Morphobank

Project 4446: A. Cau, V. Beyrand, D. F. Voeten, V. Fernandez, P. Tafforeau, K. Stein, R. Barsbold, K. Tsogtbaatar, P. J. Currie, P. Godefroit. 2017. Synchrotron scanning reveals amphibious ecomorphology in a new clade of bird-like dinosaurs. Nature. 552 (7685):395-399.

Specimen: † Halszkaraptor (unvouchered)
View: Lateral view of skeleton

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Abstract

Maniraptora includes birds and their closest relatives among theropod dinosaurs1,2,3,4,5. During the Cretaceous period, several maniraptoran lineages diverged from the ancestral coelurosaurian bauplan and evolved novel ecomorphologies, including active flight2, gigantism3, cursoriality4 and herbivory5. Propagation X-ray phase-contrast synchrotron microtomography of a well-preserved maniraptoran from Mongolia, still partially embedded in the rock matrix, revealed a mosaic of features, most of them absent among non-avian maniraptorans but shared by reptilian and avian groups with aquatic or semiaquatic ecologies6,7,8,9,10,11,12,13,14. This new theropod, Halszkaraptor escuilliei gen. et sp. nov., is related to other enigmatic Late Cretaceous maniraptorans from Mongolia15,16 in a novel clade at the root of Dromaeosauridae17. This lineage adds an amphibious ecomorphology to those evolved by maniraptorans: it acquired a predatory mode that relied mainly on neck hyperelongation for food procurement, it coupled the obligatory bipedalism of theropods with forelimb proportions that may support a swimming function, and it developed postural adaptations convergent with short-tailed birds.

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Article DOI: 10.1038/nature24679

Project DOI: 10.7934/P4446, http://dx.doi.org/10.7934/P4446

This project contains	Matrices
1 Media 2 Matrices 5 Documents 311 Taxa 1 Specimen 2620 Characters Total size of project's media files: 22.31k Download Project SDD File	Total scored cells: 115450 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 2633 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 5481 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:2/2631

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MorphoBank Project 4446

Creation Date:
15 October 2022
Publication Date:
19 December 2022
Matrix downloads: 4

Authors' Institutions

Palacky University

University of Alberta

Royal Belgian Institute of Natural Sciences

Museo Geologico ‘Giovanni Capellini’ (Bologna)

European Synchrotron Radiation Facility

Mongolian Academy of Sciences

Vrije Universiteit Brussel

Members

Bidane Martinez Huerta
Project Administrator

311

2633

115450
(115450, 0, 0)

Brooke Long-Fox
Full membership

0
(0, 0, 0)

Taxonomic Overview for Matrix 'M29073' (199 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Euparkeria Taxon name last Modified on 12/12/22	521	1252	1252	0	0	0	0	2
[2] † Achillesaurus Taxon name last Modified on 12/12/22	1680	93	93	0	0	0	0	2
[3] † Achillobator Taxon name last Modified on 12/12/22	1377	396	396	0	0	0	0	2
[4] † Adasaurus Taxon name last Modified on 12/12/22	1385	388	388	0	0	0	0	2
[5] † Albertonykus Taxon name last Modified on 12/12/22	1731	42	42	0	0	0	0	2
[6] † Albinykus Taxon name last Modified on 12/12/22	1681	92	92	0	0	0	0	2
[7] † Allosaurus fragilis Taxon name last Modified on 12/12/22	132	1641	1641	0	0	0	0	2
[8] † Alnashetri Taxon name last Modified on 12/12/22	1665	108	108	0	0	0	0	2
[9] † Alvarezsaurus Taxon name last Modified on 12/12/22	1511	262	262	0	0	0	0	2
[10] † Alxasaurus Taxon name last Modified on 12/12/22	1516	257	257	0	0	0	0	2
[11] † Ambiortus Taxon name last Modified on 12/12/22	1681	92	92	0	0	0	0	2
[12] † Anchiornis type Taxon name last Modified on 12/12/22	1550	223	223	0	0	0	0	2
[13] † Anchiornis lpmb00169 Taxon name last Modified on 12/12/22	1396	377	377	0	0	0	0	2
[14] † Anserimimus Taxon name last Modified on 12/12/22	1631	142	142	0	0	0	0	2
[15] † Anzu Taxon name last Modified on 12/12/22	1404	369	369	0	0	0	0	2
[16] † Apatoraptor Taxon name last Modified on 12/12/22	1600	173	173	0	0	0	0	2
[17] † Apsaravis Taxon name last Modified on 12/12/22	1415	358	358	0	0	0	0	2
[18] † Archaeopteryx Taxon name last Modified on 12/12/22	671	1102	1102	0	0	0	0	2
[19] † Archaeorhynchus Taxon name last Modified on 12/12/22	1386	387	387	0	0	0	0	2
[20] † Archaeornithura Taxon name last Modified on 12/12/22	1594	179	179	0	0	0	0	2
[21] † Aurornis Taxon name last Modified on 12/12/22	1378	395	395	0	0	0	0	2
[22] † Austroraptor Taxon name last Modified on 12/12/22	1469	304	304	0	0	0	0	2
[23] † Avimimus Taxon name last Modified on 12/12/22	1199	574	574	0	0	0	0	2
[24] † Balaur Taxon name last Modified on 12/12/22	1394	379	379	0	0	0	0	2
[25] † Bambiraptor Taxon name last Modified on 12/12/22	884	889	889	0	0	0	0	2
[26] † Banji Taxon name last Modified on 12/12/22	1610	163	163	0	0	0	0	2
[27] † Beipiaosaurus Taxon name last Modified on 12/12/22	1446	327	327	0	0	0	0	2
[28] † Beishanlong Taxon name last Modified on 12/12/22	1604	169	169	0	0	0	0	2
[29] † Bohaiornis Taxon name last Modified on 12/12/22	1439	334	334	0	0	0	0	2
[30] † Boluochia Taxon name last Modified on 12/12/22	1707	66	66	0	0	0	0	2
[31] † Bonapartenykus Taxon name last Modified on 12/12/22	1654	119	119	0	0	0	0	2
[32] † Borogovia Taxon name last Modified on 12/12/22	1685	88	88	0	0	0	0	2
[33] † Buitreraptor Taxon name last Modified on 12/12/22	1281	492	492	0	0	0	0	2
[34] † Byronosaurus Taxon name last Modified on 12/12/22	1488	285	285	0	0	0	0	2
[35] † Caenagnathasia Taxon name last Modified on 12/12/22	1743	30	30	0	0	0	0	2
[36] † Caenagnathus collinsi Taxon name last Modified on 12/12/22	1692	81	81	0	0	0	0	2
[37] † Cathayornis Taxon name last Modified on 12/12/22	1520	253	253	0	0	0	0	2
[38] † Caudipteryx Taxon name last Modified on 12/12/22	1049	724	724	0	0	0	0	2
[39] † Ceratonykus Taxon name last Modified on 12/12/22	1649	124	124	0	0	0	0	2
[40] † Changchengornis Taxon name last Modified on 12/12/22	1617	156	156	0	0	0	0	2
[41] † Changyuraptor Taxon name last Modified on 12/12/22	1646	127	127	0	0	0	0	2
[42] † Chirostenotes Taxon name last Modified on 12/12/22	1435	338	338	0	0	0	0	2
[43] † Citipati Taxon name last Modified on 12/12/22	913	860	860	0	0	0	0	2
[44] † Coelophysis rhodesiensis Taxon name last Modified on 12/12/22	761	1012	1012	0	0	0	0	2
[45] † Coelurus Taxon name last Modified on 12/12/22	1441	332	332	0	0	0	0	2
[46] † Concornis Taxon name last Modified on 12/12/22	1474	299	299	0	0	0	0	2
[47] † Confuciusornis Taxon name last Modified on 12/12/22	940	833	833	0	0	0	0	2
[48] † Cruralispennia Taxon name last Modified on 12/12/22	1554	219	219	0	0	0	0	2
[49] † Deinocheirus Taxon name last Modified on 12/12/22	916	857	857	0	0	0	0	2
[50] † Deinonychus Taxon name last Modified on 12/12/22	642	1131	1131	0	0	0	0	2
[51] † Dromaeosaurus Taxon name last Modified on 12/12/22	1375	398	398	0	0	0	0	2
[52] † Elmisaurus Taxon name last Modified on 12/12/22	1534	239	239	0	0	0	0	2
[53] † Enigmosaurus Taxon name last Modified on 12/12/22	1705	68	68	0	0	0	0	2
[54] † Eocathayornis Taxon name last Modified on 12/12/22	1574	199	199	0	0	0	0	2
[55] † Eoenantiornis Taxon name last Modified on 12/12/22	1494	279	279	0	0	0	0	2
[56] † Eopengornis Taxon name last Modified on 12/12/22	1539	234	234	0	0	0	0	2
[57] † Eosinopteryx Taxon name last Modified on 12/12/22	1452	321	321	0	0	0	0	2
[58] † Epichirostenotes Taxon name last Modified on 12/12/22	1589	184	184	0	0	0	0	2
[59] † Epidendrosaurus Taxon name last Modified on 12/12/22	1357	416	416	0	0	0	0	2
[60] † Epidexipteryx Taxon name last Modified on 12/12/22	1536	237	237	0	0	0	0	2
[61] † Erliansaurus Taxon name last Modified on 12/12/22	1578	195	195	0	0	0	0	2
[62] † Erlicosaurus Taxon name last Modified on 12/12/22	1260	513	513	0	0	0	0	2
[63] † Falcarius Taxon name last Modified on 12/12/22	1001	772	772	0	0	0	0	2
[64] † Feitianius Taxon name last Modified on 12/12/22	1627	146	146	0	0	0	0	2
[65] † Fukuivenator Taxon name last Modified on 12/12/22	1335	438	438	0	0	0	0	2
[66] † Gallimimus Taxon name last Modified on 12/12/22	574	1199	1199	0	0	0	0	2
[67] † Gansus yumenensis Taxon name last Modified on 12/12/22	1419	354	354	0	0	0	0	2
[68] † Ganzhousaurus Taxon name last Modified on 12/12/22	1699	74	74	0	0	0	0	2
[69] † Garudimimus Taxon name last Modified on 12/12/22	1112	661	661	0	0	0	0	2
[70] † Gigantoraptor Taxon name last Modified on 12/12/22	1537	236	236	0	0	0	0	2
[71] † Gobipteryx Taxon name last Modified on 12/12/22	1476	297	297	0	0	0	0	2
[72] † Gobivenator Taxon name last Modified on 12/12/22	1227	546	546	0	0	0	0	2
[73] † Graciliraptor Taxon name last Modified on 12/12/22	1579	194	194	0	0	0	0	2
[74] † Hagryphus Taxon name last Modified on 12/12/22	1687	86	86	0	0	0	0	2
[75] † Halszkaraptor Taxon name last Modified on 12/12/22	1134	639	639	0	0	0	0	2
[76] † Haplocheirus Taxon name last Modified on 12/12/22	1133	640	640	0	0	0	0	2
[77] † Harpymimus Taxon name last Modified on 12/12/22	1124	649	649	0	0	0	0	2
[78] † Hebeiornis Taxon name last Modified on 12/12/22	1468	305	305	0	0	0	0	2
[79] † Herrerasaurus Taxon name last Modified on 12/12/22	512	1261	1261	0	0	0	0	2
[80] † Hesperornis Taxon name last Modified on 12/12/22	1437	336	336	0	0	0	0	2
[81] † Hexing Taxon name last Modified on 12/12/22	1544	229	229	0	0	0	0	2
[82] † Heyuannia huangi Taxon name last Modified on 12/12/22	1370	403	403	0	0	0	0	2
[83] † Heyuannia yanshini Taxon name last Modified on 12/12/22	1274	499	499	0	0	0	0	2
[84] † Hollanda Taxon name last Modified on 12/12/22	1663	110	110	0	0	0	0	2
[85] † Hongshanornis Taxon name last Modified on 12/12/22	1462	311	311	0	0	0	0	2
[86] † Houornis Taxon name last Modified on 12/12/22	1623	150	150	0	0	0	0	2
[87] † Huanansaurus Taxon name last Modified on 12/12/22	1511	262	262	0	0	0	0	2
[88] † Iaceornis Taxon name last Modified on 12/12/22	1645	128	128	0	0	0	0	2
[89] † Iberomesornis Taxon name last Modified on 12/12/22	1549	224	224	0	0	0	0	2
[90] † Ichthyornis Taxon name last Modified on 12/12/22	1320	453	453	0	0	0	0	2
[91] † Incisivosaurus Taxon name last Modified on 12/12/22	1375	398	398	0	0	0	0	2
[92] † Iteravis Taxon name last Modified on 12/12/22	1426	347	347	0	0	0	0	2
[93] † Jianchangornis Taxon name last Modified on 12/12/22	1562	211	211	0	0	0	0	2
[94] † Jianchangosaurus Taxon name last Modified on 12/12/22	1476	297	297	0	0	0	0	2
[95] † Jiangxisaurus Taxon name last Modified on 12/12/22	1689	84	84	0	0	0	0	2
[96] † Jinfengopteryx Taxon name last Modified on 12/12/22	1486	287	287	0	0	0	0	2
[97] † Jixiangornis Taxon name last Modified on 12/12/22	1564	209	209	0	0	0	0	2
[98] † Khaan Taxon name last Modified on 12/12/22	951	822	822	0	0	0	0	2
[99] † Kol Taxon name last Modified on 12/12/22	1687	86	86	0	0	0	0	2
[100] † Linhenykus Taxon name last Modified on 12/12/22	1648	125	125	0	0	0	0	2
[101] † Linheraptor Taxon name last Modified on 12/12/22	1377	396	396	0	0	0	0	2
[102] † Linhevenator Taxon name last Modified on 12/12/22	1613	160	160	0	0	0	0	2
[103] † Longicrusavis Taxon name last Modified on 12/12/22	1509	264	264	0	0	0	0	2
[104] † Longipteryx Taxon name last Modified on 12/12/22	1482	291	291	0	0	0	0	2
[105] † Longirostravis Taxon name last Modified on 12/12/22	1560	213	213	0	0	0	0	2
[106] † Longusunguis Taxon name last Modified on 12/12/22	1475	298	298	0	0	0	0	2
[107] † Machairasaurus Taxon name last Modified on 12/12/22	1679	94	94	0	0	0	0	2
[108] † Mahakala Taxon name last Modified on 12/12/22	1553	220	220	0	0	0	0	2
[109] † Mei Taxon name last Modified on 12/12/22	1193	580	580	0	0	0	0	2
[110] † Meleagris Taxon name last Modified on 12/12/22	875	898	898	0	0	0	0	2
[111] † Microraptor Taxon name last Modified on 12/12/22	972	801	801	0	0	0	0	2
[112] † Microvenator Taxon name last Modified on 12/12/22	1404	369	369	0	0	0	0	2
[113] † Mononykus Taxon name last Modified on 12/12/22	1195	578	578	0	0	0	0	2
[114] † MPC d 100 42 Taxon name last Modified on 12/12/22	1222	551	551	0	0	0	0	2
[115] † MPC d 100 44 Taxon name last Modified on 12/12/22	1617	156	156	0	0	0	0	2
[116] † MPC d 100 140 Taxon name last Modified on 12/12/22	1621	152	152	0	0	0	0	2
[117] † Nankangia Taxon name last Modified on 12/12/22	1524	249	249	0	0	0	0	2
[118] † Nashiungosaurus brevispinus Taxon name last Modified on 12/12/22	1702	71	71	0	0	0	0	2
[119] † Neimongosaurus Taxon name last Modified on 12/12/22	1527	246	246	0	0	0	0	2
[120] † Nemegtomaia Taxon name last Modified on 12/12/22	1388	385	385	0	0	0	0	2
[121] † Neuquenornis Taxon name last Modified on 12/12/22	1575	198	198	0	0	0	0	2
[122] † NGMC 91 Taxon name last Modified on 12/12/22	1500	273	273	0	0	0	0	2
[123] † Nomingia Taxon name last Modified on 12/12/22	1505	268	268	0	0	0	0	2
[124] † Nothronychus graffami Taxon name last Modified on 12/12/22	1438	335	335	0	0	0	0	2
[125] † Nothronychus mckinleyi Taxon name last Modified on 12/12/22	1539	234	234	0	0	0	0	2
[126] † Nqwebasaurus Taxon name last Modified on 12/12/22	1374	399	399	0	0	0	0	2
[127] † Ornitholestes Taxon name last Modified on 12/12/22	1011	762	762	0	0	0	0	2
[128] † Ornithomimus Taxon name last Modified on 12/12/22	697	1076	1076	0	0	0	0	2
[129] † Oviraptor Taxon name last Modified on 12/12/22	1521	252	252	0	0	0	0	2
[130] † Parabohaiornis Taxon name last Modified on 12/12/22	1363	410	410	0	0	0	0	2
[131] † Parahongshanornis Taxon name last Modified on 12/12/22	1678	95	95	0	0	0	0	2
[132] † Parvicursor Taxon name last Modified on 12/12/22	1630	143	143	0	0	0	0	2
[133] † Patagonykus Taxon name last Modified on 12/12/22	1448	325	325	0	0	0	0	2
[134] † Patagopteryx Taxon name last Modified on 12/12/22	1446	327	327	0	0	0	0	2
[135] † Pelecanimimus Taxon name last Modified on 12/12/22	1431	342	342	0	0	0	0	2
[136] † Pengornis Taxon name last Modified on 12/12/22	1456	317	317	0	0	0	0	2
[137] † Philovenator Taxon name last Modified on 12/12/22	1671	102	102	0	0	0	0	2
[138] † Piscivoravis Taxon name last Modified on 12/12/22	1538	235	235	0	0	0	0	2
[139] † Protarchaeopteryx Taxon name last Modified on 12/12/22	1478	295	295	0	0	0	0	2
[140] † Protopteryx Taxon name last Modified on 12/12/22	1449	324	324	0	0	0	0	2
[141] † Pterygornis Taxon name last Modified on 12/12/22	1532	241	241	0	0	0	0	2
[142] † Qiliania Taxon name last Modified on 12/12/22	1659	114	114	0	0	0	0	2
[143] † Qiupalong Taxon name last Modified on 12/12/22	1652	121	121	0	0	0	0	2
[144] † Rahonavis Taxon name last Modified on 12/12/22	1342	431	431	0	0	0	0	2
[145] † Rapaxavis Taxon name last Modified on 12/12/22	1469	304	304	0	0	0	0	2
[146] † Rinchenia Taxon name last Modified on 12/12/22	1480	293	293	0	0	0	0	2
[147] † Sapeornis Taxon name last Modified on 12/12/22	1148	625	625	0	0	0	0	2
[148] † Saurornithoides Taxon name last Modified on 12/12/22	1497	276	276	0	0	0	0	2
[149] † Saurornitholestes Taxon name last Modified on 12/12/22	1675	98	98	0	0	0	0	2
[150] † Schizooura Taxon name last Modified on 12/12/22	1462	311	311	0	0	0	0	2
[151] † Segnosaurus Taxon name last Modified on 12/12/22	1449	324	324	0	0	0	0	2
[152] † Shanag Taxon name last Modified on 12/12/22	1679	94	94	0	0	0	0	2
[153] † Shengjingornis Taxon name last Modified on 12/12/22	1612	161	161	0	0	0	0	2
[154] † Shenqiornis Taxon name last Modified on 12/12/22	1519	254	254	0	0	0	0	2
[155] † Shenzhouraptor Taxon name last Modified on 12/12/22	1194	579	579	0	0	0	0	2
[156] † Shenzhousaurus Taxon name last Modified on 12/12/22	1403	370	370	0	0	0	0	2
[157] † Shixinggia Taxon name last Modified on 12/12/22	1676	97	97	0	0	0	0	2
[158] † Shuvuuia Taxon name last Modified on 12/12/22	1001	772	772	0	0	0	0	2
[159] † Similicaudipteryx Taxon name last Modified on 12/12/22	1329	444	444	0	0	0	0	2
[160] † Sinornis Taxon name last Modified on 12/12/22	1390	383	383	0	0	0	0	2
[161] † Sinornithoides Taxon name last Modified on 12/12/22	1329	444	444	0	0	0	0	2
[162] † Sinornithosaurus Taxon name last Modified on 12/12/22	1120	653	653	0	0	0	0	2
[163] † Sinovenator Taxon name last Modified on 12/12/22	1243	530	530	0	0	0	0	2
[164] † Sinraptor dongi Taxon name last Modified on 12/12/22	497	1276	1276	0	0	0	0	2
[165] † Sinusonasus Taxon name last Modified on 12/12/22	1505	268	268	0	0	0	0	2
[166] † Sulcavis Taxon name last Modified on 12/12/22	1473	300	300	0	0	0	0	2
[167] † Suzhousaurus Taxon name last Modified on 12/12/22	1558	215	215	0	0	0	0	2
[168] † Talos Taxon name last Modified on 12/12/22	1667	106	106	0	0	0	0	2
[169] † Tanycolagreus Taxon name last Modified on 12/12/22	1183	590	590	0	0	0	0	2
[170] † Teviornis Taxon name last Modified on 12/12/22	1702	71	71	0	0	0	0	2
[171] † Therizinosaurus Taxon name last Modified on 12/12/22	1564	209	209	0	0	0	0	2
[172] † Tianyuraptor Taxon name last Modified on 12/12/22	1490	283	283	0	0	0	0	2
[173] † Troodon Taxon name last Modified on 12/12/22	1344	429	429	0	0	0	0	2
[174] † Tsaagan Taxon name last Modified on 12/12/22	1289	484	484	0	0	0	0	2
[175] † Tugriken shireh Parvicursorinae Taxon name last Modified on 12/12/22	1581	192	192	0	0	0	0	2
[176] † Unenlagia comahuensis Taxon name last Modified on 12/12/22	1559	214	214	0	0	0	0	2
[177] † Unenlagia paynemili Taxon name last Modified on 12/12/22	1687	86	86	0	0	0	0	2
[178] † Utahraptor Taxon name last Modified on 12/12/22	1615	158	158	0	0	0	0	2
[179] † Vegavis Taxon name last Modified on 12/12/22	1696	77	77	0	0	0	0	2
[180] † Velociraptor mongoliensis Taxon name last Modified on 12/12/22	500	1273	1273	0	0	0	0	2
[181] † Vorona Taxon name last Modified on 12/12/22	1657	116	116	0	0	0	0	2
[182] † Wulatelong Taxon name last Modified on 12/12/22	1569	204	204	0	0	0	0	2
[183] † Xiaotingia Taxon name last Modified on 12/12/22	1459	314	314	0	0	0	0	2
[184] † Xixiasaurus Taxon name last Modified on 12/12/22	1601	172	172	0	0	0	0	2
[185] † Xixianykus Taxon name last Modified on 12/12/22	1615	158	158	0	0	0	0	2
[186] † Yanornis Taxon name last Modified on 12/12/22	1391	382	382	0	0	0	0	2
[187] † Yi Taxon name last Modified on 12/12/22	1640	133	133	0	0	0	0	2
[188] † Yixianornis Taxon name last Modified on 12/12/22	1369	404	404	0	0	0	0	2
[189] † Yixianosaurus Taxon name last Modified on 12/12/22	1611	162	162	0	0	0	0	2
[190] † Zanabazar Taxon name last Modified on 12/12/22	1421	352	352	0	0	0	0	2
[191] † Zhenyuanlong Taxon name last Modified on 12/12/22	1416	357	357	0	0	0	0	2
[192] † Zhongjianornis Taxon name last Modified on 12/12/22	1535	238	238	0	0	0	0	2
[193] † Zhongornis Taxon name last Modified on 12/12/22	1586	187	187	0	0	0	0	2
[194] † Zhouornis Taxon name last Modified on 12/12/22	1363	410	410	0	0	0	0	2
[195] † Zuolong Taxon name last Modified on 12/12/22	1340	433	433	0	0	0	0	2
[196] † Anchiornithin sp Taxon name last Modified on 12/12/22	1428	345	345	0	0	0	0	2
[197] † Hesperonychus Taxon name last Modified on 12/12/22	1722	51	51	0	0	0	0	2
[198] † Hulsanpes Taxon name last Modified on 12/12/22	1730	43	43	0	0	0	0	2
[199] † Neuquenraptor Taxon name last Modified on 12/12/22	1696	77	77	0	0	0	0	2

Taxonomic Overview for Matrix 'M29101' (153 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Allosaurus fragilis Taxon name last Modified on 12/12/22	67	796	793	0	0	0	0	2
[2] † Sinraptor dongi Taxon name last Modified on 12/12/22	207	653	653	0	0	0	0	2
[3] † Dromaeosaurus albertensis Taxon name last Modified on 12/12/22	610	250	250	0	0	0	0	2
[4] † Deinonychus antirrhopus Taxon name last Modified on 12/12/22	253	608	607	0	0	0	0	2
[5] † Velociraptor mongoliensis Taxon name last Modified on 12/12/22	112	750	748	0	0	0	0	2
[6] † Balaur bondoc Taxon name last Modified on 12/12/22	596	266	264	0	0	0	0	2
[7] † Tsaagan mangas Taxon name last Modified on 12/12/22	502	358	358	0	0	0	0	2
[8] † Bambiraptor feinbergorum Taxon name last Modified on 12/12/22	230	632	630	0	0	0	0	2
[9] † Tianyuraptor ostromi Taxon name last Modified on 12/12/22	680	180	180	0	0	0	0	2
[10] † Sinornithosaurus millenii Taxon name last Modified on 12/12/22	433	428	427	0	0	0	0	2
[11] † Microraptor zhaoianus Taxon name last Modified on 12/12/22	505	356	355	0	0	0	0	2
[12] † Graciliraptor lujiatunensis Taxon name last Modified on 12/12/22	760	101	100	0	0	0	0	2
[13] † Hesperonychus elizabethae Taxon name last Modified on 12/12/22	826	34	34	0	0	0	0	2
[14] † Pyroraptor olympius Taxon name last Modified on 12/12/22	838	22	22	0	0	0	0	2
[15] † Rahonavis ostromi Taxon name last Modified on 12/12/22	633	228	227	0	0	0	0	2
[16] † Buitreraptor gonzalozorum Taxon name last Modified on 12/12/22	455	409	405	0	0	0	0	2
[17] † Neuquenraptor plus Unenlagia Taxon name last Modified on 12/12/22	624	236	236	0	0	0	0	2
[18] † Austroraptor Taxon name last Modified on 12/12/22	683	179	177	0	0	0	0	2
[19] † Shanag ashile Taxon name last Modified on 12/12/22	797	63	63	0	0	0	0	2
[20] † Mahakala omnogovae Taxon name last Modified on 12/12/22	572	291	288	0	0	0	0	2
[21] † Atrociraptor marshalli Taxon name last Modified on 12/12/22	792	69	68	0	0	0	0	2
[22] † Utahraptor Taxon name last Modified on 12/12/22	728	133	132	0	0	0	0	2
[23] † Adasaurus mongoliensis Taxon name last Modified on 12/12/22	569	293	291	0	0	0	0	2
[24] † Achillobator giganticus Taxon name last Modified on 12/12/22	660	200	200	0	0	0	0	2
[25] † Saurornitholestes langsto Taxon name last Modified on 12/12/22	687	173	173	0	0	0	0	2
[26] † Saurornithoides mongolien Taxon name last Modified on 12/12/22	608	253	252	0	0	0	0	2
[27] † Zanabazar junior Taxon name last Modified on 12/12/22	601	260	259	0	0	0	0	2
[28] † Xixiasaurus Taxon name last Modified on 12/12/22	729	131	131	0	0	0	0	2
[29] † Byronosaurus jaffei Taxon name last Modified on 12/12/22	670	190	190	0	0	0	0	2
[30] † Sinornithoides youngi Taxon name last Modified on 12/12/22	560	301	300	0	0	0	0	2
[31] † Mei long Taxon name last Modified on 12/12/22	440	422	420	0	0	0	0	2
[32] † Jinfengopteryx elegans Taxon name last Modified on 12/12/22	700	160	160	0	0	0	0	2
[33] † Anchiornis huxleyi Taxon name last Modified on 12/12/22	485	376	375	0	0	0	0	2
[34] † Xiaotingia Taxon name last Modified on 12/12/22	578	283	282	0	0	0	0	2
[35] † Aurornis Taxon name last Modified on 12/12/22	595	265	265	0	0	0	0	2
[36] † Eosinopteryx Taxon name last Modified on 12/12/22	615	245	245	0	0	0	0	2
[37] † Troodon formosus Taxon name last Modified on 12/12/22	597	263	263	0	0	0	0	2
[38] † Sinovenator changii Taxon name last Modified on 12/12/22	448	414	412	0	0	0	0	2
[39] † EK troodontid IGM 100 slash 44 Taxon name last Modified on 12/12/22	813	47	47	0	0	0	0	2
[40] † IGM 100 slash 1126 Taxon name last Modified on 12/12/22	615	246	245	0	0	0	0	2
[41] † IGM 100 slash 1323 Taxon name last Modified on 12/12/22	712	149	148	0	0	0	0	2
[42] † Archaeopteryx lithographica Taxon name last Modified on 12/12/22	247	615	613	0	0	0	0	2
[43] † Confuciusornis sanctus Taxon name last Modified on 12/12/22	379	485	481	0	0	0	0	2
[44] † Jeholornis prima Taxon name last Modified on 12/12/22	484	376	376	0	0	0	0	2
[45] † Jixiangornis orientalis Taxon name last Modified on 12/12/22	518	343	342	0	0	0	0	2
[46] † Yanornis martini Taxon name last Modified on 12/12/22	568	298	292	0	0	0	0	2
[47] † Apsaravis ukhaana Taxon name last Modified on 12/12/22	560	305	300	0	0	0	0	2
[48] † Yixianornis Taxon name last Modified on 12/12/22	503	362	357	0	0	0	0	2
[49] † Sapeornis Taxon name last Modified on 12/12/22	427	436	433	0	0	0	0	2
[50] † Neuquenornis volans Taxon name last Modified on 12/12/22	715	148	145	0	0	0	0	2
[51] † Patagopteryx deferrariisi Taxon name last Modified on 12/12/22	599	267	261	0	0	0	0	2
[52] † Cathayornis Taxon name last Modified on 12/12/22	575	292	285	0	0	0	0	2
[53] † Concornis Taxon name last Modified on 12/12/22	698	169	162	0	0	0	0	2
[54] † Gobipteryx Taxon name last Modified on 12/12/22	726	137	134	0	0	0	0	2
[55] † Vorona Taxon name last Modified on 12/12/22	781	79	79	0	0	0	0	2
[56] † Songlingornis Taxon name last Modified on 12/12/22	796	66	64	0	0	0	0	2
[57] † Pengornis houi Taxon name last Modified on 12/12/22	583	284	277	0	0	0	0	2
[58] † Hesperornis Taxon name last Modified on 12/12/22	501	359	359	0	0	0	0	2
[59] † Baptornis Taxon name last Modified on 12/12/22	733	130	127	0	0	0	0	2
[60] † Ichthyornis Taxon name last Modified on 12/12/22	430	436	430	0	0	0	0	2
[61] † Iaceornis marshii Taxon name last Modified on 12/12/22	738	123	122	0	0	0	0	2
[62] † Limenavis patagonica Taxon name last Modified on 12/12/22	820	42	40	0	0	0	0	2
[63] † Lithornis Taxon name last Modified on 12/12/22	537	331	323	0	0	0	0	2
[64] † Hongshanornis longicrest Taxon name last Modified on 12/12/22	621	243	239	0	0	0	0	2
[65] † Liaoningornis longidigitu Taxon name last Modified on 12/12/22	791	69	69	0	0	0	0	2
[66] † Crypturellus undulatus Taxon name last Modified on 12/12/22	446	415	414	0	0	0	0	2
[67] † Gallus gallus Taxon name last Modified on 12/12/22	444	418	416	0	0	0	0	2
[68] † Crax pauxi Taxon name last Modified on 12/12/22	439	421	421	0	0	0	0	2
[69] † Anas platyrhynchus Taxon name last Modified on 12/12/22	458	404	402	0	0	0	0	2
[70] † Chauna torquata Taxon name last Modified on 12/12/22	439	422	421	0	0	0	0	2
[71] † Pedopenna Taxon name last Modified on 12/12/22	828	32	32	0	0	0	0	2
[72] † Epidendrosaurus Taxon name last Modified on 12/12/22	766	94	94	0	0	0	0	2
[73] † Epidexipteryx Taxon name last Modified on 12/12/22	687	175	173	0	0	0	0	2
[74] † Incisivosaurus gauthieri Taxon name last Modified on 12/12/22	575	285	285	0	0	0	0	2
[75] † Citipati osmolskae Taxon name last Modified on 12/12/22	296	567	564	0	0	0	0	2
[76] † Oviraptor philoceratops Taxon name last Modified on 12/12/22	643	217	217	0	0	0	0	2
[77] † Microvenator celer Taxon name last Modified on 12/12/22	615	245	245	0	0	0	0	2
[78] † Caudipteryx zoui Taxon name last Modified on 12/12/22	526	335	334	0	0	0	0	2
[79] † Ingenia yanshani Taxon name last Modified on 12/12/22	534	327	326	0	0	0	0	2
[80] † Rinchenia mongoliensis Taxon name last Modified on 12/12/22	689	171	171	0	0	0	0	2
[81] † Conchoraptor gracilis Taxon name last Modified on 12/12/22	532	329	328	0	0	0	0	2
[82] † Chirostenotes pergracilis Taxon name last Modified on 12/12/22	559	303	301	0	0	0	0	2
[83] † Avimimus portentosus Taxon name last Modified on 12/12/22	531	331	329	0	0	0	0	2
[84] † Falcarius Taxon name last Modified on 12/12/22	307	554	553	0	0	0	0	2
[85] † Beipiaosaurus Taxon name last Modified on 12/12/22	656	204	204	0	0	0	0	2
[86] † Segnosaurus galbinensis Taxon name last Modified on 12/12/22	646	214	214	0	0	0	0	2
[87] † Erlikosaurus andrewsi Taxon name last Modified on 12/12/22	522	338	338	0	0	0	0	2
[88] † Alxasaurus elesitaiensis Taxon name last Modified on 12/12/22	653	208	207	0	0	0	0	2
[89] † Neimongosaurus Taxon name last Modified on 12/12/22	629	232	231	0	0	0	0	2
[90] † Erliansaurus Taxon name last Modified on 12/12/22	725	135	135	0	0	0	0	2
[91] † Suzhousaurus Taxon name last Modified on 12/12/22	652	209	208	0	0	0	0	2
[92] † Nothronychus graffami Taxon name last Modified on 12/12/22	503	359	357	0	0	0	0	2
[93] † Enigmosaurus Taxon name last Modified on 12/12/22	795	65	65	0	0	0	0	2
[94] † Nanshiungosaurus brevispinus Taxon name last Modified on 12/12/22	821	39	39	0	0	0	0	2
[95] † Therizinosaurus Taxon name last Modified on 12/12/22	738	122	122	0	0	0	0	2
[96] † Haplocheirus Taxon name last Modified on 12/12/22	344	516	516	0	0	0	0	2
[97] † Alvarezsaurus calvoi Taxon name last Modified on 12/12/22	706	154	154	0	0	0	0	2
[98] † Patagonykus puertai Taxon name last Modified on 12/12/22	652	208	208	0	0	0	0	2
[99] † Achillesaurus Taxon name last Modified on 12/12/22	801	59	59	0	0	0	0	2
[100] † Mononykus olecranus Taxon name last Modified on 12/12/22	533	327	327	0	0	0	0	2
[101] † Shuvuuia deserti Taxon name last Modified on 12/12/22	250	612	610	0	0	0	0	2
[102] † Parvicursor Taxon name last Modified on 12/12/22	726	134	134	0	0	0	0	2
[103] † Albertonykus Taxon name last Modified on 12/12/22	835	25	25	0	0	0	0	2
[104] † Albinykus Taxon name last Modified on 12/12/22	791	69	69	0	0	0	0	2
[105] † Bonapartenykus Taxon name last Modified on 12/12/22	784	76	76	0	0	0	0	2
[106] † Ceratonykus Taxon name last Modified on 12/12/22	717	143	143	0	0	0	0	2
[107] † Linhenykus Taxon name last Modified on 12/12/22	696	164	164	0	0	0	0	2
[108] † Xixianykus Taxon name last Modified on 12/12/22	696	164	164	0	0	0	0	2
[109] † Nqwebasaurus Taxon name last Modified on 12/12/22	657	203	203	0	0	0	0	2
[110] † Shenzhousaurus orientalis Taxon name last Modified on 12/12/22	599	261	261	0	0	0	0	2
[111] † Ornithomimus edmonticus Taxon name last Modified on 12/12/22	337	524	523	0	0	0	0	2
[112] † Archaeornithomimus asiati Taxon name last Modified on 12/12/22	514	348	346	0	0	0	0	2
[113] † Anserimimus planinychus Taxon name last Modified on 12/12/22	731	129	129	0	0	0	0	2
[114] † Struthiomimus altus Taxon name last Modified on 12/12/22	234	627	626	0	0	0	0	2
[115] † Gallimimus bullatus Taxon name last Modified on 12/12/22	149	713	711	0	0	0	0	2
[116] † Garudimimus brevipes Taxon name last Modified on 12/12/22	347	513	513	0	0	0	0	2
[117] † Pelecanimimus polydon Taxon name last Modified on 12/12/22	610	250	250	0	0	0	0	2
[118] † Harpymimus okladnikovi Taxon name last Modified on 12/12/22	476	385	384	0	0	0	0	2
[119] † Beishanlong Taxon name last Modified on 12/12/22	662	198	198	0	0	0	0	2
[120] † Sinornithomimus Taxon name last Modified on 12/12/22	294	566	566	0	0	0	0	2
[121] † Qiupalong Taxon name last Modified on 12/12/22	749	111	111	0	0	0	0	2
[122] † Kinnareemimus Taxon name last Modified on 12/12/22	803	57	57	0	0	0	0	2
[123] † Huaxiagnathus orientalis Taxon name last Modified on 12/12/22	523	338	337	0	0	0	0	2
[124] † Sinosauropteryx prima Taxon name last Modified on 12/12/22	494	367	366	0	0	0	0	2
[125] † Compsognathus longipes Taxon name last Modified on 12/12/22	414	447	446	0	0	0	0	2
[126] † Juravenator starki Taxon name last Modified on 12/12/22	484	377	376	0	0	0	0	2
[127] † Sinocalliopteryx Taxon name last Modified on 12/12/22	454	407	406	0	0	0	0	2
[128] † Mirischia Taxon name last Modified on 12/12/22	783	77	77	0	0	0	0	2
[129] † Ornitholestes hermanni Taxon name last Modified on 12/12/22	336	525	524	0	0	0	0	2
[130] † Coelurus fragilis Taxon name last Modified on 12/12/22	631	229	229	0	0	0	0	2
[131] † Tanycolagreus Taxon name last Modified on 12/12/22	495	366	365	0	0	0	0	2
[132] † Tugulusaurus Taxon name last Modified on 12/12/22	792	68	68	0	0	0	0	2
[133] † Zuolong Taxon name last Modified on 12/12/22	489	371	371	0	0	0	0	2
[134] † Bicentenaria Taxon name last Modified on 12/12/22	679	181	181	0	0	0	0	2
[135] † Kileskus Taxon name last Modified on 12/12/22	788	72	72	0	0	0	0	2
[136] † Guanlong Taxon name last Modified on 12/12/22	167	694	693	0	0	0	0	2
[137] † Sinotyrannus Taxon name last Modified on 12/12/22	776	84	84	0	0	0	0	2
[138] † Proceratosaurus bradleyi Taxon name last Modified on 12/12/22	654	206	206	0	0	0	0	2
[139] † Dilong paradoxus Taxon name last Modified on 12/12/22	279	581	581	0	0	0	0	2
[140] † Eotyrannus lengi Taxon name last Modified on 12/12/22	584	276	276	0	0	0	0	2
[141] † Juratyrant Taxon name last Modified on 12/12/22	702	158	158	0	0	0	0	2
[142] † Xiongguanlong Taxon name last Modified on 12/12/22	545	315	315	0	0	0	0	2
[143] † Dryptosaurus Taxon name last Modified on 12/12/22	713	148	147	0	0	0	0	2
[144] † Appalachiosaurus Taxon name last Modified on 12/12/22	612	248	248	0	0	0	0	2
[145] † Bistahieversor Taxon name last Modified on 12/12/22	550	310	310	0	0	0	0	2
[146] † Albertosaurus sacrophagus Taxon name last Modified on 12/12/22	160	707	700	0	0	0	0	2
[147] † Gorgosaurus libratus Taxon name last Modified on 12/12/22	76	787	784	0	0	0	0	2
[148] † Alioramus Taxon name last Modified on 12/12/22	305	556	555	0	0	0	0	2
[149] † Teratophoneus Taxon name last Modified on 12/12/22	715	145	145	0	0	0	0	2
[150] † Daspletosaurus Taxon name last Modified on 12/12/22	180	683	680	0	0	0	0	2
[151] † Tyrannosaurus rex Taxon name last Modified on 12/12/22	52	814	808	0	0	0	0	2
[152] † Tarbosaurus baatar Taxon name last Modified on 12/12/22	83	777	777	0	0	0	0	2
[153] † Halszkaraptor escuilliei Taxon name last Modified on 12/12/22	557	306	303	0	0	0	0	2

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	8
Project downloads	4
Matrix downloads	4	Cau et al (2017) (1) (3 downloads); Cau et al (2017) (2) (1 download);