Project 4438: K. D. Angielczyk, J. Liu, W. Yang. 2021. A redescription of Kunpania scopulusa, a bidentalian dicynodont (Therapsida, Anomodontia) from the Guadalupian of northwestern China. Journal of Vertebrate Paleontology. 41 (1):null.
Abstract
Permian dicynodonts were discovered in strata exposed on the flanks of the Bogda Mountains (Xinjiang, China) in 1928. Nearly all known specimens were collected in the Guodikeng Formation (= upper Wutonggou low-order cycle); the single exception is the holotype of Kunpania scopulusa, which originated in the underlying Quanzijie Formation. The Quanzijie Formation is generally considered Capitanian in age, which would make Kunpania one of the oldest known bidentalian dicynodonts, but the type of K. scopulusa has received little attention. Here we redescribe K. scopulusa and investigate its phylogenetic relationships. Kunpania scopulusa is a valid species diagnosed by the absence of anterior palatal ridges; a robust crista oesophagea; a prominent lateral dentary shelf; a posteriorly curving projection on the angular; and a large, triangular attachment area for M. latissimus dorsi on the humerus. Our phylogenetic analysis recovered K. scopulusa as a stemward member of Dicynodontoidea and the sister taxon of the South African species Sintocephalus alticeps. Depending on the precise age of the uppermost Quanzijie Formation, three scenarios describe the potential phylogenetic implications of K. scopulusa: (1) bidentalian dicynodonts diversified before the end-Guadalupian extinction, with little of their early history preserved; (2) the main diversifications of cryptodont and dicynodontoid bidentalians occurred in the aftermath of the end-Guadalupian extinction, with K. scopulusa representing the latter; (3) K. scopulusa is a contemporary of other mid–late Wuchiapingian dicynodontoids and the initial radiation of Dicynodontoidea is not preserved in the known fossil record.Read the article »
Article DOI: 10.1080/02724634.2021.1922428
Project DOI: 10.7934/P4438, http://dx.doi.org/10.7934/P4438
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 13807 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 174 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 418 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:174/0 |
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MorphoBank Project 4438
MorphoBank Project 4438
- Creation Date:
09 October 2022 - Publication Date:
24 August 2023 - Matrix downloads: 1
This research
supported by
Authors' Institutions
- Institute of Vertebrate Paleontology and Paleoanthropology (IVPP), CAS
- Field Museum of Natural History
- Missouri University of Science and Technology
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Bidane Martinez Huerta Project Administrator | 114 | 1 | 1 | 174 | 0 | 0 | 13807 (13807, 0, 0) | 0 | 0 | 0 | ||||
Brooke Long-Fox Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29024' (114 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Biarmosuchus tener Taxon name last Modified on 10/15/22 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[2] † Hipposaurus boonstrai Taxon name last Modified on 08/24/23 | 24 | 150 | 150 | 0 | 0 | 0 | 0 | 2 |
[3] † Archaeosyodon praeventor Taxon name last Modified on 10/15/22 | 67 | 107 | 107 | 0 | 0 | 0 | 0 | 2 |
[4] † Titanophoneus potens Taxon name last Modified on 08/24/23 | 15 | 160 | 159 | 0 | 0 | 0 | 0 | 2 |
[5] † Gorgonops torvus Taxon name last Modified on 08/24/23 | 12 | 162 | 162 | 0 | 0 | 0 | 0 | 2 |
[6] † Lycosuchus vanderrieti Taxon name last Modified on 08/24/23 | 20 | 154 | 154 | 0 | 0 | 0 | 0 | 2 |
[7] † Glanosuchus macrops Taxon name last Modified on 08/24/23 | 12 | 162 | 162 | 0 | 0 | 0 | 0 | 2 |
[8] † Biseridens qilianicus Taxon name last Modified on 10/15/22 | 59 | 115 | 115 | 0 | 0 | 0 | 0 | 2 |
[9] † Anomocephalus africanus Taxon name last Modified on 10/15/22 | 146 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[10] † Tiarajudens eccentricus Taxon name last Modified on 08/24/23 | 120 | 54 | 54 | 0 | 0 | 0 | 0 | 2 |
[11] † Otsheria netzvetajevi Taxon name last Modified on 08/24/23 | 88 | 86 | 86 | 0 | 0 | 0 | 0 | 2 |
[12] † Ulemica Taxon name last Modified on 08/24/23 | 48 | 127 | 126 | 0 | 0 | 0 | 0 | 2 |
[13] † Suminia getmanovi Taxon name last Modified on 08/24/23 | 20 | 154 | 154 | 0 | 0 | 0 | 0 | 2 |
[14] † Patranomodon nyaphulii Taxon name last Modified on 08/24/23 | 44 | 130 | 130 | 0 | 0 | 0 | 0 | 2 |
[15] † Galeops whaitsi Taxon name last Modified on 10/15/22 | 56 | 118 | 118 | 0 | 0 | 0 | 0 | 2 |
[16] † Galepus jouberti Taxon name last Modified on 10/15/22 | 134 | 40 | 40 | 0 | 0 | 0 | 0 | 2 |
[17] † Galechirus scholtzi Taxon name last Modified on 10/15/22 | 117 | 57 | 57 | 0 | 0 | 0 | 0 | 2 |
[18] † Eodicynodon oelofseni Taxon name last Modified on 10/15/22 | 80 | 94 | 94 | 0 | 0 | 0 | 0 | 2 |
[19] † Eodicynodon oosthuizeni Taxon name last Modified on 10/15/22 | 7 | 167 | 167 | 0 | 0 | 0 | 0 | 2 |
[20] † Colobodectes cluveri Taxon name last Modified on 10/15/22 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 2 |
[21] † Lanthanostegus mohoii Taxon name last Modified on 08/24/23 | 115 | 59 | 59 | 0 | 0 | 0 | 0 | 2 |
[22] † Eosimops newtoni Taxon name last Modified on 10/15/22 | 17 | 159 | 157 | 0 | 0 | 0 | 0 | 2 |
[23] † Diictodon feliceps Taxon name last Modified on 10/15/22 | 9 | 168 | 165 | 0 | 0 | 0 | 0 | 2 |
[24] † Robertia broomiana Taxon name last Modified on 08/24/23 | 13 | 161 | 161 | 0 | 0 | 0 | 0 | 2 |
[25] † Prosictodon dubei Taxon name last Modified on 08/24/23 | 49 | 125 | 125 | 0 | 0 | 0 | 0 | 2 |
[26] † Endothiodon bathystoma Taxon name last Modified on 10/15/22 | 15 | 162 | 159 | 0 | 0 | 0 | 0 | 2 |
[27] † Endothiodon tolani Taxon name last Modified on 10/15/22 | 48 | 127 | 126 | 0 | 0 | 0 | 0 | 2 |
[28] † Niassodon mfumukasi Taxon name last Modified on 08/24/23 | 44 | 130 | 130 | 0 | 0 | 0 | 0 | 2 |
[29] † Brachyprosopus broomi Taxon name last Modified on 10/15/22 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[30] † Pristerodon mackayi Taxon name last Modified on 08/24/23 | 8 | 166 | 166 | 0 | 0 | 0 | 0 | 2 |
[31] † Emydops Taxon name last Modified on 10/16/22 | 9 | 166 | 165 | 0 | 0 | 0 | 0 | 2 |
[32] † Digalodon rubidgei Taxon name last Modified on 10/15/22 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[33] † Dicynodontoides Taxon name last Modified on 10/16/22 | 9 | 165 | 165 | 0 | 0 | 0 | 0 | 2 |
[34] † Kombuisia frerensis Taxon name last Modified on 08/24/23 | 55 | 119 | 119 | 0 | 0 | 0 | 0 | 2 |
[35] † Myosaurus gracilis Taxon name last Modified on 08/24/23 | 28 | 146 | 146 | 0 | 0 | 0 | 0 | 2 |
[36] † Cistecephalus microrhinus Taxon name last Modified on 10/15/22 | 12 | 164 | 162 | 0 | 0 | 0 | 0 | 2 |
[37] † Cistecephaloides boonstrai Taxon name last Modified on 10/15/22 | 52 | 122 | 122 | 0 | 0 | 0 | 0 | 2 |
[38] † Kawingasaurus fossilis Taxon name last Modified on 08/24/23 | 35 | 139 | 139 | 0 | 0 | 0 | 0 | 2 |
[39] † Daqingshanodon limbus Taxon name last Modified on 10/15/22 | 52 | 122 | 122 | 0 | 0 | 0 | 0 | 2 |
[40] † Keyseria benjamini Taxon name last Modified on 08/24/23 | 75 | 99 | 99 | 0 | 0 | 0 | 0 | 2 |
[41] † Rhachiocephalus magnus Taxon name last Modified on 08/24/23 | 19 | 156 | 155 | 0 | 0 | 0 | 0 | 2 |
[42] † Kitchinganomodon crassus Taxon name last Modified on 08/24/23 | 40 | 134 | 134 | 0 | 0 | 0 | 0 | 2 |
[43] † Oudenodon bainii Taxon name last Modified on 08/24/23 | 7 | 168 | 167 | 0 | 0 | 0 | 0 | 2 |
[44] † Tropidostoma dubium Taxon name last Modified on 08/24/23 | 18 | 156 | 156 | 0 | 0 | 0 | 0 | 2 |
[45] † Australobarbarus Taxon name last Modified on 10/16/22 | 42 | 132 | 132 | 0 | 0 | 0 | 0 | 2 |
[46] † Syops vanhoepeni Taxon name last Modified on 08/24/23 | 87 | 87 | 87 | 0 | 0 | 0 | 0 | 2 |
[47] † Odontocyclops whaitsi Taxon name last Modified on 08/24/23 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[48] † Idelesaurus tataricus Taxon name last Modified on 08/24/23 | 42 | 132 | 132 | 0 | 0 | 0 | 0 | 2 |
[49] † Aulacephalodon bainii Taxon name last Modified on 10/15/22 | 7 | 172 | 167 | 0 | 0 | 0 | 0 | 2 |
[50] † Pelanomodon moschops Taxon name last Modified on 08/24/23 | 64 | 111 | 110 | 0 | 0 | 0 | 0 | 2 |
[51] † Geikia locusticeps Taxon name last Modified on 10/15/22 | 36 | 138 | 138 | 0 | 0 | 0 | 0 | 2 |
[52] † Geikia elginensis Taxon name last Modified on 08/24/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 2 |
[53] † Elph borealis Taxon name last Modified on 10/15/22 | 60 | 114 | 114 | 0 | 0 | 0 | 0 | 2 |
[54] † Interpresosaurus blomi Taxon name last Modified on 08/24/23 | 130 | 44 | 44 | 0 | 0 | 0 | 0 | 2 |
[55] † Katumbia parringtoni Taxon name last Modified on 08/24/23 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[56] † Delectosaurus arefjevi Taxon name last Modified on 10/15/22 | 67 | 108 | 107 | 0 | 0 | 0 | 0 | 2 |
[57] † Dicynodon lacerticeps Taxon name last Modified on 10/15/22 | 37 | 138 | 137 | 0 | 0 | 0 | 0 | 2 |
[58] † Daptocephalus leoniceps Taxon name last Modified on 10/15/22 | 27 | 147 | 147 | 0 | 0 | 0 | 0 | 2 |
[59] † Dinanomodon gilli Taxon name last Modified on 10/15/22 | 51 | 123 | 123 | 0 | 0 | 0 | 0 | 2 |
[60] † Peramodon amalitzkii Taxon name last Modified on 08/24/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 2 |
[61] † Vivaxosaurus trautscholdi Taxon name last Modified on 08/24/23 | 33 | 143 | 141 | 0 | 0 | 0 | 0 | 2 |
[62] † Jimusaria sinkiangensis Taxon name last Modified on 08/24/23 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[63] † Sintocephalus alticeps Taxon name last Modified on 08/24/23 | 94 | 80 | 80 | 0 | 0 | 0 | 0 | 2 |
[64] † Basilodon woodwardi Taxon name last Modified on 10/15/22 | 63 | 111 | 111 | 0 | 0 | 0 | 0 | 2 |
[65] † Turfanodon bogdaensis Taxon name last Modified on 08/24/23 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 2 |
[66] † Gordonia traquairi Taxon name last Modified on 08/24/23 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 2 |
[67] † Euptychognathus bathyrhynchus Taxon name last Modified on 10/15/22 | 61 | 113 | 113 | 0 | 0 | 0 | 0 | 2 |
[68] † Lystrosaurus hedini Taxon name last Modified on 08/24/23 | 19 | 155 | 155 | 0 | 0 | 0 | 0 | 2 |
[69] † Lystrosaurus maccaigi Taxon name last Modified on 08/24/23 | 35 | 139 | 139 | 0 | 0 | 0 | 0 | 2 |
[70] † Lystrosaurus curvatus Taxon name last Modified on 08/24/23 | 34 | 140 | 140 | 0 | 0 | 0 | 0 | 2 |
[71] † Lystrosaurus declivis Taxon name last Modified on 08/24/23 | 14 | 160 | 160 | 0 | 0 | 0 | 0 | 2 |
[72] † Lystrosaurus murrayi Taxon name last Modified on 08/24/23 | 14 | 160 | 160 | 0 | 0 | 0 | 0 | 2 |
[73] † Shansiodon Taxon name last Modified on 08/24/23 | 33 | 141 | 141 | 0 | 0 | 0 | 0 | 2 |
[74] † Tetragonias njalilus Taxon name last Modified on 08/24/23 | 10 | 165 | 164 | 0 | 0 | 0 | 0 | 2 |
[75] † Vinceria andina Taxon name last Modified on 08/24/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 2 |
[76] † Rhinodicynodon gracile Taxon name last Modified on 08/24/23 | 41 | 133 | 133 | 0 | 0 | 0 | 0 | 2 |
[77] † Dinodontosaurus Taxon name last Modified on 10/16/22 | 17 | 157 | 157 | 0 | 0 | 0 | 0 | 2 |
[78] † Angonisaurus cruickshanki Taxon name last Modified on 10/15/22 | 45 | 129 | 129 | 0 | 0 | 0 | 0 | 2 |
[79] † Kannemeyeria simocephalus Taxon name last Modified on 08/24/23 | 12 | 162 | 162 | 0 | 0 | 0 | 0 | 2 |
[80] † Kannemeyeria lophorhinus Taxon name last Modified on 08/24/23 | 46 | 128 | 128 | 0 | 0 | 0 | 0 | 2 |
[81] † Dolichuranus primaevus Taxon name last Modified on 10/15/22 | 32 | 142 | 142 | 0 | 0 | 0 | 0 | 2 |
[82] † Sinokannemeyeria Taxon name last Modified on 08/24/23 | 26 | 148 | 148 | 0 | 0 | 0 | 0 | 2 |
[83] † Parakannemeyeria Taxon name last Modified on 08/24/23 | 24 | 151 | 150 | 0 | 0 | 0 | 0 | 2 |
[84] † Xiyukannemeyeria brevirostris Taxon name last Modified on 08/24/23 | 46 | 128 | 128 | 0 | 0 | 0 | 0 | 2 |
[85] † Rabidosaurus cristatus Taxon name last Modified on 08/24/23 | 102 | 72 | 72 | 0 | 0 | 0 | 0 | 2 |
[86] † Rhadiodromus Taxon name last Modified on 08/24/23 | 81 | 94 | 93 | 0 | 0 | 0 | 0 | 2 |
[87] † Wadiasaurus indicus Taxon name last Modified on 08/24/23 | 64 | 111 | 110 | 0 | 0 | 0 | 0 | 2 |
[88] † Shaanbeikannemeyeria Taxon name last Modified on 08/24/23 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 2 |
[89] † Rechnisaurus cristarhynchus Taxon name last Modified on 08/24/23 | 100 | 74 | 74 | 0 | 0 | 0 | 0 | 2 |
[90] † Uralokannemeyeria vjuschkovi Taxon name last Modified on 08/24/23 | 96 | 78 | 78 | 0 | 0 | 0 | 0 | 2 |
[91] † Zambiasaurus submersus Taxon name last Modified on 08/24/23 | 111 | 63 | 63 | 0 | 0 | 0 | 0 | 2 |
[92] † Placerias hesternus Taxon name last Modified on 08/24/23 | 51 | 123 | 123 | 0 | 0 | 0 | 0 | 2 |
[93] † Moghreberia nmachouensis Taxon name last Modified on 08/24/23 | 76 | 98 | 98 | 0 | 0 | 0 | 0 | 2 |
[94] † Stahleckeria potens Taxon name last Modified on 08/24/23 | 11 | 163 | 163 | 0 | 0 | 0 | 0 | 2 |
[95] † Sangusaurus parringtonii Taxon name last Modified on 08/24/23 | 40 | 134 | 134 | 0 | 0 | 0 | 0 | 2 |
[96] † Eubrachiosaurus browni Taxon name last Modified on 10/15/22 | 163 | 11 | 11 | 0 | 0 | 0 | 0 | 2 |
[97] † Jachaleria Taxon name last Modified on 08/24/23 | 44 | 130 | 130 | 0 | 0 | 0 | 0 | 2 |
[98] † Ischigualastia jenseni Taxon name last Modified on 08/24/23 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 2 |
[99] † Rastodon procurvidens Taxon name last Modified on 08/24/23 | 72 | 102 | 102 | 0 | 0 | 0 | 0 | 2 |
[100] † Sauroscaptor tharavati Taxon name last Modified on 08/24/23 | 80 | 94 | 94 | 0 | 0 | 0 | 0 | 2 |
[101] † Bulbasaurus phylloxyron Taxon name last Modified on 10/15/22 | 52 | 126 | 122 | 0 | 0 | 0 | 0 | 2 |
[102] † Compsodon helmoedi Taxon name last Modified on 10/15/22 | 64 | 110 | 110 | 0 | 0 | 0 | 0 | 2 |
[103] † Pentasaurus goggai Taxon name last Modified on 08/24/23 | 150 | 24 | 24 | 0 | 0 | 0 | 0 | 2 |
[104] † Thliptosaurus imperforatus Taxon name last Modified on 08/24/23 | 106 | 68 | 68 | 0 | 0 | 0 | 0 | 2 |
[105] † Ufudocyclops mukanelai Taxon name last Modified on 08/24/23 | 63 | 111 | 111 | 0 | 0 | 0 | 0 | 2 |
[106] † Dicynodon angielczyki Taxon name last Modified on 10/15/22 | 39 | 135 | 135 | 0 | 0 | 0 | 0 | 2 |
[107] † Daptocephalus huenei Taxon name last Modified on 10/15/22 | 65 | 109 | 109 | 0 | 0 | 0 | 0 | 2 |
[108] † Kunpania scopulusa Taxon name last Modified on 08/24/23 | 93 | 81 | 81 | 0 | 0 | 0 | 0 | 2 |
[109] † Taoheodon baizhijuni Taxon name last Modified on 08/24/23 | 67 | 107 | 107 | 0 | 0 | 0 | 0 | 2 |
[110] LPB 1993 3" Taxon name last Modified on 10/15/22 | 96 | 78 | 78 | 0 | 0 | 0 | 0 | 2 |
[111] LPB 1993 2" Taxon name last Modified on 10/15/22 | 99 | 75 | 75 | 0 | 0 | 0 | 0 | 2 |
[112] LPB 1995 9" Taxon name last Modified on 10/15/22 | 91 | 83 | 83 | 0 | 0 | 0 | 0 | 2 |
[113] † Kembawacela kitchingi Taxon name last Modified on 08/24/23 | 38 | 136 | 136 | 0 | 0 | 0 | 0 | 2 |
[114] † Abajudon kaayai Taxon name last Modified on 10/15/22 | 51 | 123 | 123 | 0 | 0 | 0 | 0 | 2 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 1 | |
Matrix downloads | 1 | Angielczyk et al. (2021) (1 download); |