Morphobank

Project 4397: D. Geraads, S. Zouhri. 2021. A new late Miocene elasmotheriine rhinoceros from Morocco. Acta Palaeontologica Polonica. 66 (4):753-765.

Specimen: † Teleoceras fossiger (unvouchered)
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Abstract

We describe here the first definite representative of the subfamily Elasmotheriinae in North Africa. It comes from theupper Miocene site of Skoura near Ouarzazate, on the southern slope of the Central High Atlas in Morocco. It consistsof a virtually complete skull with articulated mandible and a few fragmentary postcranial remains, making it by far thebest known elasmotheriine from the African late Miocene. We assign it to a new taxon, Eoazara xerrii gen. et sp. nov.The skull is characterized by long nasal bones indicating a strong horn and long, anteriorly expanded, edentulous premaxillae.Compared to other Rhinocerotidae, the face is moderately elongated; the lower incisors are of medium size; andthe premolar row is short. The upper molars have a strongly pinched protocone, a long antecrochet, and an unexpandedcentral valley. Eoazara xerrii gen. et sp. nov. is at a lower evolutionary grade than the Chinese species of Ningxiatherium and Parelasmotherium, but probably comparable to the very incomplete remains from the East African late Mioceneforms. We regard Eoazara as a member of a chiefly Eurasian clade, rather than as a survivor of a hypothetical Africanelasmotheriine branch. Parsimony analysis confirms the monophyly of the Elasmotheriinae, but that of the remainingRhinocerotidae is questionable.

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Article DOI: 10.4202/app.00904.2021

Project DOI: 10.7934/P4397, http://dx.doi.org/10.7934/P4397

This project contains	Matrices
10 Media 1 Matrix 75 Taxa 12 Specimens 223 Characters Total size of project's media files: 41.68M Download Project SDD File	Total scored cells: 11333 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 223 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 537 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:15/208

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MorphoBank Project 4397

Creation Date:
09 September 2022
Publication Date:
16 September 2022
Project views: 9109
Media downloads: 1
Matrix downloads: 2

Authors' Institutions

Muséum national d'Histoire naturelle, Paris (Natural History Museum, Paris)

Centre National de la Recherche Scientifique (CNRS)

Sorbonne Université

Hassan II University of Casablanca

Members

MorphoBank Curator
Project Administrator

223

11333
(10568, 0, 765)

Taxonomic Overview for Matrix 'M28982' (75 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] Tapirus terrestris Last Modified in 09/16/22	15	186	186	0	22	0	0	1
[2] † Aceratherium incisivum Last Modified in 09/16/22	44	172	172	0	7	0	0	1
[3] † Acerorhinus fuguensis Last Modified in 09/16/22	43	170	170	0	10	0	0	1
[4] † Acerorhinus hezhengensis Last Modified in 09/16/22	35	176	176	0	12	0	0	1
[5] † Acerorhinus palaeosinensis Last Modified in 09/16/22	69	146	146	0	8	0	0	1
[6] † Acerorhinus tsaidamensis Last Modified in 09/16/22	34	176	176	0	13	0	0	1
[7] † Acerorhinus yuanmouensis Last Modified in 09/16/22	80	134	134	0	9	0	0	1
[8] † Acerorhinus zernowi Last Modified in 09/16/22	59	155	155	0	9	0	0	1
[9] † Alicornops complanatum Last Modified in 09/16/22	102	114	114	0	7	0	0	1
[10] † Alicornops simorrense Last Modified in 09/16/22	35	179	179	0	9	0	0	1
[11] † Aprotodon fatehjangense Last Modified in 09/16/22	89	130	130	0	4	0	0	1
[12] † Begertherium grimmi Last Modified in 09/16/22	85	128	128	0	10	0	0	1
[13] † Brachypotherium brachypus Last Modified in 09/16/22	33	177	177	0	13	0	0	1
[14] † Brachypotherium minor Last Modified in 09/16/22	92	119	119	0	12	0	0	1
[15] † Caementodon caucasicum Last Modified in 09/16/22	94	119	119	0	10	0	0	1
[16] Ceratotherium simum Last Modified in 09/16/22	18	187	187	0	18	0	0	1
[17] † Chilotheridium pattersoni Last Modified in 09/16/22	96	119	119	0	8	0	0	1
[18] † Chilotherium anderssoni Last Modified in 09/16/22	16	193	193	0	14	0	0	1
[19] † Chilotherium wimani Last Modified in 09/16/22	26	182	182	0	15	0	0	1
[20] † Coelodonta antiquitatis Last Modified in 09/16/22	28	181	181	0	14	0	0	1
[21] † Diaceratherium aginense Last Modified in 09/16/22	72	142	142	0	9	0	0	1
[22] † Diceratherium armatum Last Modified in 09/16/22	16	195	195	0	12	0	0	1
[23] Dicerorhinus sumatrensis Last Modified in 09/16/22	30	179	179	0	14	0	0	1
[24] Diceros bicornis Last Modified in 09/16/22	13	189	189	0	21	0	0	1
[25] † Elasmotherium sibiricum Last Modified in 09/16/22	43	149	149	0	31	0	0	1
[26] † Eoazara xerrii Last Modified in 09/16/22	86	122	122	0	15	0	0	1
[27] † Gaindatherium browni Last Modified in 09/16/22	57	153	153	0	13	0	0	1
[28] † Hispanotherium beonense Last Modified in 09/16/22	38	177	177	0	8	0	0	1
[29] † Hispanotherium corcolense Last Modified in 09/16/22	93	125	125	0	5	0	0	1
[30] † Hispanotherium matritense Last Modified in 09/16/22	75	142	142	0	6	0	0	1
[31] † Hispanotherium tungurense Last Modified in 09/16/22	28	184	184	0	11	0	0	1
[32] † Hoploaceratherium tetradactylum Last Modified in 09/16/22	37	177	177	0	9	0	0	1
[33] † Hyrachyus eximius Last Modified in 09/16/22	32	168	168	0	23	0	0	1
[34] † Iranotherium morgani Last Modified in 09/16/22	42	167	167	0	14	0	0	1
[35] † Lartetotherium sansaniense Last Modified in 09/16/22	44	166	166	0	13	0	0	1
[36] † Menoceras arikarense Last Modified in 09/16/22	18	194	194	0	11	0	0	1
[37] † Mesaceratherium gaimersheimense Last Modified in 09/16/22	69	144	144	0	10	0	0	1
[38] † Mesaceratherium paulhiacense Last Modified in 09/16/22	94	122	122	0	7	0	0	1
[39] † Mesaceratherium welcommi Last Modified in 09/16/22	73	138	138	0	12	0	0	1
[40] † Molassitherium albigense Last Modified in 09/16/22	39	174	174	0	10	0	0	1
[41] † Ningxiatherium euryrhinus Last Modified in 09/16/22	99	117	117	0	7	0	0	1
[42] † Ningxiatherium longirhinus Last Modified in 09/16/22	103	113	113	0	7	0	0	1
[43] † Parelasmotherium lingxiaense Last Modified in 09/16/22	90	124	124	0	9	0	0	1
[44] † Plesiaceratherium gracile Last Modified in 09/16/22	51	159	159	0	13	0	0	1
[45] † Plesiaceratherium mirallesi Last Modified in 09/16/22	49	165	165	0	9	0	0	1
[46] † Pleuroceros blanfordi Last Modified in 09/16/22	48	168	168	0	7	0	0	1
[47] † Pleuroceros pleuroceros Last Modified in 09/16/22	61	156	156	0	6	0	0	1
[48] † Prosantorhinus douvillei Last Modified in 09/16/22	50	163	163	0	10	0	0	1
[49] † Protaceratherium minutum Last Modified in 09/16/22	55	158	158	0	10	0	0	1
[50] Rhinoceros sondaicus Last Modified in 09/16/22	15	192	192	0	16	0	0	1
[51] Rhinoceros unicornis Last Modified in 09/16/22	17	192	192	0	14	0	0	1
[52] † Ronzotherium filholi Last Modified in 09/16/22	19	190	190	0	14	0	0	1
[53] † Rusingaceros leakeyi Last Modified in 09/16/22	92	123	123	0	8	0	0	1
[54] † Shansirhinus ringstromi Last Modified in 09/16/22	41	170	170	0	12	0	0	1
[55] † Subhyracodon occidentalis Last Modified in 09/16/22	43	171	171	0	9	0	0	1
[56] † Teleoceras fossiger Last Modified in 09/16/22	18	190	190	0	15	0	0	1
[57] † Trigonias osborni Last Modified in 09/16/22	15	192	192	0	16	0	0	1
[58] † Turkanatherium acutirostratum Last Modified in 09/16/22	88	126	126	0	9	0	0	1
[59] † Victoriaceros kenyensis Last Modified in 09/16/22	52	157	157	0	14	0	0	1
[60] † Aceratherium huadeensis Last Modified in 09/16/22	141	77	77	0	5	0	0	1
[61] † Acerorhinus lufengensis Last Modified in 09/16/22	131	86	86	0	6	0	0	1
[62] † Alicornops alfambrense Last Modified in 09/16/22	186	36	36	0	1	0	0	1
[63] † Begertherium borissiaki Last Modified in 09/16/22	155	65	65	0	3	0	0	1
[64] † Beliajevina tekkayai Last Modified in 09/16/22	118	102	102	0	3	0	0	1
[65] † Bugtirhinus praecursor Last Modified in 09/16/22	135	85	85	0	3	0	0	1
[66] † Caementodon oettingenae Last Modified in 09/16/22	126	94	94	0	3	0	0	1
[67] † Elasmotherium caucasicum Last Modified in 09/16/22	111	90	90	0	22	0	0	1
[68] † Kenyatherium bishopi Last Modified in 09/16/22	184	38	38	0	1	0	0	1
[69] † Ougandatherium napakense Last Modified in 09/16/22	146	69	69	0	8	0	0	1
[70] † Parelasmotherium schansiense Last Modified in 09/16/22	183	39	39	0	1	0	0	1
[71] † Procoelodonta mongoliense Last Modified in 09/16/22	142	77	77	0	4	0	0	1
[72] † Shennongtherium hyposodontus Last Modified in 09/16/22	184	37	37	0	2	0	0	1
[73] † Sinotherium lagrelii Last Modified in 09/16/22	121	95	95	0	7	0	0	1
[74] † Subchilotherium intermedium Last Modified in 09/16/22	115	102	102	0	6	0	0	1
[75] † Victoriaceros hooijeri Last Modified in 09/16/22	116	100	100	0	7	0	0	1

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	9109
Project overview	446
Media views	2281	Media search (794 views); M846869 (152 views); M846868 (143 views); M846871 (127 views); M846870 (159 views); M846863 (151 views); M846862 (143 views); M846872 (149 views); M846867 (150 views); M846873 (156 views); M846866 (157 views);
Matrix views	267	Matrix landing page (247 views); matrix (20 views);
Taxon list	2742
Views for media list	550
Bibliography	205
Specimen list	2613
Documents list	5

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	4
Matrix downloads	2	matrix (2 downloads);
Project downloads	1
Media downloads	1	M846872 (1 download);