Morphobank

Project 4327: D. Foffa, E. M. Dunne, S. J. Nesbitt, R. J. Butler, N. C. Fraser, S. L. Brusatte, A. Farnsworth, D. J. Lunt, P. J. Valdes, S. Walsh, P. M. Barrett. 2022. Scleromochlus and the early evolution of Pterosauromorpha. Nature. 610 (7931):313-318.

Specimen: † Scleromochlus taylori Woodward, 1907
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Abstract

Pterosaurs, the first vertebrates to evolve powered flight, were key components of Mesozoic terrestrial ecosystems, from their sudden appearance in the Late Triassic until their demise at the end of the Cretaceous1–6. However, the origin and early evolution of pterosaurs are poorly understood, due to a substantial stratigraphic and morphological gap between these reptiles and their closest relatives6, Lagerpetidae7. Scleromochlus taylori, a tiny reptile from the early Late Triassic of Scotland discovered over a century ago, was hypothesized to be a key taxon closely related to pterosaurs8, but its poor preservation has limited prior studies and resulted in controversy over its phylogenetic position, with some even doubting its identification as an archosaur9. Here we use micro-computed tomographic scans to provide the first accurate whole skeleton reconstruction and a revised diagnosis of Scleromochlus, revealing new anatomical details that conclusively identify it as a close pterosaur relative1 within Pterosauromorpha (the lagerpetid + pterosaur clade). Scleromochlus is anatomically more similar to lagerpetids than to pterosaurs and retains numerous features that were likely present in very early diverging members of Avemetatarsalia (bird-line archosaurs). These results support the hypothesis that the first flying reptiles evolved from tiny, likely facultatively bipedal, cursorial ancestors1.

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Article DOI: 10.1038/s41586-022-05284-x

Project DOI: 10.7934/P4327, http://dx.doi.org/10.7934/P4327

This project contains	Matrices
1 Media 1 Matrix 6 Documents 198 Taxa 1 Specimen 823 Characters Total size of project's media files: 453.12k Download Project SDD File	Total scored cells: 68622 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 823 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1910 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:716/107

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MorphoBank Project 4327

Creation Date:
19 July 2022
Publication Date:
07 October 2022
Project views: 1814
Media downloads: 1
Matrix downloads: 18

Authors' Institutions

Natural History Museum, London

National Museums of Scotland, Royal Museum, Edinburgh

Chinese Academy of Sciences

University of Birmingham

University of Edinburgh

Virginia Tech (Virginia Polytechnic Institute and State University)

University of Bristol

Friedrich-Alexander-Universität Erlangen-Nürnberg

Institute of Tibetan Plateau Research

Members

Davide Foffa
Project Administrator

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Taxonomic Overview for Matrix 'M29034' (197 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Petrolacosaurus kansensis Last Modified in 10/26/22	105	621	603	0	99	0	0	1
[2] † Acerosodontosaurus piveteaui Last Modified in 10/26/22	531	231	224	0	62	0	0	1
[3] † Youngina capensis Last Modified in 10/26/22	101	638	591	0	89	0	0	1
[4] † Paliguana whitei Last Modified in 10/26/22	692	103	101	0	28	0	0	1
[5] † Planocephalosaurus robinsonae Last Modified in 10/26/22	269	462	449	0	94	0	0	1
[6] † Gephyrosaurus bridensis Last Modified in 10/26/22	251	498	469	0	79	0	0	1
[7] † Cteniogenys sp Last Modified in 10/26/22	459	313	306	0	56	0	0	1
[8] † Simoedosaurus lemoinei Last Modified in 10/26/22	185	560	554	0	83	0	0	1
[9] † Aenigmastropheus parringtoni Last Modified in 10/26/22	778	45	43	0	1	0	0	1
[10] † Protorosaurus speneri Last Modified in 10/26/22	223	536	487	0	78	0	0	1
[11] † Dinocephalosaurus orientalis Last Modified in 10/26/22	403	327	323	0	97	0	0	1
[12] † Amotosaurus rotfeldensis Last Modified in 10/26/22	424	345	337	0	60	0	0	1
[13] † Macrocnemus bassanii Last Modified in 10/26/22	269	478	445	0	81	0	0	1
[14] † Macrocnemus obristi Last Modified in 10/26/22	729	76	75	0	19	0	0	1
[15] † Macrocnemus fuyuanensis Last Modified in 10/26/22	345	407	399	0	79	0	0	1
[16] † Fuyuansaurus acutirostris Last Modified in 10/26/22	663	137	133	0	26	0	0	1
[17] † Pectodens zhenyuensis Last Modified in 10/26/22	506	256	247	0	70	0	0	1
[18] † Augustaburiania vatagini Last Modified in 10/26/22	737	79	79	0	7	0	0	1
[19] † Protanystropheus antiquus Last Modified in 10/26/22	802	22	20	0	1	0	0	1
[20] † Trachelosaurus fischeri Last Modified in 10/26/22	728	91	80	0	15	0	0	1
[21] † Tanystropheus longobardicus Last Modified in 10/26/22	106	628	570	0	106	0	0	1
[22] † Tanystropheus haasi Last Modified in 10/26/22	812	10	10	0	1	0	0	1
[23] † Langobadisaurus pandolfii Last Modified in 10/26/22	475	294	280	0	67	0	0	1
[24] † Tanytrachelos ahynis Last Modified in 10/26/22	451	331	310	0	62	0	0	1
[25] † Jesairosaurus lehmani Last Modified in 10/26/22	410	346	329	0	78	0	0	1
[26] † Pamelaria dolichotrachela Last Modified in 10/26/22	198	568	563	0	62	0	0	1
[27] † Azendohsaurus madagaskarensis Last Modified in 10/26/22	87	670	659	0	72	0	0	1
[28] † Azendohsaurus laaroussi Last Modified in 10/26/22	745	68	67	0	11	0	0	1
[29] † Shringasaurus indicus Last Modified in 10/26/22	376	402	394	0	53	0	0	1
[30] † Malerisaurus robinsonae Last Modified in 10/26/22	488	312	300	0	35	0	0	1
[31] † Trilophosaurus buettneri Last Modified in 10/26/22	103	634	600	0	92	0	0	1
[32] † Spinosuchus caseanus Last Modified in 10/26/22	418	342	338	0	67	0	0	1
[33] † Teraterpeton hrynewichorum Last Modified in 10/26/22	378	372	368	0	77	0	0	1
[34] † Arctosaurus osborni Last Modified in 10/26/22	809	13	13	0	1	0	0	1
[35] † Noteosuchus colletti Last Modified in 10/26/22	604	202	190	0	20	0	0	1
[36] † Mesosuchus browni Last Modified in 10/26/22	80	668	635	0	79	0	0	1
[37] † Howesia browni Last Modified in 10/26/22	396	390	363	0	43	0	0	1
[38] † Eohyosaurus wolvaardti Last Modified in 10/26/22	645	163	156	0	21	0	0	1
[39] † Rhynchosaurus articeps Last Modified in 10/26/22	210	536	521	0	81	0	0	1
[40] † Bentonyx sidensis Last Modified in 10/26/22	531	237	236	0	56	0	0	1
[41] † Mesodapedon kuttyi Last Modified in 10/26/22	775	38	38	0	10	0	0	1
[42] † Ammorhynchus navajoi Last Modified in 10/26/22	752	48	47	0	24	0	0	1
[43] † Stenaulorhynchus stockleyi Last Modified in 10/26/22	145	599	597	0	81	0	0	1
[44] † Brasinorhynchus mariantensis Last Modified in 10/26/22	487	286	283	0	53	0	0	1
[45] † Chanares rhynchosaur Last Modified in 10/26/22	766	49	46	0	11	0	0	1
[46] † Langeronyx brodiei Last Modified in 10/26/22	635	145	144	0	44	0	0	1
[47] † Fodonyx spenceri Last Modified in 10/26/22	662	140	140	0	21	0	0	1
[48] † Isalorhynchus genovefae Last Modified in 10/26/22	231	523	519	0	71	0	0	1
[49] † Hyperodapedon gordoni Last Modified in 10/26/22	123	611	606	0	94	0	0	1
[50] † Hyperodapedon huxleyi Last Modified in 10/26/22	120	611	609	0	94	0	0	1
[51] † Teyumbaita sulcognathus Last Modified in 10/26/22	246	497	494	0	83	0	0	1
[52] † Eorasaurus olsoni Last Modified in 10/26/22	794	29	26	0	2	0	0	1
[53] † Prolacertoides jimusarensis Last Modified in 10/26/22	719	74	72	0	31	0	0	1
[54] † Prolacerta broomi Last Modified in 10/26/22	38	725	679	0	73	0	0	1
[55] † K australiensis combined Last Modified in 10/26/22	691	105	100	0	32	0	0	1
[56] † Boreopricea funerea Last Modified in 10/26/22	455	325	300	0	50	0	0	1
[57] † Teyujagua paradoxa Last Modified in 10/26/22	508	262	260	0	55	0	0	1
[58] † Antarctanax shackletoni Last Modified in 10/26/22	751	67	62	0	6	0	0	1
[59] † Archosaurus rossicus holotype Last Modified in 10/26/22	801	21	21	0	1	0	0	1
[60] † Archosaurus complete Last Modified in 10/26/22	722	103	99	0	2	0	0	1
[61] † Proterosuchus fergusi Last Modified in 10/26/22	177	622	607	0	37	0	0	1
[62] † Proterosuchus goweri Last Modified in 10/26/22	529	282	278	0	16	0	0	1
[63] † Proterosuchus alexanderi Last Modified in 10/26/22	256	528	524	0	42	0	0	1
[64] † Chasmatosaurus yuani Last Modified in 10/26/22	192	594	580	0	50	0	0	1
[65] † Panchet proterosuchid Last Modified in 10/26/22	636	183	173	0	14	0	0	1
[66] † Tasmaniosaurus triassicus Last Modified in 10/26/22	690	126	119	0	14	0	0	1
[67] † Vonhuenia fredericki Last Modified in 10/26/22	799	21	21	0	3	0	0	1
[68] † C rossicus combined Last Modified in 10/26/22	775	46	45	0	3	0	0	1
[69] † C magnus combined Last Modified in 10/26/22	793	29	28	0	2	0	0	1
[70] † Chasmatosuchus vjushkovi Last Modified in 10/26/22	796	28	26	0	1	0	0	1
[71] † Koilamasuchus gonzalezdiazi Last Modified in 10/26/22	790	32	29	0	4	0	0	1
[72] † Kalisuchus rewanensis holotype Last Modified in 10/26/22	768	48	43	0	12	0	0	1
[73] † NMQR 3570 Last Modified in 10/26/22	777	35	35	0	11	0	0	1
[74] † Fugusuchus hejiapanensis Last Modified in 10/26/22	580	224	221	0	21	0	0	1
[75] † Sarmatosuchus otschevi Last Modified in 10/26/22	557	264	256	0	10	0	0	1
[76] † Guchengosuchus shiguaiensis Last Modified in 10/26/22	599	206	198	0	26	0	0	1
[77] † Cuyosuchus huenei Last Modified in 10/26/22	613	208	198	0	12	0	0	1
[78] † Garjainia prima Last Modified in 10/26/22	155	633	627	0	40	0	0	1
[79] † Garjainia madiba combined Last Modified in 10/26/22	433	381	362	0	22	0	0	1
[80] † Erythrosuchus africanus Last Modified in 10/26/22	95	685	672	0	48	0	0	1
[81] † Shansisuchus shansisuchus Last Modified in 10/26/22	219	566	558	0	45	0	0	1
[82] † Shansisuchus kuyeheensis Last Modified in 10/26/22	723	94	88	0	12	0	0	1
[83] † Chalishevia cothurnata Last Modified in 10/26/22	707	108	105	0	11	0	0	1
[84] † Youngosuchus sinensis Last Modified in 10/26/22	446	363	356	0	21	0	0	1
[85] † Uralosaurus combined Last Modified in 10/26/22	783	34	33	0	7	0	0	1
[86] † Vancleavea campi Last Modified in 10/26/22	265	491	486	0	70	0	0	1
[87] † Litorosuchus somnii Last Modified in 10/26/22	343	454	448	0	32	0	0	1
[88] † Asperoris mnyama Last Modified in 10/26/22	698	117	102	0	17	0	0	1
[89] † Euparkeria capensis Last Modified in 10/26/22	48	753	741	0	34	0	0	1
[90] † Dorosuchus neoetus Last Modified in 10/26/22	698	122	122	0	3	0	0	1
[91] † Halazhaisuchus qiaoensis Last Modified in 10/26/22	719	96	95	0	9	0	0	1
[92] † Osmolskina czatkoviensis Last Modified in 10/26/22	792	24	24	0	7	0	0	1
[93] † Osmolskina complete Last Modified in 10/26/22	335	470	460	0	28	0	0	1
[94] † Yarasuchus deccanensis Last Modified in 10/26/22	590	223	219	0	13	0	0	1
[95] † Dongusuchus efremovi Last Modified in 10/26/22	781	40	40	0	2	0	0	1
[96] † Teleocrater rhadinus Last Modified in 10/26/22	486	319	318	0	19	0	0	1
[97] † Spondylosoma absconditum Last Modified in 10/26/22	737	83	82	0	4	0	0	1
[98] † Proterochampsa barrionuevoi Last Modified in 10/26/22	271	506	497	0	55	0	0	1
[99] † Proterochampsa nodosa Last Modified in 10/26/22	570	234	228	0	20	0	0	1
[100] † Tropidosuchus romeri Last Modified in 10/26/22	178	613	598	0	40	0	0	1
[101] † Cerritosaurus binsfeldi Last Modified in 10/26/22	568	235	230	0	25	0	0	1
[102] † Gualosuchus reigi Last Modified in 10/26/22	262	523	514	0	45	0	0	1
[103] † Chanaresuchus bonapartei Last Modified in 10/26/22	98	687	672	0	49	0	0	1
[104] † Pseudochampsa ischigualastensis Last Modified in 10/26/22	354	439	433	0	36	0	0	1
[105] † Rhadinosuchus gracilis Last Modified in 10/26/22	689	128	121	0	13	0	0	1
[106] † Archeopelta arborensis Last Modified in 10/26/22	706	113	113	0	4	0	0	1
[107] † Tarjadia ruthae Last Modified in 10/26/22	299	498	494	0	30	0	0	1
[108] † Pagosvenator candelariensis Last Modified in 10/26/22	645	176	175	0	3	0	0	1
[109] † Erpetosuchus granti Last Modified in 10/26/22	450	348	344	0	27	0	0	1
[110] † Erpetosuchus sp Last Modified in 10/26/22	718	99	94	0	11	0	0	1
[111] † Dyoplax arenaceus Last Modified in 10/26/22	763	59	57	0	3	0	0	1
[112] † Jaxtasuchus salomoni Last Modified in 10/26/22	596	216	208	0	19	0	0	1
[113] † Doswellia kaltenbachi Last Modified in 10/26/22	407	368	362	0	54	0	0	1
[114] † Triopticus primus Last Modified in 10/26/22	718	102	93	0	7	0	0	1
[115] † Parasuchus angustifrons Last Modified in 10/26/22	542	251	248	0	33	0	0	1
[116] † Parasuchus hislopi Last Modified in 10/26/22	112	678	669	0	40	0	0	1
[117] † Nicrosaurus kapffi Last Modified in 10/26/22	271	513	505	0	46	0	0	1
[118] † Smilosuchus spp. Last Modified in 10/26/22	139	638	624	0	54	0	0	1
[119] † Diandongosuchus fuyuanensis Last Modified in 10/26/22	347	453	451	0	25	0	0	1
[120] † Wannia scurriensis Last Modified in 10/26/22	613	185	173	0	32	0	0	1
[121] † Ebrachosuchus neukami Last Modified in 10/26/22	505	286	282	0	35	0	0	1
[122] † Paleorhinus sawini Last Modified in 10/26/22	542	256	252	0	29	0	0	1
[123] † Angistorhinus talainti Last Modified in 10/26/22	473	316	313	0	37	0	0	1
[124] † Mystriosuchus planirostris Last Modified in 10/26/22	327	474	455	0	41	0	0	1
[125] † Machaeroprosopus pristinus Last Modified in 10/26/22	347	447	428	0	48	0	0	1
[126] † Redondasaurus spp Last Modified in 10/26/22	473	323	311	0	39	0	0	1
[127] † Ornithosuchus woodwardi Last Modified in 10/26/22	258	534	527	0	34	0	0	1
[128] † Riojasuchus tenuisceps Last Modified in 10/26/22	132	656	644	0	41	0	0	1
[129] † Venaticosuchus rusconii Last Modified in 10/26/22	625	182	179	0	19	0	0	1
[130] † Dynamosuchus collisensis Last Modified in 10/26/22	541	282	279	0	3	0	0	1
[131] † Nundasuchus songeaensis Last Modified in 10/26/22	528	287	282	0	13	0	0	1
[132] † Turfanosuchus dabanensis Last Modified in 10/26/22	263	537	530	0	28	0	0	1
[133] † Gracilisuchus stipanicicorum Last Modified in 10/26/22	156	639	627	0	40	0	0	1
[134] † Yonghesuchus sangbiensis Last Modified in 10/26/22	611	195	192	0	20	0	0	1
[135] † Parringtonia gracilis Last Modified in 10/26/22	340	461	455	0	28	0	0	1
[136] † Otter sandstone archosaur Last Modified in 10/26/22	772	54	48	0	3	0	0	1
[137] † Aetosauroides scagliai Last Modified in 10/26/22	264	539	529	0	29	0	0	1
[138] † Aetosaurus ferratus Last Modified in 10/26/22	219	570	565	0	39	0	0	1
[139] † Stagonolepis robertsoni Last Modified in 10/26/22	96	686	686	0	41	0	0	1
[140] † Ticinosuchus ferox Last Modified in 10/26/22	489	318	311	0	22	0	0	1
[141] † Prestosuchus nyassicus Last Modified in 10/26/22	656	165	159	0	8	0	0	1
[142] † Dagasuchus santacruzensis Last Modified in 10/26/22	792	29	29	0	2	0	0	1
[143] † Qianosuchus mixtus Last Modified in 10/26/22	321	472	470	0	32	0	0	1
[144] † Arizonasaurus babbitti Last Modified in 10/26/22	375	414	409	0	39	0	0	1
[145] † Lotosaurus adentus Last Modified in 10/26/22	317	432	426	0	80	0	0	1
[146] † Ctenosauriscus koeneni Last Modified in 10/26/22	773	34	34	0	16	0	0	1
[147] † Hypselorhachis mirabilis Last Modified in 10/26/22	796	26	25	0	2	0	0	1
[148] † Waldhaus poposauroid Last Modified in 10/26/22	789	34	33	0	1	0	0	1
[149] † Vytshegdosuchus zbeshartensis Last Modified in 10/26/22	813	9	9	0	1	0	0	1
[150] † Bystrowisuchus flerovi Last Modified in 10/26/22	805	19	16	0	2	0	0	1
[151] † Bromsgroveia walkeri Last Modified in 10/26/22	782	40	40	0	1	0	0	1
[152] † Xilousuchus sapingensis Last Modified in 10/26/22	580	225	224	0	18	0	0	1
[153] † Moenkopi poposauroid Last Modified in 10/26/22	777	44	40	0	6	0	0	1
[154] † Mandasuchus tanyauchen Last Modified in 10/26/22	527	283	278	0	18	0	0	1
[155] † Luperosuchus fractus Last Modified in 10/26/22	699	123	121	0	3	0	0	1
[156] † Decuriasuchus quartacolonia Last Modified in 10/26/22	383	413	408	0	32	0	0	1
[157] † Batrachotomus kupferzellensis Last Modified in 10/26/22	168	626	617	0	34	0	0	1
[158] † Prestosuchus chiniquensis Last Modified in 10/26/22	90	703	698	0	35	0	0	1
[159] † Austriadraco dallavecchiai Last Modified in 10/26/22	640	184	171	0	12	0	0	1
[160] † Preondactylus buffarinii Last Modified in 10/26/22	570	215	212	0	41	0	0	1
[161] † Austriadactylus cristatus Last Modified in 10/26/22	591	206	202	0	30	0	0	1
[162] † Seazzadactylus venieri Last Modified in 10/26/22	558	229	217	0	45	0	0	1
[163] † Raeticodactylus filisurensis Last Modified in 10/26/22	576	226	218	0	29	0	0	1
[164] † Carniadactylus rosenfeldi Last Modified in 10/26/22	481	283	277	0	65	0	0	1
[165] † Peteinosaurus zambellii Last Modified in 10/26/22	582	210	204	0	37	0	0	1
[166] † Caelestiventus hanseni Last Modified in 10/26/22	644	147	143	0	36	0	0	1
[167] † Dimorphodon macronyx Last Modified in 10/26/22	353	407	403	0	67	0	0	1
[168] † Eudimorphodon ranzii Last Modified in 10/26/22	487	298	295	0	41	0	0	1
[169] † Rhamphorhynchus muensteri Last Modified in 10/26/22	168	538	531	0	124	0	0	1
[170] † Cacibupteryx caribensis Last Modified in 10/26/22	657	141	139	0	27	0	0	1
[171] † Allkaruen koi Last Modified in 10/26/22	690	123	121	0	12	0	0	1
[172] † Lagerpeton chanarensis Last Modified in 10/26/22	533	259	256	0	34	0	0	1
[173] † Ixalerpeton polesinensis Last Modified in 10/26/22	451	344	338	0	34	0	0	1
[174] † Kongonaphon kely Last Modified in 10/26/22	741	74	70	0	12	0	0	1
[175] † PVSJ 883 Last Modified in 10/26/22	813	9	9	0	1	0	0	1
[176] † Dromomeron romeri Last Modified in 10/26/22	651	151	143	0	29	0	0	1
[177] † Dromomeron gregorii Last Modified in 10/26/22	684	135	135	0	4	0	0	1
[178] † Dromomeron gigas Last Modified in 10/26/22	787	31	31	0	5	0	0	1
[179] † Lagosuchus talampayensis Last Modified in 10/26/22	409	386	379	0	35	0	0	1
[180] † Lutungutali sitwensis Last Modified in 10/26/22	786	39	34	0	3	0	0	1
[181] † Asilisaurus kongwe Last Modified in 10/26/22	312	484	476	0	35	0	0	1
[182] † Nyasasaurus parringtoni Last Modified in 10/26/22	772	49	48	0	3	0	0	1
[183] † Lewisuchus admixtus Last Modified in 10/26/22	277	525	517	0	29	0	0	1
[184] † Silesaurus opolensis Last Modified in 10/26/22	175	612	603	0	45	0	0	1
[185] † Heterodontosaurus tucki Last Modified in 10/26/22	88	654	645	0	83	0	0	1
[186] † Lesothosaurus diagnosticus Last Modified in 10/26/22	99	656	646	0	75	0	0	1
[187] † Herrerasaurus ischigualastensis Last Modified in 10/26/22	72	706	691	0	56	0	0	1
[188] † Eoraptor lunensis Last Modified in 10/26/22	164	617	612	0	46	0	0	1
[189] † Buriolestes schultzi Last Modified in 10/26/22	106	666	661	0	56	0	0	1
[190] † Saturnalia tupiniquim Last Modified in 10/26/22	305	476	469	0	49	0	0	1
[191] † Tawa hallae Last Modified in 10/26/22	211	554	552	0	58	0	0	1
[192] † Coelophysis bauri Last Modified in 10/26/22	63	695	688	0	71	0	0	1
[193] † Megapnosaurus rhodesiensis Last Modified in 10/26/22	84	673	667	0	72	0	0	1
[194] † Scleromochlus taylori i Last Modified in 10/26/22	466	326	314	0	43	0	0	1
[195] † Scleromochlus taylori ii Last Modified in 10/26/22	466	334	322	0	35	0	0	1
[196] † Scleromochlus taylori iii Last Modified in 10/26/22	463	338	326	0	34	0	0	1
[197] † Scleromochlus taylori noankle Last Modified in 10/26/22	480	321	309	0	34	0	0	1

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	1814
Project overview	219
Documents list	101
Bibliography	59
Taxon list	626
Media views	381	Media search (316 views); M847705 (65 views);
Views for media list	72
Specimen list	124
Matrix views	232	Matrix landing page (219 views); dataset1_Parsimony_TNT_Foffaetal_2022 (13 views);

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	41
Document downloads	13	SI Dataset 2. (4 downloads); SI Dataset 1 (3 downloads); SI Dataset 3 (2 downloads); SI Dataset 4 (2 downloads); SI Dataset 5 (1 download); SI Dataset 6 (1 download);
Project downloads	9
Media downloads	1	M847705 (1 download);
Matrix downloads	18	dataset1_Parsimony_TNT_Foffaetal_2022 (18 downloads);