05/31/22
    237676031002
    [18] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    79464605002
    [19] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    257777028002
    [20] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178282031002
    [21] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148585031002
    [22] † Houcaris saron
    Last Modified in 12/09/21
    178282031002
    [23] † Parapeytoia yunnanensis protocerebral
    Last Modified in 12/09/21
    753333022002
    [24] † Kylinxia zhangi
    Last Modified in 12/09/21
    492727054002
    [25] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104242078002
    [26] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104242078002
    [27] † Leanchoilia superlata
    Last Modified in 05/31/22
    153433082002
    [28] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74040083002
    [29] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144040076002
    [30] † Kuamaia lata
    Last Modified in 05/31/22
    144040076002
    [31] Limulus polyphemus
    Last Modified in 05/31/22
    14646083002
    [32] Triops cancriformis
    Last Modified in 05/31/22
    15050079002
    [33] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144343073002
    [34] † Onychodictyon ferox
    Last Modified in 05/31/22
    145252064002
    [35] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244040066002
    [36] † Diania cactiformis
    Last Modified in 05/31/22
    204141069002
    [37] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313232067002
    [38] † Paucipodia inermis
    Last Modified in 05/31/22
    163636078002
    [39] † Microdictyon sinicum
    Last Modified in 05/31/22
    144545071002
    [40] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224343065002
    [41] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194141070002
    [42] † Hallucigenia fortis
    Last Modified in 05/31/22
    174646067002
    [43] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463030054002
    [44] † Luolishania longicruris
    Last Modified in 05/31/22
    175151062002
    [45] † Collinsium ciliosum
    Last Modified in 05/31/22
    145252064002
    [46] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542626050002
    [47] † Collins monster emu bay
    Last Modified in 05/31/22
    483535047002
    [48] † Acinocricus stichus
    Last Modified in 05/31/22
    552828047002
    [49] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661616048002
    [50] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292828073002
    [51] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283030072002
    [52] Euperipatoides sp
    Last Modified in 05/31/22
    05454076002
    [53] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15454075002
    [54] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15454075002
    [55] Batillipes pennaki
    Last Modified in 05/31/22
    35555072002
    [56] Echiniscus testudo
    Last Modified in 05/31/22
    26060068002
    [57] Actinarctus doryphorus
    Last Modified in 05/31/22
    45858068002
    [58] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05656074002
    [59] Hypsibius dujardini
    Last Modified in 05/31/22
    05656074002


    Taxonomic Overview for Matrix 'M27725' (59 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    023230107002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122424094002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273434069002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253535070002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552323052002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116363056002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244848058002
    [8] † nmw.2021.3g.8
    Last Modified in 12/09/21
    572626047002
    [9] † Mieridduryn bonniae
    Last Modified in 12/09/21
    414444045002
    [10] † Utaurora comosa
    Last Modified in 05/31/22
    483030052002
    [11] † Opabinia regalis
    Last Modified in 05/31/22
    245353053002
    [12] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    336767030002
    [13] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495757024002
    [14] † Cambroraster falcatus
    Last Modified in 05/31/22
    178383030002
    [15] † Hurdia triangulata
    Last Modified in 05/31/22
    178686027002
    [16] † Hurdia victoria
    Last Modified in 05/31/22
    178686027002
    [17] † Peytoia nathorsti
    Last Modified in 05/31/22
    237676031002
    [18] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    79464605002
    [19] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    257777028002
    [20] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178282031002
    [21] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148585031002
    [22] † Houcaris saron
    Last Modified in 12/09/21
    178282031002
    [23] † Parapeytoia yunnanensis deutocerebral
    Last Modified in 12/09/21
    961414020002
    [24] † Kylinxia zhangi
    Last Modified in 12/09/21
    492727054002
    [25] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104242078002
    [26] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104242078002
    [27] † Leanchoilia superlata
    Last Modified in 05/31/22
    153433082002
    [28] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74040083002
    [29] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144040076002
    [30] † Kuamaia lata
    Last Modified in 05/31/22
    144040076002
    [31] Limulus polyphemus
    Last Modified in 05/31/22
    14646083002
    [32] Triops cancriformis
    Last Modified in 05/31/22
    15050079002
    [33] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144343073002
    [34] † Onychodictyon ferox
    Last Modified in 05/31/22
    145252064002
    [35] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244040066002
    [36] † Diania cactiformis
    Last Modified in 05/31/22
    204141069002
    [37] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313232067002
    [38] † Paucipodia inermis
    Last Modified in 05/31/22
    163636078002
    [39] † Microdictyon sinicum
    Last Modified in 05/31/22
    144545071002
    [40] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224343065002
    [41] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194141070002
    [42] † Hallucigenia fortis
    Last Modified in 05/31/22
    174646067002
    [43] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463030054002
    [44] † Luolishania longicruris
    Last Modified in 05/31/22
    175151062002
    [45] † Collinsium ciliosum
    Last Modified in 05/31/22
    145252064002
    [46] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542626050002
    [47] † Collins monster emu bay
    Last Modified in 05/31/22
    483535047002
    [48] † Acinocricus stichus
    Last Modified in 05/31/22
    552828047002
    [49] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661616048002
    [50] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292828073002
    [51] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283030072002
    [52] Euperipatoides sp
    Last Modified in 05/31/22
    05454076002
    [53] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15454075002
    [54] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15454075002
    [55] Batillipes pennaki
    Last Modified in 05/31/22
    35555072002
    [56] Echiniscus testudo
    Last Modified in 05/31/22
    26060068002
    [57] Actinarctus doryphorus
    Last Modified in 05/31/22
    45858068002
    [58] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05656074002
    [59] Hypsibius dujardini
    Last Modified in 05/31/22
    05656074002


    Taxonomic Overview for Matrix 'M28784' (56 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    024240107002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122525094002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273535069002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253636070002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552424052002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116357057002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244744060002
    [8] CB chimera
    Last Modified in 05/31/22
    274745057002
    [9] † Utaurora comosa
    Last Modified in 05/31/22
    473131053002
    [10] † Opabinia regalis
    Last Modified in 05/31/22
    235453054002
    [11] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    326868031002
    [12] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495757025002
    [13] † Cambroraster falcatus
    Last Modified in 05/31/22
    178383031002
    [14] † Hurdia triangulata
    Last Modified in 05/31/22
    178686028002
    [15] † Hurdia victoria
    Last Modified in 05/31/22
    178685028002
    [16] † Peytoia nathorsti
    Last Modified in 05/31/22
    237676032002
    [17] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    78474706002
    [18] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    247877029002
    [19] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178282032002
    [20] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148585032002
    [21] Anomalocaris saron
    Last Modified in 05/31/22
    178282032002
    [22] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104343078002
    [23] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104343078002
    [24] † Leanchoilia superlata
    Last Modified in 05/31/22
    153534082002
    [25] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74141083002
    [26] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144141076002
    [27] † Kuamaia lata
    Last Modified in 05/31/22
    144141076002
    [28] Limulus polyphemus
    Last Modified in 05/31/22
    14747083002
    [29] Triops cancriformis
    Last Modified in 05/31/22
    15151079002
    [30] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144444073002
    [31] † Onychodictyon ferox
    Last Modified in 05/31/22
    145353064002
    [32] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244141066002
    [33] † Diania cactiformis
    Last Modified in 05/31/22
    204242069002
    [34] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313333067002
    [35] † Paucipodia inermis
    Last Modified in 05/31/22
    163737078002
    [36] † Microdictyon sinicum
    Last Modified in 05/31/22
    144646071002
    [37] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224444065002
    [38] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194242070002
    [39] † Hallucigenia fortis
    Last Modified in 05/31/22
    174747067002
    [40] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463131054002
    [41] † Luolishania longicruris
    Last Modified in 05/31/22
    175252062002
    [42] † Collinsium ciliosum
    Last Modified in 05/31/22
    145353064002
    [43] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542727050002
    [44] † Collins monster emu bay
    Last Modified in 05/31/22
    483636047002
    [45] † Acinocricus stichus
    Last Modified in 05/31/22
    552929047002
    [46] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661717048002
    [47] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292929073002
    [48] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283131072002
    [49] Euperipatoides sp
    Last Modified in 05/31/22
    05555076002
    [50] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15555075002
    [51] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15555075002
    [52] Batillipes pennaki
    Last Modified in 05/31/22
    35656072002
    [53] Echiniscus testudo
    Last Modified in 05/31/22
    26161068002
    [54] Actinarctus doryphorus
    Last Modified in 05/31/22
    45959068002
    [55] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05757074002
    [56] Hypsibius dujardini
    Last Modified in 05/31/22
    05757074002


    Taxonomic Overview for Matrix 'M28785' (56 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    024240106002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122525093002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273535068002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253636069002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552424051002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116363056002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244747059002
    [8] CB chimera
    Last Modified in 05/31/22
    264848056002
    [9] † Utaurora comosa
    Last Modified in 05/31/22
    443131055002
    [10] † Opabinia regalis
    Last Modified in 05/31/22
    205454056002
    [11] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    326868030002
    [12] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495757024002
    [13] † Cambroraster falcatus
    Last Modified in 05/31/22
    178383030002
    [14] † Hurdia triangulata
    Last Modified in 05/31/22
    178686027002
    [15] † Hurdia victoria
    Last Modified in 05/31/22
    178686027002
    [16] † Peytoia nathorsti
    Last Modified in 05/31/22
    237676031002
    [17] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    78474705002
    [18] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    247878028002
    [19] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178282031002
    [20] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148585031002
    [21] Anomalocaris saron
    Last Modified in 05/31/22
    178282031002
    [22] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104343077002
    [23] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104343077002
    [24] † Leanchoilia superlata
    Last Modified in 05/31/22
    153534081002
    [25] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74141082002
    [26] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144141075002
    [27] † Kuamaia lata
    Last Modified in 05/31/22
    144141075002
    [28] Limulus polyphemus
    Last Modified in 05/31/22
    14747082002
    [29] Triops cancriformis
    Last Modified in 05/31/22
    15151078002
    [30] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144444072002
    [31] † Onychodictyon ferox
    Last Modified in 05/31/22
    145353063002
    [32] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244141065002
    [33] † Diania cactiformis
    Last Modified in 05/31/22
    204242068002
    [34] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313333066002
    [35] † Paucipodia inermis
    Last Modified in 05/31/22
    163737077002
    [36] † Microdictyon sinicum
    Last Modified in 05/31/22
    144646070002
    [37] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224444064002
    [38] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194242069002
    [39] † Hallucigenia fortis
    Last Modified in 05/31/22
    174747066002
    [40] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463131053002
    [41] † Luolishania longicruris
    Last Modified in 05/31/22
    175252061002
    [42] † Collinsium ciliosum
    Last Modified in 05/31/22
    145353063002
    [43] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542727049002
    [44] † Collins monster emu bay
    Last Modified in 05/31/22
    483636046002
    [45] † Acinocricus stichus
    Last Modified in 05/31/22
    552929046002
    [46] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661717047002
    [47] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292929072002
    [48] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283131071002
    [49] Euperipatoides sp
    Last Modified in 05/31/22
    05555075002
    [50] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15555074002
    [51] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15555074002
    [52] Batillipes pennaki
    Last Modified in 05/31/22
    35656071002
    [53] Echiniscus testudo
    Last Modified in 05/31/22
    26161067002
    [54] Actinarctus doryphorus
    Last Modified in 05/31/22
    45959067002
    [55] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05757073002
    [56] Hypsibius dujardini
    Last Modified in 05/31/22
    05757073002


    Taxonomic Overview for Matrix 'M28786' (56 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    023230107002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122424094002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273434069002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253535070002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552323052002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116256057002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244643060002
    [8] CB chimera
    Last Modified in 05/31/22
    315351047002
    [9] † Utaurora comosa
    Last Modified in 05/31/22
    473030053002
    [10] † Opabinia regalis
    Last Modified in 05/31/22
    235352054002
    [11] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    326767031002
    [12] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495656025002
    [13] † Cambroraster falcatus
    Last Modified in 05/31/22
    178282031002
    [14] † Hurdia triangulata
    Last Modified in 05/31/22
    178585028002
    [15] † Hurdia victoria
    Last Modified in 05/31/22
    178584028002
    [16] † Peytoia nathorsti
    Last Modified in 05/31/22
    237575032002
    [17] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    78464606002
    [18] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    247776029002
    [19] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178181032002
    [20] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148484032002
    [21] Anomalocaris saron
    Last Modified in 05/31/22
    178181032002
    [22] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104242078002
    [23] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104242078002
    [24] † Leanchoilia superlata
    Last Modified in 05/31/22
    153433082002
    [25] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74040083002
    [26] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144040076002
    [27] † Kuamaia lata
    Last Modified in 05/31/22
    144040076002
    [28] Limulus polyphemus
    Last Modified in 05/31/22
    14646083002
    [29] Triops cancriformis
    Last Modified in 05/31/22
    15050079002
    [30] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144343073002
    [31] † Onychodictyon ferox
    Last Modified in 05/31/22
    145252064002
    [32] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244040066002
    [33] † Diania cactiformis
    Last Modified in 05/31/22
    204141069002
    [34] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313232067002
    [35] † Paucipodia inermis
    Last Modified in 05/31/22
    163636078002
    [36] † Microdictyon sinicum
    Last Modified in 05/31/22
    144545071002
    [37] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224343065002
    [38] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194141070002
    [39] † Hallucigenia fortis
    Last Modified in 05/31/22
    174646067002
    [40] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463030054002
    [41] † Luolishania longicruris
    Last Modified in 05/31/22
    175151062002
    [42] † Collinsium ciliosum
    Last Modified in 05/31/22
    145252064002
    [43] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542626050002
    [44] † Collins monster emu bay
    Last Modified in 05/31/22
    483535047002
    [45] † Acinocricus stichus
    Last Modified in 05/31/22
    552828047002
    [46] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661616048002
    [47] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292828073002
    [48] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283030072002
    [49] Euperipatoides sp
    Last Modified in 05/31/22
    05454076002
    [50] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15454075002
    [51] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15454075002
    [52] Batillipes pennaki
    Last Modified in 05/31/22
    35555072002
    [53] Echiniscus testudo
    Last Modified in 05/31/22
    26060068002
    [54] Actinarctus doryphorus
    Last Modified in 05/31/22
    45858068002
    [55] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05656074002
    [56] Hypsibius dujardini
    Last Modified in 05/31/22
    05656074002


    Taxonomic Overview for Matrix 'M28787' (56 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] Tubiluchus troglodytes
    Last Modified in 05/31/22
    023230106002
    [2] † Cricocosmia jinningensis
    Last Modified in 05/31/22
    122424093002
    [3] † Megadictyon haikouensis
    Last Modified in 05/31/22
    273434068002
    [4] † Jianshanopodia decora
    Last Modified in 05/31/22
    253535069002
    [5] † Hadranax augustus
    Last Modified in 05/31/22
    552323051002
    [6] † Kerygmachela kierkegaardi
    Last Modified in 05/31/22
    116357055002
    [7] † Pambdelurion whittingtoni
    Last Modified in 05/31/22
    244744058002
    [8] CB chimera
    Last Modified in 05/31/22
    315351046002
    [9] † Utaurora comosa
    Last Modified in 05/31/22
    473030052002
    [10] † Opabinia regalis
    Last Modified in 05/31/22
    235352053002
    [11] † Aegirocassis benmoulae
    Last Modified in 05/31/22
    326767030002
    [12] † Buccaspinea cooperi
    Last Modified in 05/31/22
    495656024002
    [13] † Cambroraster falcatus
    Last Modified in 05/31/22
    178282030002
    [14] † Hurdia triangulata
    Last Modified in 05/31/22
    178585027002
    [15] † Hurdia victoria
    Last Modified in 05/31/22
    178584027002
    [16] † Peytoia nathorsti
    Last Modified in 05/31/22
    237575031002
    [17] † Schinderhannes bartelsi
    Last Modified in 05/31/22
    78464605002
    [18] † Amplectobelua symbrachiata
    Last Modified in 05/31/22
    247776028002
    [19] † Lyrarapax unguispinus
    Last Modified in 05/31/22
    178181031002
    [20] † Anomalocaris canadensis
    Last Modified in 05/31/22
    148484031002
    [21] Anomalocaris saron
    Last Modified in 05/31/22
    178181031002
    [22] † Fuxianhuia xiaoshibaensis
    Last Modified in 05/31/22
    104242077002
    [23] † Chengjiangocaris kunmingensis
    Last Modified in 05/31/22
    104242077002
    [24] † Leanchoilia superlata
    Last Modified in 05/31/22
    153433081002
    [25] † Alalcomenaeus sp
    Last Modified in 05/31/22
    74040082002
    [26] † Misszhouia longicaudata
    Last Modified in 05/31/22
    144040075002
    [27] † Kuamaia lata
    Last Modified in 05/31/22
    144040075002
    [28] Limulus polyphemus
    Last Modified in 05/31/22
    14646082002
    [29] Triops cancriformis
    Last Modified in 05/31/22
    15050078002
    [30] † Aysheaia pedunculata
    Last Modified in 05/31/22
    144343072002
    [31] † Onychodictyon ferox
    Last Modified in 05/31/22
    145252063002
    [32] † Onychodictyon gracilis
    Last Modified in 05/31/22
    244040065002
    [33] † Diania cactiformis
    Last Modified in 05/31/22
    204141068002
    [34] † Xenusion auerswaldae
    Last Modified in 05/31/22
    313232066002
    [35] † Paucipodia inermis
    Last Modified in 05/31/22
    163636077002
    [36] † Microdictyon sinicum
    Last Modified in 05/31/22
    144545070002
    [37] † Cardiodictyon catenulum
    Last Modified in 05/31/22
    224343064002
    [38] † Hallucigenia sparsa
    Last Modified in 05/31/22
    194141069002
    [39] † Hallucigenia fortis
    Last Modified in 05/31/22
    174646066002
    [40] † Hallucigenia hongmeia
    Last Modified in 05/31/22
    463030053002
    [41] † Luolishania longicruris
    Last Modified in 05/31/22
    175151061002
    [42] † Collinsium ciliosum
    Last Modified in 05/31/22
    145252063002
    [43] † Collinsovermis monstruosus
    Last Modified in 05/31/22
    542626049002
    [44] † Collins monster emu bay
    Last Modified in 05/31/22
    483535046002
    [45] † Acinocricus stichus
    Last Modified in 05/31/22
    552828046002
    [46] † Orstenotubulus evamuellerae
    Last Modified in 05/31/22
    661616047002
    [47] † Antennacanthopodia gracilis
    Last Modified in 05/31/22
    292828072002
    [48] † Tertiapatus dominicanus
    Last Modified in 05/31/22
    283030071002
    [49] Euperipatoides sp
    Last Modified in 05/31/22
    05454075002
    [50] Metaperipatus blainvillei
    Last Modified in 05/31/22
    15454074002
    [51] Epiperipatus biolleyi
    Last Modified in 05/31/22
    15454074002
    [52] Batillipes pennaki
    Last Modified in 05/31/22
    35555071002
    [53] Echiniscus testudo
    Last Modified in 05/31/22
    26060067002
    [54] Actinarctus doryphorus
    Last Modified in 05/31/22
    45858067002
    [55] Macrobiotus cf. harmsworthi
    Last Modified in 05/31/22
    05656073002
    [56] Hypsibius dujardini
    Last Modified in 05/31/22
    05656073002


    Project views

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    Total project views4549
    Project overview380
    Matrix views531Matrix landing page (345 views); Matrix for Supplementary Figure 5 (38 views); Matrix for Supplementary Figure 11d (17 views); Matrix for Supplementary Figure 11c (17 views); Matrix for Supplementary Figure 6 (16 views); Matrix for Supplementary Figure 11b (22 views); Matrix for Supplementary Figure 9 (35 views); Matrix for Supplementary Figure 11a (19 views); Matrix for Supplementary Figure 10 (22 views);
    Media views827Media search (699 views); M848214 (128 views);
    Views for media list251
    Specimen list403
    Taxon list1977
    Bibliography176
    Documents list4




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    Total downloads from project14
    Project downloads4
    Matrix downloads10Matrix for Supplementary Figure 9 (2 downloads); Matrix for Supplementary Figure 5 (2 downloads); Matrix for Supplementary Figure 6 (1 download); Matrix for Supplementary Figure 10 (1 download); Matrix for Supplementary Figure 11a (1 download); Matrix for Supplementary Figure 11b (1 download); Matrix for Supplementary Figure 11c (1 download); Matrix for Supplementary Figure 11d (1 download);