Project 423: G. Giribet, G. D. Edgecombe, W. C. Wheeler, C. Babbitt. 2002. Phylogeny and Systematic Position of Opiliones: A Combined Analysis of Chelicerate Relationships Using Morphological and Molecular Data. Cladistics. 18 (1):5-70.
Abstract
The ordinal level phylogeny of the Arachnida and the suprafamilial level phylogeny of the Opiliones were studied on the basis of a combined analysis of 253 morphological characters, the complete sequence of the 18S rRNA gene, and the D3 region of the 28S rRNA gene. Molecular data were collected for 63 terminal taxa. Morphological data were collected for 35 exemplar taxa of Opiliones, but groundplans were applied to some of the remaining chelicerate groups. Six extinct terminals, including Paleozoic scorpions, are scored for morphological characters. The data were analyzed using strict parsimony for the morphological data matrix and via direct optimization for the molecular and combined data matrices. A sensitivity analysis of 15 parameter sets was undertaken, and character congruence was used as the optimality criterion to choose among competing hypotheses. The results obtained are unstable for the high-level chelicerate relationships (except for Tetrapulmonata, Pedipalpi, and Camarostomata), and the sister group of the Opiliones is not clearly established, although the monophyly of Dromopoda is supported under many parameter sets. However, the internal phylogeny of the Opiliones is robust to parameter choice and allows the discarding of previous hypotheses of opilionid phylogeny such as the “Cyphopalpatores” or “Palpatores.” The topology obtained is congruent with the previous hypothesis of “Palpatores” paraphyly as follows: (Cyphophthalmi (Eupnoi (Dyspnoi + Laniatores))). Resolution within the Eupnoi, Dyspnoi, and Laniatores (the latter two united as Dyspnolaniatores nov.) is also stable to the superfamily level, permitting a new classification system for the Opiliones.Read the article »
Project DOI: 10.7934/P423, http://dx.doi.org/10.7934/P423
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 12393 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 253 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 540 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:253/0 |
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MorphoBank Project 423
MorphoBank Project 423
- Creation Date:
17 October 2010 - Publication Date:
13 February 2011 - Matrix downloads: 20
Authors' Institutions
- Harvard University
- American Museum of Natural History (AMNH)
- Australian Museum
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 60 | 0 | 1 | 253 | 0 | 0 | 12393 (10242, 0, 2151) | 0 | 0 | 0 | ||||
Maureen Admin Full membership | 1 | 1 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Gonzalo Giribet Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Gonzalo Giribet Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M983' (60 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Trilobita Taxon name last Modified on 12/05/13 | 111 | 57 | 57 | 0 | 85 | 0 | 0 | 4 |
[2] Palaeoisopus Taxon name last Modified on 12/05/13 | 136 | 57 | 57 | 0 | 60 | 0 | 0 | 4 |
[3] Pycnogonida Taxon name last Modified on 12/05/13 | 71 | 109 | 109 | 0 | 73 | 0 | 0 | 4 |
[4] Eurypterida Taxon name last Modified on 12/05/13 | 110 | 96 | 96 | 0 | 47 | 0 | 0 | 4 |
[5] Limulus Taxon name last Modified on 12/05/13 | 42 | 161 | 161 | 0 | 50 | 0 | 0 | 4 |
[6] Carcinoscorpius Taxon name last Modified on 12/05/13 | 44 | 159 | 159 | 0 | 50 | 0 | 0 | 4 |
[7] Palaeophonidae Taxon name last Modified on 12/05/13 | 126 | 80 | 80 | 0 | 47 | 0 | 0 | 4 |
[8] Proscorpius Taxon name last Modified on 12/05/13 | 122 | 84 | 84 | 0 | 47 | 0 | 0 | 4 |
[9] Androctonus Taxon name last Modified on 12/05/13 | 32 | 176 | 176 | 0 | 45 | 0 | 0 | 4 |
[10] Belisarius Taxon name last Modified on 12/05/13 | 38 | 166 | 166 | 0 | 49 | 0 | 0 | 4 |
[11] Siro Taxon name last Modified on 12/05/13 | 33 | 191 | 191 | 0 | 29 | 0 | 0 | 4 |
[12] Parasiro Taxon name last Modified on 12/05/13 | 48 | 176 | 176 | 0 | 29 | 0 | 0 | 4 |
[13] Stylocellus Taxon name last Modified on 12/05/13 | 49 | 178 | 178 | 0 | 26 | 0 | 0 | 4 |
[14] StylocellusJP Taxon name last Modified on 12/05/13 | 49 | 178 | 178 | 0 | 26 | 0 | 0 | 4 |
[15] Dalquestia Taxon name last Modified on 12/05/13 | 48 | 186 | 186 | 0 | 19 | 0 | 0 | 4 |
[16] Odiellus Taxon name last Modified on 12/05/13 | 35 | 196 | 196 | 0 | 22 | 0 | 0 | 4 |
[17] Phalangium Taxon name last Modified on 12/05/13 | 34 | 197 | 197 | 0 | 22 | 0 | 0 | 4 |
[18] Opilio Taxon name last Modified on 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 4 |
[19] Astrobunus Taxon name last Modified on 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 4 |
[20] Nelima Taxon name last Modified on 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 4 |
[21] Leiobunum Taxon name last Modified on 12/05/13 | 32 | 199 | 199 | 0 | 22 | 0 | 0 | 4 |
[22] Hadrobunus Taxon name last Modified on 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 4 |
[23] Caddo Taxon name last Modified on 12/05/13 | 48 | 186 | 186 | 0 | 19 | 0 | 0 | 4 |
[24] Ischyropsalis Taxon name last Modified on 12/05/13 | 35 | 191 | 191 | 0 | 27 | 0 | 0 | 4 |
[25] Hesperonemastoma Taxon name last Modified on 12/05/13 | 47 | 180 | 180 | 0 | 26 | 0 | 0 | 4 |
[26] Ceratolasma Taxon name last Modified on 12/05/13 | 48 | 181 | 181 | 0 | 24 | 0 | 0 | 4 |
[27] Sabacon Taxon name last Modified on 12/05/13 | 47 | 182 | 182 | 0 | 24 | 0 | 0 | 4 |
[28] Taracus Taxon name last Modified on 12/05/13 | 47 | 182 | 182 | 0 | 24 | 0 | 0 | 4 |
[29] Dicranolasma Taxon name last Modified on 12/05/13 | 47 | 180 | 180 | 0 | 26 | 0 | 0 | 4 |
[30] Centetostoma Taxon name last Modified on 12/05/13 | 50 | 177 | 177 | 0 | 26 | 0 | 0 | 4 |
[31] Ortholasma Taxon name last Modified on 12/05/13 | 48 | 180 | 180 | 0 | 25 | 0 | 0 | 4 |
[32] Nemastoma Taxon name last Modified on 12/05/13 | 33 | 191 | 191 | 0 | 29 | 0 | 0 | 4 |
[33] Nipponopsalis Taxon name last Modified on 12/05/13 | 48 | 180 | 180 | 0 | 25 | 0 | 0 | 4 |
[34] Trogulus Taxon name last Modified on 12/05/13 | 34 | 190 | 190 | 0 | 29 | 0 | 0 | 4 |
[35] Equitius Taxon name last Modified on 12/05/13 | 47 | 184 | 184 | 0 | 22 | 0 | 0 | 4 |
[36] Triaenobunus Taxon name last Modified on 12/05/13 | 48 | 183 | 183 | 0 | 22 | 0 | 0 | 4 |
[37] Zuma Taxon name last Modified on 12/05/13 | 48 | 183 | 183 | 0 | 22 | 0 | 0 | 4 |
[38] Oncopus Taxon name last Modified on 12/05/13 | 47 | 184 | 183 | 0 | 23 | 0 | 0 | 4 |
[39] Gnomulus Taxon name last Modified on 12/05/13 | 47 | 184 | 183 | 0 | 23 | 0 | 0 | 4 |
[40] Scotolemon Taxon name last Modified on 12/05/13 | 35 | 192 | 192 | 0 | 26 | 0 | 0 | 4 |
[41] Maiorerus Taxon name last Modified on 12/05/13 | 35 | 190 | 190 | 0 | 28 | 0 | 0 | 4 |
[42] Bishopella Taxon name last Modified on 12/05/13 | 35 | 192 | 192 | 0 | 26 | 0 | 0 | 4 |
[43] Gnidia Taxon name last Modified on 12/05/13 | 33 | 194 | 194 | 0 | 26 | 0 | 0 | 4 |
[44] Pachyloides Taxon name last Modified on 12/05/13 | 34 | 193 | 193 | 0 | 26 | 0 | 0 | 4 |
[45] Hoplobunus Taxon name last Modified on 12/05/13 | 35 | 190 | 190 | 0 | 28 | 0 | 0 | 4 |
[46] Solifugae Taxon name last Modified on 12/05/13 | 34 | 157 | 157 | 0 | 62 | 0 | 0 | 4 |
[47] Palpigradi Taxon name last Modified on 12/05/13 | 13 | 184 | 184 | 0 | 56 | 0 | 0 | 4 |
[48] Liphistius Taxon name last Modified on 12/05/13 | 3 | 206 | 206 | 0 | 44 | 0 | 0 | 4 |
[49] Aphonopelma Taxon name last Modified on 12/05/13 | 9 | 200 | 200 | 0 | 44 | 0 | 0 | 4 |
[50] Nesticus Taxon name last Modified on 12/05/13 | 7 | 198 | 198 | 0 | 48 | 0 | 0 | 4 |
[51] Amblypygi Taxon name last Modified on 12/05/13 | 4 | 206 | 206 | 0 | 43 | 0 | 0 | 4 |
[52] Mastigoproctus Taxon name last Modified on 12/05/13 | 6 | 205 | 205 | 0 | 42 | 0 | 0 | 4 |
[53] Schizomida Taxon name last Modified on 12/05/13 | 9 | 198 | 198 | 0 | 46 | 0 | 0 | 4 |
[54] Ricinulei Taxon name last Modified on 12/05/13 | 38 | 159 | 159 | 0 | 56 | 0 | 0 | 4 |
[55] Pseudoscorpiones Taxon name last Modified on 12/05/13 | 34 | 163 | 163 | 0 | 56 | 0 | 0 | 4 |
[56] Allonothrus Taxon name last Modified on 12/05/13 | 57 | 150 | 150 | 0 | 46 | 0 | 0 | 4 |
[57] Acarus Taxon name last Modified on 12/05/13 | 44 | 150 | 150 | 0 | 59 | 0 | 0 | 4 |
[58] Opilioacarus Taxon name last Modified on 12/05/13 | 57 | 151 | 151 | 0 | 45 | 0 | 0 | 4 |
[59] Rhipicephalus Taxon name last Modified on 12/05/13 | 44 | 148 | 148 | 0 | 61 | 0 | 0 | 4 |
[60] Trigonotarbida Taxon name last Modified on 12/05/13 | 106 | 104 | 104 | 0 | 43 | 0 | 0 | 4 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 406 | |
Matrix downloads | 20 | Phylogeny and Systematic Position of Opiliones: A Combined Analysis of Chelicerate Relationships Using Morphological and Molecular Data (20 downloads); |
Project downloads | 385 | |
Document downloads | 1 | copyright information for exemplar media (1 download); |