Project 4400: I. Yáñez, D. Pol, J. Leardi, O. Alcober, R. Martinez. 2021. An enigmatic new archosauriform from the Carnian-Norian Ischigualasto Formation of Northwestern Argentina. Acta Palaeontologica Polonica. 66 (3):509-533.
Abstract
In this contribution we introduce a new Late Triassic archosaur, Incertovenator longicollum gen. et sp. nov., with an unusual combination of character states that are present in certain early avemetatarsalian and pseudosuchian archosaur clades. The holotype consists of a partial postcranial skeleton, preserving most of the axial skeleton and displaying a marked anteroposterior elongation in the cervical vertebrae. We include I. longicollum gen. et sp. nov. into one of the most comprehensive early archosaur phylogenetic data sets available, and recover it as either an early diverging avemetatarsalian, closely associated with the clade Aphanosauria and Ornithodira, or as an early diverging loricatan closely related to Mandasuchus tanyauchen in the most parsimonious trees. We further evaluate which alternative phylogenetic positions can I. longicollum gen. et sp. nov. take in the suboptimal trees, and determined which character states support those alternative positions in comparison with those of the unconstrained analysis. The analyses recover the new taxon in three main general phylogenetic placements within Archosauria, as well as one position outside this clade, highlighting widespread morphological evolutionary convergence towards neck elongation in several clades of Triassic archosauriforms.Read the article »
Article DOI: 10.4202/app.00806.2020
Project DOI: 10.7934/P4400, http://dx.doi.org/10.7934/P4400
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 22907 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 419 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 916 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:400/19 |
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MorphoBank Project 4400
MorphoBank Project 4400
- Creation Date:
12 September 2022 - Publication Date:
31 October 2022 - Matrix downloads: 3
Authors' Institutions
- Museo Paleontológico Egidio Feruglio
- Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)
- Universidad de Buenos Aires
- Universidad Nacional de San Juan
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| MorphoBank Curator Project Administrator | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| MorphoBank Curator Full membership | 99 | 1 | 1 | 419 | 0 | 0 | 22907 (22815, 0, 92) | 0 | 0 | 0 | ||||
| Imanol Yáñez Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M29022' (99 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] † Mesosuchus browni Taxon name last Modified on 10/31/22 | 84 | 335 | 335 | 0 | 0 | 0 | 0 | 3 |
| [2] † Prolacerta broomi Taxon name last Modified on 10/31/22 | 56 | 363 | 363 | 0 | 0 | 0 | 0 | 3 |
| [3] † Archosaurus rossicus Taxon name last Modified on 10/31/22 | 414 | 5 | 5 | 0 | 0 | 0 | 0 | 3 |
| [4] † Proterosuchus Taxon name last Modified on 10/31/22 | 51 | 368 | 368 | 0 | 0 | 0 | 0 | 3 |
| [5] † Erythrosuchus africanus Taxon name last Modified on 10/31/22 | 50 | 369 | 369 | 0 | 0 | 0 | 0 | 3 |
| [6] † Vancleavea campi Taxon name last Modified on 10/31/22 | 134 | 285 | 285 | 0 | 0 | 0 | 0 | 3 |
| [7] † Chanaresuchus bonapartei Taxon name last Modified on 10/31/22 | 53 | 367 | 366 | 0 | 0 | 0 | 0 | 3 |
| [8] † Tropidosuchus romeri Taxon name last Modified on 10/31/22 | 103 | 317 | 316 | 0 | 0 | 0 | 0 | 3 |
| [9] † Euparkeria capensis Taxon name last Modified on 10/31/22 | 18 | 402 | 401 | 0 | 0 | 0 | 0 | 3 |
| [10] † Parasuchus hislopi Taxon name last Modified on 10/31/22 | 66 | 353 | 353 | 0 | 0 | 0 | 0 | 3 |
| [11] † Smilosuchus gregorii Taxon name last Modified on 10/31/22 | 76 | 343 | 343 | 0 | 0 | 0 | 0 | 3 |
| [12] † Pseudopalatus pristinus Taxon name last Modified on 10/31/22 | 33 | 386 | 386 | 0 | 0 | 0 | 0 | 3 |
| [13] † Gracilisuchus stipanicicorum Taxon name last Modified on 10/31/22 | 108 | 315 | 310 | 0 | 1 | 0 | 0 | 3 |
| [14] † Turfanosuchus dabanensis Taxon name last Modified on 10/31/22 | 147 | 273 | 272 | 0 | 0 | 0 | 0 | 3 |
| [15] † Yonghesuchus sangbiensis Taxon name last Modified on 10/31/22 | 328 | 91 | 90 | 0 | 1 | 0 | 0 | 3 |
| [16] † Incertovenator longicollum Taxon name last Modified on 10/31/22 | 382 | 35 | 35 | 0 | 2 | 0 | 0 | 3 |
| [17] † Ornithosuchus longidens Taxon name last Modified on 10/31/22 | 140 | 279 | 279 | 0 | 0 | 0 | 0 | 3 |
| [18] † Riojasuchus tenuisceps Taxon name last Modified on 10/31/22 | 62 | 357 | 357 | 0 | 0 | 0 | 0 | 3 |
| [19] † Revueltosaurus callenderi Taxon name last Modified on 10/31/22 | 47 | 372 | 372 | 0 | 0 | 0 | 0 | 3 |
| [20] † Stagonolepis robertsoni Taxon name last Modified on 10/31/22 | 101 | 318 | 318 | 0 | 0 | 0 | 0 | 3 |
| [21] † Aetosaurus ferratus Taxon name last Modified on 10/31/22 | 152 | 268 | 267 | 0 | 0 | 0 | 0 | 3 |
| [22] † Longosuchus meadei Taxon name last Modified on 10/31/22 | 70 | 349 | 349 | 0 | 0 | 0 | 0 | 3 |
| [23] † Ticinosuchus ferox Taxon name last Modified on 10/31/22 | 211 | 208 | 208 | 0 | 0 | 0 | 0 | 3 |
| [24] † Qianosuchus mixtus Taxon name last Modified on 10/31/22 | 131 | 288 | 288 | 0 | 0 | 0 | 0 | 3 |
| [25] † Xilousuchus sapingensis Taxon name last Modified on 10/31/22 | 300 | 119 | 119 | 0 | 0 | 0 | 0 | 3 |
| [26] † Arizonasaurus babbitti Taxon name last Modified on 10/31/22 | 163 | 256 | 256 | 0 | 0 | 0 | 0 | 3 |
| [27] † Poposaurus gracilis holotype Taxon name last Modified on 10/31/22 | 380 | 38 | 38 | 0 | 1 | 0 | 0 | 3 |
| [28] † Poposaurus gracilis yale Taxon name last Modified on 10/31/22 | 202 | 217 | 216 | 0 | 1 | 0 | 0 | 3 |
| [29] † Lotosaurus adentus Taxon name last Modified on 10/31/22 | 103 | 316 | 316 | 0 | 0 | 0 | 0 | 3 |
| [30] † Sillosuchus longicervix Taxon name last Modified on 10/31/22 | 338 | 80 | 80 | 0 | 1 | 0 | 0 | 3 |
| [31] † Effigia okeeffeae Taxon name last Modified on 10/31/22 | 98 | 320 | 320 | 0 | 1 | 0 | 0 | 3 |
| [32] † Shuvosaurus inexpectatus Taxon name last Modified on 10/31/22 | 107 | 311 | 311 | 0 | 1 | 0 | 0 | 3 |
| [33] † Prestosuchus chiniquensis Taxon name last Modified on 10/31/22 | 245 | 174 | 174 | 0 | 0 | 0 | 0 | 3 |
| [34] † UFRGS 0156 T Taxon name last Modified on 10/31/22 | 251 | 168 | 168 | 0 | 0 | 0 | 0 | 3 |
| [35] † UFRGS 152 T Taxon name last Modified on 10/31/22 | 151 | 268 | 268 | 0 | 0 | 0 | 0 | 3 |
| [36] † Combined prestosuchus Taxon name last Modified on 10/31/22 | 61 | 358 | 358 | 0 | 0 | 0 | 0 | 3 |
| [37] † Saurosuchus galilei Taxon name last Modified on 10/31/22 | 109 | 310 | 310 | 0 | 0 | 0 | 0 | 3 |
| [38] † Batrachotomus kuperferzellensis Taxon name last Modified on 10/31/22 | 63 | 356 | 356 | 0 | 0 | 0 | 0 | 3 |
| [39] † Fasolasuchus tenax Taxon name last Modified on 10/31/22 | 256 | 163 | 163 | 0 | 0 | 0 | 0 | 3 |
| [40] † Rauisuchus triradentes Taxon name last Modified on 10/31/22 | 285 | 134 | 134 | 0 | 0 | 0 | 0 | 3 |
| [41] † Polonosuchus silesiacus Taxon name last Modified on 10/31/22 | 300 | 119 | 119 | 0 | 0 | 0 | 0 | 3 |
| [42] † Postosuchus kirkpatricki Taxon name last Modified on 10/31/22 | 76 | 343 | 343 | 0 | 0 | 0 | 0 | 3 |
| [43] † Postosuchus alisonae Taxon name last Modified on 10/31/22 | 249 | 170 | 170 | 0 | 0 | 0 | 0 | 3 |
| [44] † CM 73372 Taxon name last Modified on 10/31/22 | 255 | 163 | 163 | 0 | 1 | 0 | 0 | 3 |
| [45] † Hesperosuchus agilis AMNH-6758 Taxon name last Modified on 10/31/22 | 200 | 219 | 219 | 0 | 0 | 0 | 0 | 3 |
| [46] † Dromicosuchus grallator Taxon name last Modified on 10/31/22 | 145 | 274 | 274 | 0 | 0 | 0 | 0 | 3 |
| [47] † Hesperosuchus agilis UCMP-12947 Taxon name last Modified on 10/31/22 | 131 | 288 | 288 | 0 | 0 | 0 | 0 | 3 |
| [48] † Dibothrosuchus elaphros Taxon name last Modified on 10/31/22 | 198 | 220 | 220 | 0 | 1 | 0 | 0 | 3 |
| [49] † Terrestrisuchus gracilis Taxon name last Modified on 10/31/22 | 100 | 318 | 318 | 0 | 1 | 0 | 0 | 3 |
| [50] † Sphenosuchus acutus Taxon name last Modified on 10/31/22 | 201 | 218 | 218 | 0 | 0 | 0 | 0 | 3 |
| [51] † Litargosuchus leptorhynchus Taxon name last Modified on 10/31/22 | 229 | 190 | 190 | 0 | 0 | 0 | 0 | 3 |
| [52] † Kayentasuchus walkeri Taxon name last Modified on 10/31/22 | 310 | 108 | 108 | 0 | 1 | 0 | 0 | 3 |
| [53] † Orthosuchus stormbergi Taxon name last Modified on 10/31/22 | 96 | 322 | 322 | 0 | 1 | 0 | 0 | 3 |
| [54] Alligator mississippiensis Taxon name last Modified on 10/31/22 | 46 | 374 | 372 | 0 | 1 | 0 | 0 | 3 |
| [55] † Protosuchus haughtoni Taxon name last Modified on 10/31/22 | 260 | 159 | 159 | 0 | 0 | 0 | 0 | 3 |
| [56] † Protosuchus richardsoni Taxon name last Modified on 10/31/22 | 54 | 364 | 364 | 0 | 1 | 0 | 0 | 3 |
| [57] † Eudimorphodon ranzii Taxon name last Modified on 10/31/22 | 229 | 190 | 190 | 0 | 0 | 0 | 0 | 3 |
| [58] † Dimorphodon macronyx Taxon name last Modified on 10/31/22 | 219 | 200 | 200 | 0 | 0 | 0 | 0 | 3 |
| [59] † Lagerpeton chanarensis Taxon name last Modified on 10/31/22 | 288 | 131 | 131 | 0 | 0 | 0 | 0 | 3 |
| [60] † Dromomeron gregorii Taxon name last Modified on 10/31/22 | 376 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [61] † Dromomeron romeri Taxon name last Modified on 10/31/22 | 356 | 63 | 63 | 0 | 0 | 0 | 0 | 3 |
| [62] † Marasuchus lilloensis Taxon name last Modified on 10/31/22 | 215 | 204 | 204 | 0 | 0 | 0 | 0 | 3 |
| [63] † Eucoelophysis baldwini Taxon name last Modified on 10/31/22 | 392 | 27 | 27 | 0 | 0 | 0 | 0 | 3 |
| [64] † Sacisaurus agudoensis Taxon name last Modified on 10/31/22 | 327 | 92 | 92 | 0 | 0 | 0 | 0 | 3 |
| [65] † Lewisuchus admixtus Taxon name last Modified on 10/31/22 | 286 | 133 | 133 | 0 | 0 | 0 | 0 | 3 |
| [66] † Pseudolagosuchus majori Taxon name last Modified on 10/31/22 | 322 | 97 | 97 | 0 | 0 | 0 | 0 | 3 |
| [67] † Lewisuchus pseudolagosuchus Taxon name last Modified on 10/31/22 | 191 | 228 | 228 | 0 | 0 | 0 | 0 | 3 |
| [68] † Eocursor parvus Taxon name last Modified on 10/31/22 | 276 | 142 | 142 | 0 | 1 | 0 | 0 | 3 |
| [69] † Silesaurus opolensis Taxon name last Modified on 10/31/22 | 80 | 340 | 339 | 0 | 0 | 0 | 0 | 3 |
| [70] † Pisanosaurus mertii Taxon name last Modified on 10/31/22 | 331 | 88 | 88 | 0 | 0 | 0 | 0 | 3 |
| [71] † Heterodontosaurus tucki Taxon name last Modified on 10/31/22 | 74 | 346 | 344 | 0 | 1 | 0 | 0 | 3 |
| [72] † Lesothosaurus diagnosticus Taxon name last Modified on 10/31/22 | 88 | 331 | 330 | 0 | 1 | 0 | 0 | 3 |
| [73] † Scutellosaurus lawleri Taxon name last Modified on 10/31/22 | 248 | 171 | 170 | 0 | 1 | 0 | 0 | 3 |
| [74] † Herrerasaurus ischigualastensis Taxon name last Modified on 10/31/22 | 69 | 349 | 349 | 0 | 1 | 0 | 0 | 3 |
| [75] † Staurikosaurus pricei Taxon name last Modified on 10/31/22 | 301 | 117 | 117 | 0 | 1 | 0 | 0 | 3 |
| [76] † Eoraptor lunensis Taxon name last Modified on 10/31/22 | 134 | 285 | 284 | 0 | 1 | 0 | 0 | 3 |
| [77] † Saturnalia tupiniquim Taxon name last Modified on 10/31/22 | 222 | 196 | 196 | 0 | 1 | 0 | 0 | 3 |
| [78] † Plateosaurus engelhardti Taxon name last Modified on 10/31/22 | 30 | 388 | 388 | 0 | 1 | 0 | 0 | 3 |
| [79] † Efraasia minor Taxon name last Modified on 10/31/22 | 95 | 323 | 323 | 0 | 1 | 0 | 0 | 3 |
| [80] † Tawa hallae Taxon name last Modified on 10/31/22 | 71 | 347 | 347 | 0 | 1 | 0 | 0 | 3 |
| [81] † Coelophysis bauri Taxon name last Modified on 10/31/22 | 52 | 366 | 366 | 0 | 1 | 0 | 0 | 3 |
| [82] † Dilophosaurus wetherelli Taxon name last Modified on 10/31/22 | 76 | 342 | 342 | 0 | 1 | 0 | 0 | 3 |
| [83] † Allosaurus fragilis Taxon name last Modified on 10/31/22 | 41 | 377 | 377 | 0 | 1 | 0 | 0 | 3 |
| [84] † Velociraptor mongoliensis Taxon name last Modified on 10/31/22 | 83 | 336 | 335 | 0 | 1 | 0 | 0 | 3 |
| [85] † Parringtonia gracilis Taxon name last Modified on 10/31/22 | 381 | 38 | 38 | 0 | 0 | 0 | 0 | 3 |
| [86] † Erpetosuchus granti Taxon name last Modified on 10/31/22 | 266 | 153 | 153 | 0 | 0 | 0 | 0 | 3 |
| [87] † Asilisaurus kongwe Taxon name last Modified on 10/31/22 | 227 | 177 | 177 | 0 | 15 | 0 | 0 | 3 |
| [88] † Asilisaurus kongwe skeleton Taxon name last Modified on 10/31/22 | 165 | 238 | 238 | 0 | 16 | 0 | 0 | 3 |
| [89] † Asilisaurus kongwe combined Taxon name last Modified on 10/31/22 | 136 | 268 | 267 | 0 | 16 | 0 | 0 | 3 |
| [90] † Teleocrater holotype Taxon name last Modified on 10/31/22 | 338 | 81 | 81 | 0 | 0 | 0 | 0 | 3 |
| [91] † Teleocrater 2015 quarry Taxon name last Modified on 10/31/22 | 258 | 161 | 161 | 0 | 0 | 0 | 0 | 3 |
| [92] † Teleocrater combined Taxon name last Modified on 10/31/22 | 248 | 171 | 171 | 0 | 0 | 0 | 0 | 3 |
| [93] † Dongusuchus efremovi Taxon name last Modified on 10/31/22 | 391 | 28 | 28 | 0 | 0 | 0 | 0 | 3 |
| [94] † Spondylosoma absconditum Taxon name last Modified on 10/31/22 | 382 | 37 | 37 | 0 | 0 | 0 | 0 | 3 |
| [95] † Yarasuchus deccanensis Taxon name last Modified on 10/31/22 | 287 | 129 | 129 | 0 | 3 | 0 | 0 | 3 |
| [96] † Scleromochlus taylori Taxon name last Modified on 10/31/22 | 334 | 85 | 85 | 0 | 0 | 0 | 0 | 3 |
| [97] † Mandasuchus total Taxon name last Modified on 10/31/22 | 262 | 155 | 154 | 0 | 3 | 0 | 0 | 3 |
| [98] † Nundasuchus Taxon name last Modified on 10/31/22 | 224 | 191 | 191 | 0 | 4 | 0 | 0 | 3 |
| [99] † Trialestes romeri Taxon name last Modified on 10/31/22 | 249 | 168 | 166 | 0 | 4 | 0 | 0 | 3 |
Project downloads
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 4 | |
| Matrix downloads | 3 | Matrix from Yanez et al 2021 - (prepared by MorphoBank curators from original submission) (3 downloads); |
| Project downloads | 1 |