Morphobank

Project 4397: D. Geraads, S. Zouhri. 2021. A new late Miocene elasmotheriine rhinoceros from Morocco. Acta Palaeontologica Polonica. 66 (4):753-765.

Specimen: † Teleoceras fossiger (unvouchered)
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Abstract

We describe here the first definite representative of the subfamily Elasmotheriinae in North Africa. It comes from theupper Miocene site of Skoura near Ouarzazate, on the southern slope of the Central High Atlas in Morocco. It consistsof a virtually complete skull with articulated mandible and a few fragmentary postcranial remains, making it by far thebest known elasmotheriine from the African late Miocene. We assign it to a new taxon, Eoazara xerrii gen. et sp. nov.The skull is characterized by long nasal bones indicating a strong horn and long, anteriorly expanded, edentulous premaxillae.Compared to other Rhinocerotidae, the face is moderately elongated; the lower incisors are of medium size; andthe premolar row is short. The upper molars have a strongly pinched protocone, a long antecrochet, and an unexpandedcentral valley. Eoazara xerrii gen. et sp. nov. is at a lower evolutionary grade than the Chinese species of Ningxiatherium and Parelasmotherium, but probably comparable to the very incomplete remains from the East African late Mioceneforms. We regard Eoazara as a member of a chiefly Eurasian clade, rather than as a survivor of a hypothetical Africanelasmotheriine branch. Parsimony analysis confirms the monophyly of the Elasmotheriinae, but that of the remainingRhinocerotidae is questionable.

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Article DOI: 10.4202/app.00904.2021

Project DOI: 10.7934/P4397, http://dx.doi.org/10.7934/P4397

This project contains	Matrices
10 Media 1 Matrix 75 Taxa 12 Specimens 223 Characters Total size of project's media files: 41.68M Download Project SDD File	Total scored cells: 11333 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 223 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 537 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:15/208

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MorphoBank Project 4397

Creation Date:
09 September 2022
Publication Date:
16 September 2022
Media downloads: 1
Matrix downloads: 5

Authors' Institutions

Muséum national d'Histoire naturelle, Paris (Natural History Museum, Paris)

Centre National de la Recherche Scientifique (CNRS)

Sorbonne Université

Hassan II University of Casablanca

Members

MorphoBank Curator
Project Administrator

223

11333
(10568, 0, 765)

Taxonomic Overview for Matrix 'M28982' (75 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] Tapirus terrestris Taxon name last Modified on 09/16/22	15	186	186	0	22	0	0	1
[2] † Aceratherium incisivum Taxon name last Modified on 09/16/22	44	172	172	0	7	0	0	1
[3] † Acerorhinus fuguensis Taxon name last Modified on 09/16/22	43	170	170	0	10	0	0	1
[4] † Acerorhinus hezhengensis Taxon name last Modified on 09/16/22	35	176	176	0	12	0	0	1
[5] † Acerorhinus palaeosinensis Taxon name last Modified on 09/16/22	69	146	146	0	8	0	0	1
[6] † Acerorhinus tsaidamensis Taxon name last Modified on 09/16/22	34	176	176	0	13	0	0	1
[7] † Acerorhinus yuanmouensis Taxon name last Modified on 09/16/22	80	134	134	0	9	0	0	1
[8] † Acerorhinus zernowi Taxon name last Modified on 09/16/22	59	155	155	0	9	0	0	1
[9] † Alicornops complanatum Taxon name last Modified on 09/16/22	102	114	114	0	7	0	0	1
[10] † Alicornops simorrense Taxon name last Modified on 09/16/22	35	179	179	0	9	0	0	1
[11] † Aprotodon fatehjangense Taxon name last Modified on 09/16/22	89	130	130	0	4	0	0	1
[12] † Begertherium grimmi Taxon name last Modified on 09/16/22	85	128	128	0	10	0	0	1
[13] † Brachypotherium brachypus Taxon name last Modified on 09/16/22	33	177	177	0	13	0	0	1
[14] † Brachypotherium minor Taxon name last Modified on 09/16/22	92	119	119	0	12	0	0	1
[15] † Caementodon caucasicum Taxon name last Modified on 09/16/22	94	119	119	0	10	0	0	1
[16] Ceratotherium simum Taxon name last Modified on 09/16/22	18	187	187	0	18	0	0	1
[17] † Chilotheridium pattersoni Taxon name last Modified on 09/16/22	96	119	119	0	8	0	0	1
[18] † Chilotherium anderssoni Taxon name last Modified on 09/16/22	16	193	193	0	14	0	0	1
[19] † Chilotherium wimani Taxon name last Modified on 09/16/22	26	182	182	0	15	0	0	1
[20] † Coelodonta antiquitatis Taxon name last Modified on 09/16/22	28	181	181	0	14	0	0	1
[21] † Diaceratherium aginense Taxon name last Modified on 09/16/22	72	142	142	0	9	0	0	1
[22] † Diceratherium armatum Taxon name last Modified on 09/16/22	16	195	195	0	12	0	0	1
[23] Dicerorhinus sumatrensis Taxon name last Modified on 09/16/22	30	179	179	0	14	0	0	1
[24] Diceros bicornis Taxon name last Modified on 09/16/22	13	189	189	0	21	0	0	1
[25] † Elasmotherium sibiricum Taxon name last Modified on 09/16/22	43	149	149	0	31	0	0	1
[26] † Eoazara xerrii Taxon name last Modified on 09/16/22	86	122	122	0	15	0	0	1
[27] † Gaindatherium browni Taxon name last Modified on 09/16/22	57	153	153	0	13	0	0	1
[28] † Hispanotherium beonense Taxon name last Modified on 09/16/22	38	177	177	0	8	0	0	1
[29] † Hispanotherium corcolense Taxon name last Modified on 09/16/22	93	125	125	0	5	0	0	1
[30] † Hispanotherium matritense Taxon name last Modified on 09/16/22	75	142	142	0	6	0	0	1
[31] † Hispanotherium tungurense Taxon name last Modified on 09/16/22	28	184	184	0	11	0	0	1
[32] † Hoploaceratherium tetradactylum Taxon name last Modified on 09/16/22	37	177	177	0	9	0	0	1
[33] † Hyrachyus eximius Taxon name last Modified on 09/16/22	32	168	168	0	23	0	0	1
[34] † Iranotherium morgani Taxon name last Modified on 09/16/22	42	167	167	0	14	0	0	1
[35] † Lartetotherium sansaniense Taxon name last Modified on 09/16/22	44	166	166	0	13	0	0	1
[36] † Menoceras arikarense Taxon name last Modified on 09/16/22	18	194	194	0	11	0	0	1
[37] † Mesaceratherium gaimersheimense Taxon name last Modified on 09/16/22	69	144	144	0	10	0	0	1
[38] † Mesaceratherium paulhiacense Taxon name last Modified on 09/16/22	94	122	122	0	7	0	0	1
[39] † Mesaceratherium welcommi Taxon name last Modified on 09/16/22	73	138	138	0	12	0	0	1
[40] † Molassitherium albigense Taxon name last Modified on 09/16/22	39	174	174	0	10	0	0	1
[41] † Ningxiatherium euryrhinus Taxon name last Modified on 09/16/22	99	117	117	0	7	0	0	1
[42] † Ningxiatherium longirhinus Taxon name last Modified on 09/16/22	103	113	113	0	7	0	0	1
[43] † Parelasmotherium lingxiaense Taxon name last Modified on 09/16/22	90	124	124	0	9	0	0	1
[44] † Plesiaceratherium gracile Taxon name last Modified on 09/16/22	51	159	159	0	13	0	0	1
[45] † Plesiaceratherium mirallesi Taxon name last Modified on 09/16/22	49	165	165	0	9	0	0	1
[46] † Pleuroceros blanfordi Taxon name last Modified on 09/16/22	48	168	168	0	7	0	0	1
[47] † Pleuroceros pleuroceros Taxon name last Modified on 09/16/22	61	156	156	0	6	0	0	1
[48] † Prosantorhinus douvillei Taxon name last Modified on 09/16/22	50	163	163	0	10	0	0	1
[49] † Protaceratherium minutum Taxon name last Modified on 09/16/22	55	158	158	0	10	0	0	1
[50] Rhinoceros sondaicus Taxon name last Modified on 09/16/22	15	192	192	0	16	0	0	1
[51] Rhinoceros unicornis Taxon name last Modified on 09/16/22	17	192	192	0	14	0	0	1
[52] † Ronzotherium filholi Taxon name last Modified on 09/16/22	19	190	190	0	14	0	0	1
[53] † Rusingaceros leakeyi Taxon name last Modified on 09/16/22	92	123	123	0	8	0	0	1
[54] † Shansirhinus ringstromi Taxon name last Modified on 09/16/22	41	170	170	0	12	0	0	1
[55] † Subhyracodon occidentalis Taxon name last Modified on 09/16/22	43	171	171	0	9	0	0	1
[56] † Teleoceras fossiger Taxon name last Modified on 09/16/22	18	190	190	0	15	0	0	1
[57] † Trigonias osborni Taxon name last Modified on 09/16/22	15	192	192	0	16	0	0	1
[58] † Turkanatherium acutirostratum Taxon name last Modified on 09/16/22	88	126	126	0	9	0	0	1
[59] † Victoriaceros kenyensis Taxon name last Modified on 09/16/22	52	157	157	0	14	0	0	1
[60] † Aceratherium huadeensis Taxon name last Modified on 09/16/22	141	77	77	0	5	0	0	1
[61] † Acerorhinus lufengensis Taxon name last Modified on 09/16/22	131	86	86	0	6	0	0	1
[62] † Alicornops alfambrense Taxon name last Modified on 09/16/22	186	36	36	0	1	0	0	1
[63] † Begertherium borissiaki Taxon name last Modified on 09/16/22	155	65	65	0	3	0	0	1
[64] † Beliajevina tekkayai Taxon name last Modified on 09/16/22	118	102	102	0	3	0	0	1
[65] † Bugtirhinus praecursor Taxon name last Modified on 09/16/22	135	85	85	0	3	0	0	1
[66] † Caementodon oettingenae Taxon name last Modified on 09/16/22	126	94	94	0	3	0	0	1
[67] † Elasmotherium caucasicum Taxon name last Modified on 09/16/22	111	90	90	0	22	0	0	1
[68] † Kenyatherium bishopi Taxon name last Modified on 09/16/22	184	38	38	0	1	0	0	1
[69] † Ougandatherium napakense Taxon name last Modified on 09/16/22	146	69	69	0	8	0	0	1
[70] † Parelasmotherium schansiense Taxon name last Modified on 09/16/22	183	39	39	0	1	0	0	1
[71] † Procoelodonta mongoliense Taxon name last Modified on 09/16/22	142	77	77	0	4	0	0	1
[72] † Shennongtherium hyposodontus Taxon name last Modified on 09/16/22	184	37	37	0	2	0	0	1
[73] † Sinotherium lagrelii Taxon name last Modified on 09/16/22	121	95	95	0	7	0	0	1
[74] † Subchilotherium intermedium Taxon name last Modified on 09/16/22	115	102	102	0	6	0	0	1
[75] † Victoriaceros hooijeri Taxon name last Modified on 09/16/22	116	100	100	0	7	0	0	1

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	7
Matrix downloads	5	matrix (5 downloads);
Project downloads	1
Media downloads	1	M846872 (1 download);