Morphobank

Project 4446: A. Cau, V. Beyrand, D. F. Voeten, V. Fernandez, P. Tafforeau, K. Stein, R. Barsbold, K. Tsogtbaatar, P. J. Currie, P. Godefroit. 2017. Synchrotron scanning reveals amphibious ecomorphology in a new clade of bird-like dinosaurs. Nature. 552 (7685):395-399.

Specimen: † Halszkaraptor (unvouchered)
View: Lateral view of skeleton

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Abstract

Maniraptora includes birds and their closest relatives among theropod dinosaurs1,2,3,4,5. During the Cretaceous period, several maniraptoran lineages diverged from the ancestral coelurosaurian bauplan and evolved novel ecomorphologies, including active flight2, gigantism3, cursoriality4 and herbivory5. Propagation X-ray phase-contrast synchrotron microtomography of a well-preserved maniraptoran from Mongolia, still partially embedded in the rock matrix, revealed a mosaic of features, most of them absent among non-avian maniraptorans but shared by reptilian and avian groups with aquatic or semiaquatic ecologies6,7,8,9,10,11,12,13,14. This new theropod, Halszkaraptor escuilliei gen. et sp. nov., is related to other enigmatic Late Cretaceous maniraptorans from Mongolia15,16 in a novel clade at the root of Dromaeosauridae17. This lineage adds an amphibious ecomorphology to those evolved by maniraptorans: it acquired a predatory mode that relied mainly on neck hyperelongation for food procurement, it coupled the obligatory bipedalism of theropods with forelimb proportions that may support a swimming function, and it developed postural adaptations convergent with short-tailed birds.

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Article DOI: 10.1038/nature24679

Project DOI: 10.7934/P4446, http://dx.doi.org/10.7934/P4446

This project contains	Matrices
1 Media 2 Matrices 5 Documents 311 Taxa 1 Specimen 2620 Characters Total size of project's media files: 22.31k Download Project SDD File	Total scored cells: 115450 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 2633 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 5481 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:2/2631

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MorphoBank Project 4446

Creation Date:
15 October 2022
Publication Date:
19 December 2022
Project views: 5208
Matrix downloads: 3

Authors' Institutions

Palacky University

University of Alberta

Royal Belgian Institute of Natural Sciences

Museo Geologico ‘Giovanni Capellini’ (Bologna)

European Synchrotron Radiation Facility

Mongolian Academy of Sciences

Vrije Universiteit Brussel

Members

Bidane Martinez Huerta
Project Administrator

311

2633

115450
(115450, 0, 0)

Brooke Long-Fox
Full membership

0
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Taxonomic Overview for Matrix 'M29073' (199 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Euparkeria Last Modified in 12/12/22	521	1252	1252	0	0	0	0	2
[2] † Achillesaurus Last Modified in 12/12/22	1680	93	93	0	0	0	0	2
[3] † Achillobator Last Modified in 12/12/22	1377	396	396	0	0	0	0	2
[4] † Adasaurus Last Modified in 12/12/22	1385	388	388	0	0	0	0	2
[5] † Albertonykus Last Modified in 12/12/22	1731	42	42	0	0	0	0	2
[6] † Albinykus Last Modified in 12/12/22	1681	92	92	0	0	0	0	2
[7] † Allosaurus fragilis Last Modified in 12/12/22	132	1641	1641	0	0	0	0	2
[8] † Alnashetri Last Modified in 12/12/22	1665	108	108	0	0	0	0	2
[9] † Alvarezsaurus Last Modified in 12/12/22	1511	262	262	0	0	0	0	2
[10] † Alxasaurus Last Modified in 12/12/22	1516	257	257	0	0	0	0	2
[11] † Ambiortus Last Modified in 12/12/22	1681	92	92	0	0	0	0	2
[12] † Anchiornis type Last Modified in 12/12/22	1550	223	223	0	0	0	0	2
[13] † Anchiornis lpmb00169 Last Modified in 12/12/22	1396	377	377	0	0	0	0	2
[14] † Anserimimus Last Modified in 12/12/22	1631	142	142	0	0	0	0	2
[15] † Anzu Last Modified in 12/12/22	1404	369	369	0	0	0	0	2
[16] † Apatoraptor Last Modified in 12/12/22	1600	173	173	0	0	0	0	2
[17] † Apsaravis Last Modified in 12/12/22	1415	358	358	0	0	0	0	2
[18] † Archaeopteryx Last Modified in 12/12/22	671	1102	1102	0	0	0	0	2
[19] † Archaeorhynchus Last Modified in 12/12/22	1386	387	387	0	0	0	0	2
[20] † Archaeornithura Last Modified in 12/12/22	1594	179	179	0	0	0	0	2
[21] † Aurornis Last Modified in 12/12/22	1378	395	395	0	0	0	0	2
[22] † Austroraptor Last Modified in 12/12/22	1469	304	304	0	0	0	0	2
[23] † Avimimus Last Modified in 12/12/22	1199	574	574	0	0	0	0	2
[24] † Balaur Last Modified in 12/12/22	1394	379	379	0	0	0	0	2
[25] † Bambiraptor Last Modified in 12/12/22	884	889	889	0	0	0	0	2
[26] † Banji Last Modified in 12/12/22	1610	163	163	0	0	0	0	2
[27] † Beipiaosaurus Last Modified in 12/12/22	1446	327	327	0	0	0	0	2
[28] † Beishanlong Last Modified in 12/12/22	1604	169	169	0	0	0	0	2
[29] † Bohaiornis Last Modified in 12/12/22	1439	334	334	0	0	0	0	2
[30] † Boluochia Last Modified in 12/12/22	1707	66	66	0	0	0	0	2
[31] † Bonapartenykus Last Modified in 12/12/22	1654	119	119	0	0	0	0	2
[32] † Borogovia Last Modified in 12/12/22	1685	88	88	0	0	0	0	2
[33] † Buitreraptor Last Modified in 12/12/22	1281	492	492	0	0	0	0	2
[34] † Byronosaurus Last Modified in 12/12/22	1488	285	285	0	0	0	0	2
[35] † Caenagnathasia Last Modified in 12/12/22	1743	30	30	0	0	0	0	2
[36] † Caenagnathus collinsi Last Modified in 12/12/22	1692	81	81	0	0	0	0	2
[37] † Cathayornis Last Modified in 12/12/22	1520	253	253	0	0	0	0	2
[38] † Caudipteryx Last Modified in 12/12/22	1049	724	724	0	0	0	0	2
[39] † Ceratonykus Last Modified in 12/12/22	1649	124	124	0	0	0	0	2
[40] † Changchengornis Last Modified in 12/12/22	1617	156	156	0	0	0	0	2
[41] † Changyuraptor Last Modified in 12/12/22	1646	127	127	0	0	0	0	2
[42] † Chirostenotes Last Modified in 12/12/22	1435	338	338	0	0	0	0	2
[43] † Citipati Last Modified in 12/12/22	913	860	860	0	0	0	0	2
[44] † Coelophysis rhodesiensis Last Modified in 12/12/22	761	1012	1012	0	0	0	0	2
[45] † Coelurus Last Modified in 12/12/22	1441	332	332	0	0	0	0	2
[46] † Concornis Last Modified in 12/12/22	1474	299	299	0	0	0	0	2
[47] † Confuciusornis Last Modified in 12/12/22	940	833	833	0	0	0	0	2
[48] † Cruralispennia Last Modified in 12/12/22	1554	219	219	0	0	0	0	2
[49] † Deinocheirus Last Modified in 12/12/22	916	857	857	0	0	0	0	2
[50] † Deinonychus Last Modified in 12/12/22	642	1131	1131	0	0	0	0	2
[51] † Dromaeosaurus Last Modified in 12/12/22	1375	398	398	0	0	0	0	2
[52] † Elmisaurus Last Modified in 12/12/22	1534	239	239	0	0	0	0	2
[53] † Enigmosaurus Last Modified in 12/12/22	1705	68	68	0	0	0	0	2
[54] † Eocathayornis Last Modified in 12/12/22	1574	199	199	0	0	0	0	2
[55] † Eoenantiornis Last Modified in 12/12/22	1494	279	279	0	0	0	0	2
[56] † Eopengornis Last Modified in 12/12/22	1539	234	234	0	0	0	0	2
[57] † Eosinopteryx Last Modified in 12/12/22	1452	321	321	0	0	0	0	2
[58] † Epichirostenotes Last Modified in 12/12/22	1589	184	184	0	0	0	0	2
[59] † Epidendrosaurus Last Modified in 12/12/22	1357	416	416	0	0	0	0	2
[60] † Epidexipteryx Last Modified in 12/12/22	1536	237	237	0	0	0	0	2
[61] † Erliansaurus Last Modified in 12/12/22	1578	195	195	0	0	0	0	2
[62] † Erlicosaurus Last Modified in 12/12/22	1260	513	513	0	0	0	0	2
[63] † Falcarius Last Modified in 12/12/22	1001	772	772	0	0	0	0	2
[64] † Feitianius Last Modified in 12/12/22	1627	146	146	0	0	0	0	2
[65] † Fukuivenator Last Modified in 12/12/22	1335	438	438	0	0	0	0	2
[66] † Gallimimus Last Modified in 12/12/22	574	1199	1199	0	0	0	0	2
[67] † Gansus yumenensis Last Modified in 12/12/22	1419	354	354	0	0	0	0	2
[68] † Ganzhousaurus Last Modified in 12/12/22	1699	74	74	0	0	0	0	2
[69] † Garudimimus Last Modified in 12/12/22	1112	661	661	0	0	0	0	2
[70] † Gigantoraptor Last Modified in 12/12/22	1537	236	236	0	0	0	0	2
[71] † Gobipteryx Last Modified in 12/12/22	1476	297	297	0	0	0	0	2
[72] † Gobivenator Last Modified in 12/12/22	1227	546	546	0	0	0	0	2
[73] † Graciliraptor Last Modified in 12/12/22	1579	194	194	0	0	0	0	2
[74] † Hagryphus Last Modified in 12/12/22	1687	86	86	0	0	0	0	2
[75] † Halszkaraptor Last Modified in 12/12/22	1134	639	639	0	0	0	0	2
[76] † Haplocheirus Last Modified in 12/12/22	1133	640	640	0	0	0	0	2
[77] † Harpymimus Last Modified in 12/12/22	1124	649	649	0	0	0	0	2
[78] † Hebeiornis Last Modified in 12/12/22	1468	305	305	0	0	0	0	2
[79] † Herrerasaurus Last Modified in 12/12/22	512	1261	1261	0	0	0	0	2
[80] † Hesperornis Last Modified in 12/12/22	1437	336	336	0	0	0	0	2
[81] † Hexing Last Modified in 12/12/22	1544	229	229	0	0	0	0	2
[82] † Heyuannia huangi Last Modified in 12/12/22	1370	403	403	0	0	0	0	2
[83] † Heyuannia yanshini Last Modified in 12/12/22	1274	499	499	0	0	0	0	2
[84] † Hollanda Last Modified in 12/12/22	1663	110	110	0	0	0	0	2
[85] † Hongshanornis Last Modified in 12/12/22	1462	311	311	0	0	0	0	2
[86] † Houornis Last Modified in 12/12/22	1623	150	150	0	0	0	0	2
[87] † Huanansaurus Last Modified in 12/12/22	1511	262	262	0	0	0	0	2
[88] † Iaceornis Last Modified in 12/12/22	1645	128	128	0	0	0	0	2
[89] † Iberomesornis Last Modified in 12/12/22	1549	224	224	0	0	0	0	2
[90] † Ichthyornis Last Modified in 12/12/22	1320	453	453	0	0	0	0	2
[91] † Incisivosaurus Last Modified in 12/12/22	1375	398	398	0	0	0	0	2
[92] † Iteravis Last Modified in 12/12/22	1426	347	347	0	0	0	0	2
[93] † Jianchangornis Last Modified in 12/12/22	1562	211	211	0	0	0	0	2
[94] † Jianchangosaurus Last Modified in 12/12/22	1476	297	297	0	0	0	0	2
[95] † Jiangxisaurus Last Modified in 12/12/22	1689	84	84	0	0	0	0	2
[96] † Jinfengopteryx Last Modified in 12/12/22	1486	287	287	0	0	0	0	2
[97] † Jixiangornis Last Modified in 12/12/22	1564	209	209	0	0	0	0	2
[98] † Khaan Last Modified in 12/12/22	951	822	822	0	0	0	0	2
[99] † Kol Last Modified in 12/12/22	1687	86	86	0	0	0	0	2
[100] † Linhenykus Last Modified in 12/12/22	1648	125	125	0	0	0	0	2
[101] † Linheraptor Last Modified in 12/12/22	1377	396	396	0	0	0	0	2
[102] † Linhevenator Last Modified in 12/12/22	1613	160	160	0	0	0	0	2
[103] † Longicrusavis Last Modified in 12/12/22	1509	264	264	0	0	0	0	2
[104] † Longipteryx Last Modified in 12/12/22	1482	291	291	0	0	0	0	2
[105] † Longirostravis Last Modified in 12/12/22	1560	213	213	0	0	0	0	2
[106] † Longusunguis Last Modified in 12/12/22	1475	298	298	0	0	0	0	2
[107] † Machairasaurus Last Modified in 12/12/22	1679	94	94	0	0	0	0	2
[108] † Mahakala Last Modified in 12/12/22	1553	220	220	0	0	0	0	2
[109] † Mei Last Modified in 12/12/22	1193	580	580	0	0	0	0	2
[110] † Meleagris Last Modified in 12/12/22	875	898	898	0	0	0	0	2
[111] † Microraptor Last Modified in 12/12/22	972	801	801	0	0	0	0	2
[112] † Microvenator Last Modified in 12/12/22	1404	369	369	0	0	0	0	2
[113] † Mononykus Last Modified in 12/12/22	1195	578	578	0	0	0	0	2
[114] † MPC d 100 42 Last Modified in 12/12/22	1222	551	551	0	0	0	0	2
[115] † MPC d 100 44 Last Modified in 12/12/22	1617	156	156	0	0	0	0	2
[116] † MPC d 100 140 Last Modified in 12/12/22	1621	152	152	0	0	0	0	2
[117] † Nankangia Last Modified in 12/12/22	1524	249	249	0	0	0	0	2
[118] † Nashiungosaurus brevispinus Last Modified in 12/12/22	1702	71	71	0	0	0	0	2
[119] † Neimongosaurus Last Modified in 12/12/22	1527	246	246	0	0	0	0	2
[120] † Nemegtomaia Last Modified in 12/12/22	1388	385	385	0	0	0	0	2
[121] † Neuquenornis Last Modified in 12/12/22	1575	198	198	0	0	0	0	2
[122] † NGMC 91 Last Modified in 12/12/22	1500	273	273	0	0	0	0	2
[123] † Nomingia Last Modified in 12/12/22	1505	268	268	0	0	0	0	2
[124] † Nothronychus graffami Last Modified in 12/12/22	1438	335	335	0	0	0	0	2
[125] † Nothronychus mckinleyi Last Modified in 12/12/22	1539	234	234	0	0	0	0	2
[126] † Nqwebasaurus Last Modified in 12/12/22	1374	399	399	0	0	0	0	2
[127] † Ornitholestes Last Modified in 12/12/22	1011	762	762	0	0	0	0	2
[128] † Ornithomimus Last Modified in 12/12/22	697	1076	1076	0	0	0	0	2
[129] † Oviraptor Last Modified in 12/12/22	1521	252	252	0	0	0	0	2
[130] † Parabohaiornis Last Modified in 12/12/22	1363	410	410	0	0	0	0	2
[131] † Parahongshanornis Last Modified in 12/12/22	1678	95	95	0	0	0	0	2
[132] † Parvicursor Last Modified in 12/12/22	1630	143	143	0	0	0	0	2
[133] † Patagonykus Last Modified in 12/12/22	1448	325	325	0	0	0	0	2
[134] † Patagopteryx Last Modified in 12/12/22	1446	327	327	0	0	0	0	2
[135] † Pelecanimimus Last Modified in 12/12/22	1431	342	342	0	0	0	0	2
[136] † Pengornis Last Modified in 12/12/22	1456	317	317	0	0	0	0	2
[137] † Philovenator Last Modified in 12/12/22	1671	102	102	0	0	0	0	2
[138] † Piscivoravis Last Modified in 12/12/22	1538	235	235	0	0	0	0	2
[139] † Protarchaeopteryx Last Modified in 12/12/22	1478	295	295	0	0	0	0	2
[140] † Protopteryx Last Modified in 12/12/22	1449	324	324	0	0	0	0	2
[141] † Pterygornis Last Modified in 12/12/22	1532	241	241	0	0	0	0	2
[142] † Qiliania Last Modified in 12/12/22	1659	114	114	0	0	0	0	2
[143] † Qiupalong Last Modified in 12/12/22	1652	121	121	0	0	0	0	2
[144] † Rahonavis Last Modified in 12/12/22	1342	431	431	0	0	0	0	2
[145] † Rapaxavis Last Modified in 12/12/22	1469	304	304	0	0	0	0	2
[146] † Rinchenia Last Modified in 12/12/22	1480	293	293	0	0	0	0	2
[147] † Sapeornis Last Modified in 12/12/22	1148	625	625	0	0	0	0	2
[148] † Saurornithoides Last Modified in 12/12/22	1497	276	276	0	0	0	0	2
[149] † Saurornitholestes Last Modified in 12/12/22	1675	98	98	0	0	0	0	2
[150] † Schizooura Last Modified in 12/12/22	1462	311	311	0	0	0	0	2
[151] † Segnosaurus Last Modified in 12/12/22	1449	324	324	0	0	0	0	2
[152] † Shanag Last Modified in 12/12/22	1679	94	94	0	0	0	0	2
[153] † Shengjingornis Last Modified in 12/12/22	1612	161	161	0	0	0	0	2
[154] † Shenqiornis Last Modified in 12/12/22	1519	254	254	0	0	0	0	2
[155] † Shenzhouraptor Last Modified in 12/12/22	1194	579	579	0	0	0	0	2
[156] † Shenzhousaurus Last Modified in 12/12/22	1403	370	370	0	0	0	0	2
[157] † Shixinggia Last Modified in 12/12/22	1676	97	97	0	0	0	0	2
[158] † Shuvuuia Last Modified in 12/12/22	1001	772	772	0	0	0	0	2
[159] † Similicaudipteryx Last Modified in 12/12/22	1329	444	444	0	0	0	0	2
[160] † Sinornis Last Modified in 12/12/22	1390	383	383	0	0	0	0	2
[161] † Sinornithoides Last Modified in 12/12/22	1329	444	444	0	0	0	0	2
[162] † Sinornithosaurus Last Modified in 12/12/22	1120	653	653	0	0	0	0	2
[163] † Sinovenator Last Modified in 12/12/22	1243	530	530	0	0	0	0	2
[164] † Sinraptor dongi Last Modified in 12/12/22	497	1276	1276	0	0	0	0	2
[165] † Sinusonasus Last Modified in 12/12/22	1505	268	268	0	0	0	0	2
[166] † Sulcavis Last Modified in 12/12/22	1473	300	300	0	0	0	0	2
[167] † Suzhousaurus Last Modified in 12/12/22	1558	215	215	0	0	0	0	2
[168] † Talos Last Modified in 12/12/22	1667	106	106	0	0	0	0	2
[169] † Tanycolagreus Last Modified in 12/12/22	1183	590	590	0	0	0	0	2
[170] † Teviornis Last Modified in 12/12/22	1702	71	71	0	0	0	0	2
[171] † Therizinosaurus Last Modified in 12/12/22	1564	209	209	0	0	0	0	2
[172] † Tianyuraptor Last Modified in 12/12/22	1490	283	283	0	0	0	0	2
[173] † Troodon Last Modified in 12/12/22	1344	429	429	0	0	0	0	2
[174] † Tsaagan Last Modified in 12/12/22	1289	484	484	0	0	0	0	2
[175] † Tugriken shireh Parvicursorinae Last Modified in 12/12/22	1581	192	192	0	0	0	0	2
[176] † Unenlagia comahuensis Last Modified in 12/12/22	1559	214	214	0	0	0	0	2
[177] † Unenlagia paynemili Last Modified in 12/12/22	1687	86	86	0	0	0	0	2
[178] † Utahraptor Last Modified in 12/12/22	1615	158	158	0	0	0	0	2
[179] † Vegavis Last Modified in 12/12/22	1696	77	77	0	0	0	0	2
[180] † Velociraptor mongoliensis Last Modified in 12/12/22	500	1273	1273	0	0	0	0	2
[181] † Vorona Last Modified in 12/12/22	1657	116	116	0	0	0	0	2
[182] † Wulatelong Last Modified in 12/12/22	1569	204	204	0	0	0	0	2
[183] † Xiaotingia Last Modified in 12/12/22	1459	314	314	0	0	0	0	2
[184] † Xixiasaurus Last Modified in 12/12/22	1601	172	172	0	0	0	0	2
[185] † Xixianykus Last Modified in 12/12/22	1615	158	158	0	0	0	0	2
[186] † Yanornis Last Modified in 12/12/22	1391	382	382	0	0	0	0	2
[187] † Yi Last Modified in 12/12/22	1640	133	133	0	0	0	0	2
[188] † Yixianornis Last Modified in 12/12/22	1369	404	404	0	0	0	0	2
[189] † Yixianosaurus Last Modified in 12/12/22	1611	162	162	0	0	0	0	2
[190] † Zanabazar Last Modified in 12/12/22	1421	352	352	0	0	0	0	2
[191] † Zhenyuanlong Last Modified in 12/12/22	1416	357	357	0	0	0	0	2
[192] † Zhongjianornis Last Modified in 12/12/22	1535	238	238	0	0	0	0	2
[193] † Zhongornis Last Modified in 12/12/22	1586	187	187	0	0	0	0	2
[194] † Zhouornis Last Modified in 12/12/22	1363	410	410	0	0	0	0	2
[195] † Zuolong Last Modified in 12/12/22	1340	433	433	0	0	0	0	2
[196] † Anchiornithin sp Last Modified in 12/12/22	1428	345	345	0	0	0	0	2
[197] † Hesperonychus Last Modified in 12/12/22	1722	51	51	0	0	0	0	2
[198] † Hulsanpes Last Modified in 12/12/22	1730	43	43	0	0	0	0	2
[199] † Neuquenraptor Last Modified in 12/12/22	1696	77	77	0	0	0	0	2

Taxonomic Overview for Matrix 'M29101' (153 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Allosaurus fragilis Last Modified in 12/12/22	67	796	793	0	0	0	0	2
[2] † Sinraptor dongi Last Modified in 12/12/22	207	653	653	0	0	0	0	2
[3] † Dromaeosaurus albertensis Last Modified in 12/12/22	610	250	250	0	0	0	0	2
[4] † Deinonychus antirrhopus Last Modified in 12/12/22	253	608	607	0	0	0	0	2
[5] † Velociraptor mongoliensis Last Modified in 12/12/22	112	750	748	0	0	0	0	2
[6] † Balaur bondoc Last Modified in 12/12/22	596	266	264	0	0	0	0	2
[7] † Tsaagan mangas Last Modified in 12/12/22	502	358	358	0	0	0	0	2
[8] † Bambiraptor feinbergorum Last Modified in 12/12/22	230	632	630	0	0	0	0	2
[9] † Tianyuraptor ostromi Last Modified in 12/12/22	680	180	180	0	0	0	0	2
[10] † Sinornithosaurus millenii Last Modified in 12/12/22	433	428	427	0	0	0	0	2
[11] † Microraptor zhaoianus Last Modified in 12/12/22	505	356	355	0	0	0	0	2
[12] † Graciliraptor lujiatunensis Last Modified in 12/12/22	760	101	100	0	0	0	0	2
[13] † Hesperonychus elizabethae Last Modified in 12/12/22	826	34	34	0	0	0	0	2
[14] † Pyroraptor olympius Last Modified in 12/12/22	838	22	22	0	0	0	0	2
[15] † Rahonavis ostromi Last Modified in 12/12/22	633	228	227	0	0	0	0	2
[16] † Buitreraptor gonzalozorum Last Modified in 12/12/22	455	409	405	0	0	0	0	2
[17] † Neuquenraptor plus Unenlagia Last Modified in 12/12/22	624	236	236	0	0	0	0	2
[18] † Austroraptor Last Modified in 12/12/22	683	179	177	0	0	0	0	2
[19] † Shanag ashile Last Modified in 12/12/22	797	63	63	0	0	0	0	2
[20] † Mahakala omnogovae Last Modified in 12/12/22	572	291	288	0	0	0	0	2
[21] † Atrociraptor marshalli Last Modified in 12/12/22	792	69	68	0	0	0	0	2
[22] † Utahraptor Last Modified in 12/12/22	728	133	132	0	0	0	0	2
[23] † Adasaurus mongoliensis Last Modified in 12/12/22	569	293	291	0	0	0	0	2
[24] † Achillobator giganticus Last Modified in 12/12/22	660	200	200	0	0	0	0	2
[25] † Saurornitholestes langsto Last Modified in 12/12/22	687	173	173	0	0	0	0	2
[26] † Saurornithoides mongolien Last Modified in 12/12/22	608	253	252	0	0	0	0	2
[27] † Zanabazar junior Last Modified in 12/12/22	601	260	259	0	0	0	0	2
[28] † Xixiasaurus Last Modified in 12/12/22	729	131	131	0	0	0	0	2
[29] † Byronosaurus jaffei Last Modified in 12/12/22	670	190	190	0	0	0	0	2
[30] † Sinornithoides youngi Last Modified in 12/12/22	560	301	300	0	0	0	0	2
[31] † Mei long Last Modified in 12/12/22	440	422	420	0	0	0	0	2
[32] † Jinfengopteryx elegans Last Modified in 12/12/22	700	160	160	0	0	0	0	2
[33] † Anchiornis huxleyi Last Modified in 12/12/22	485	376	375	0	0	0	0	2
[34] † Xiaotingia Last Modified in 12/12/22	578	283	282	0	0	0	0	2
[35] † Aurornis Last Modified in 12/12/22	595	265	265	0	0	0	0	2
[36] † Eosinopteryx Last Modified in 12/12/22	615	245	245	0	0	0	0	2
[37] † Troodon formosus Last Modified in 12/12/22	597	263	263	0	0	0	0	2
[38] † Sinovenator changii Last Modified in 12/12/22	448	414	412	0	0	0	0	2
[39] † EK troodontid IGM 100 slash 44 Last Modified in 12/12/22	813	47	47	0	0	0	0	2
[40] † IGM 100 slash 1126 Last Modified in 12/12/22	615	246	245	0	0	0	0	2
[41] † IGM 100 slash 1323 Last Modified in 12/12/22	712	149	148	0	0	0	0	2
[42] † Archaeopteryx lithographica Last Modified in 12/12/22	247	615	613	0	0	0	0	2
[43] † Confuciusornis sanctus Last Modified in 12/12/22	379	485	481	0	0	0	0	2
[44] † Jeholornis prima Last Modified in 12/12/22	484	376	376	0	0	0	0	2
[45] † Jixiangornis orientalis Last Modified in 12/12/22	518	343	342	0	0	0	0	2
[46] † Yanornis martini Last Modified in 12/12/22	568	298	292	0	0	0	0	2
[47] † Apsaravis ukhaana Last Modified in 12/12/22	560	305	300	0	0	0	0	2
[48] † Yixianornis Last Modified in 12/12/22	503	362	357	0	0	0	0	2
[49] † Sapeornis Last Modified in 12/12/22	427	436	433	0	0	0	0	2
[50] † Neuquenornis volans Last Modified in 12/12/22	715	148	145	0	0	0	0	2
[51] † Patagopteryx deferrariisi Last Modified in 12/12/22	599	267	261	0	0	0	0	2
[52] † Cathayornis Last Modified in 12/12/22	575	292	285	0	0	0	0	2
[53] † Concornis Last Modified in 12/12/22	698	169	162	0	0	0	0	2
[54] † Gobipteryx Last Modified in 12/12/22	726	137	134	0	0	0	0	2
[55] † Vorona Last Modified in 12/12/22	781	79	79	0	0	0	0	2
[56] † Songlingornis Last Modified in 12/12/22	796	66	64	0	0	0	0	2
[57] † Pengornis houi Last Modified in 12/12/22	583	284	277	0	0	0	0	2
[58] † Hesperornis Last Modified in 12/12/22	501	359	359	0	0	0	0	2
[59] † Baptornis Last Modified in 12/12/22	733	130	127	0	0	0	0	2
[60] † Ichthyornis Last Modified in 12/12/22	430	436	430	0	0	0	0	2
[61] † Iaceornis marshii Last Modified in 12/12/22	738	123	122	0	0	0	0	2
[62] † Limenavis patagonica Last Modified in 12/12/22	820	42	40	0	0	0	0	2
[63] † Lithornis Last Modified in 12/12/22	537	331	323	0	0	0	0	2
[64] † Hongshanornis longicrest Last Modified in 12/12/22	621	243	239	0	0	0	0	2
[65] † Liaoningornis longidigitu Last Modified in 12/12/22	791	69	69	0	0	0	0	2
[66] † Crypturellus undulatus Last Modified in 12/12/22	446	415	414	0	0	0	0	2
[67] † Gallus gallus Last Modified in 12/12/22	444	418	416	0	0	0	0	2
[68] † Crax pauxi Last Modified in 12/12/22	439	421	421	0	0	0	0	2
[69] † Anas platyrhynchus Last Modified in 12/12/22	458	404	402	0	0	0	0	2
[70] † Chauna torquata Last Modified in 12/12/22	439	422	421	0	0	0	0	2
[71] † Pedopenna Last Modified in 12/12/22	828	32	32	0	0	0	0	2
[72] † Epidendrosaurus Last Modified in 12/12/22	766	94	94	0	0	0	0	2
[73] † Epidexipteryx Last Modified in 12/12/22	687	175	173	0	0	0	0	2
[74] † Incisivosaurus gauthieri Last Modified in 12/12/22	575	285	285	0	0	0	0	2
[75] † Citipati osmolskae Last Modified in 12/12/22	296	567	564	0	0	0	0	2
[76] † Oviraptor philoceratops Last Modified in 12/12/22	643	217	217	0	0	0	0	2
[77] † Microvenator celer Last Modified in 12/12/22	615	245	245	0	0	0	0	2
[78] † Caudipteryx zoui Last Modified in 12/12/22	526	335	334	0	0	0	0	2
[79] † Ingenia yanshani Last Modified in 12/12/22	534	327	326	0	0	0	0	2
[80] † Rinchenia mongoliensis Last Modified in 12/12/22	689	171	171	0	0	0	0	2
[81] † Conchoraptor gracilis Last Modified in 12/12/22	532	329	328	0	0	0	0	2
[82] † Chirostenotes pergracilis Last Modified in 12/12/22	559	303	301	0	0	0	0	2
[83] † Avimimus portentosus Last Modified in 12/12/22	531	331	329	0	0	0	0	2
[84] † Falcarius Last Modified in 12/12/22	307	554	553	0	0	0	0	2
[85] † Beipiaosaurus Last Modified in 12/12/22	656	204	204	0	0	0	0	2
[86] † Segnosaurus galbinensis Last Modified in 12/12/22	646	214	214	0	0	0	0	2
[87] † Erlikosaurus andrewsi Last Modified in 12/12/22	522	338	338	0	0	0	0	2
[88] † Alxasaurus elesitaiensis Last Modified in 12/12/22	653	208	207	0	0	0	0	2
[89] † Neimongosaurus Last Modified in 12/12/22	629	232	231	0	0	0	0	2
[90] † Erliansaurus Last Modified in 12/12/22	725	135	135	0	0	0	0	2
[91] † Suzhousaurus Last Modified in 12/12/22	652	209	208	0	0	0	0	2
[92] † Nothronychus graffami Last Modified in 12/12/22	503	359	357	0	0	0	0	2
[93] † Enigmosaurus Last Modified in 12/12/22	795	65	65	0	0	0	0	2
[94] † Nanshiungosaurus brevispinus Last Modified in 12/12/22	821	39	39	0	0	0	0	2
[95] † Therizinosaurus Last Modified in 12/12/22	738	122	122	0	0	0	0	2
[96] † Haplocheirus Last Modified in 12/12/22	344	516	516	0	0	0	0	2
[97] † Alvarezsaurus calvoi Last Modified in 12/12/22	706	154	154	0	0	0	0	2
[98] † Patagonykus puertai Last Modified in 12/12/22	652	208	208	0	0	0	0	2
[99] † Achillesaurus Last Modified in 12/12/22	801	59	59	0	0	0	0	2
[100] † Mononykus olecranus Last Modified in 12/12/22	533	327	327	0	0	0	0	2
[101] † Shuvuuia deserti Last Modified in 12/12/22	250	612	610	0	0	0	0	2
[102] † Parvicursor Last Modified in 12/12/22	726	134	134	0	0	0	0	2
[103] † Albertonykus Last Modified in 12/12/22	835	25	25	0	0	0	0	2
[104] † Albinykus Last Modified in 12/12/22	791	69	69	0	0	0	0	2
[105] † Bonapartenykus Last Modified in 12/12/22	784	76	76	0	0	0	0	2
[106] † Ceratonykus Last Modified in 12/12/22	717	143	143	0	0	0	0	2
[107] † Linhenykus Last Modified in 12/12/22	696	164	164	0	0	0	0	2
[108] † Xixianykus Last Modified in 12/12/22	696	164	164	0	0	0	0	2
[109] † Nqwebasaurus Last Modified in 12/12/22	657	203	203	0	0	0	0	2
[110] † Shenzhousaurus orientalis Last Modified in 12/12/22	599	261	261	0	0	0	0	2
[111] † Ornithomimus edmonticus Last Modified in 12/12/22	337	524	523	0	0	0	0	2
[112] † Archaeornithomimus asiati Last Modified in 12/12/22	514	348	346	0	0	0	0	2
[113] † Anserimimus planinychus Last Modified in 12/12/22	731	129	129	0	0	0	0	2
[114] † Struthiomimus altus Last Modified in 12/12/22	234	627	626	0	0	0	0	2
[115] † Gallimimus bullatus Last Modified in 12/12/22	149	713	711	0	0	0	0	2
[116] † Garudimimus brevipes Last Modified in 12/12/22	347	513	513	0	0	0	0	2
[117] † Pelecanimimus polydon Last Modified in 12/12/22	610	250	250	0	0	0	0	2
[118] † Harpymimus okladnikovi Last Modified in 12/12/22	476	385	384	0	0	0	0	2
[119] † Beishanlong Last Modified in 12/12/22	662	198	198	0	0	0	0	2
[120] † Sinornithomimus Last Modified in 12/12/22	294	566	566	0	0	0	0	2
[121] † Qiupalong Last Modified in 12/12/22	749	111	111	0	0	0	0	2
[122] † Kinnareemimus Last Modified in 12/12/22	803	57	57	0	0	0	0	2
[123] † Huaxiagnathus orientalis Last Modified in 12/12/22	523	338	337	0	0	0	0	2
[124] † Sinosauropteryx prima Last Modified in 12/12/22	494	367	366	0	0	0	0	2
[125] † Compsognathus longipes Last Modified in 12/12/22	414	447	446	0	0	0	0	2
[126] † Juravenator starki Last Modified in 12/12/22	484	377	376	0	0	0	0	2
[127] † Sinocalliopteryx Last Modified in 12/12/22	454	407	406	0	0	0	0	2
[128] † Mirischia Last Modified in 12/12/22	783	77	77	0	0	0	0	2
[129] † Ornitholestes hermanni Last Modified in 12/12/22	336	525	524	0	0	0	0	2
[130] † Coelurus fragilis Last Modified in 12/12/22	631	229	229	0	0	0	0	2
[131] † Tanycolagreus Last Modified in 12/12/22	495	366	365	0	0	0	0	2
[132] † Tugulusaurus Last Modified in 12/12/22	792	68	68	0	0	0	0	2
[133] † Zuolong Last Modified in 12/12/22	489	371	371	0	0	0	0	2
[134] † Bicentenaria Last Modified in 12/12/22	679	181	181	0	0	0	0	2
[135] † Kileskus Last Modified in 12/12/22	788	72	72	0	0	0	0	2
[136] † Guanlong Last Modified in 12/12/22	167	694	693	0	0	0	0	2
[137] † Sinotyrannus Last Modified in 12/12/22	776	84	84	0	0	0	0	2
[138] † Proceratosaurus bradleyi Last Modified in 12/12/22	654	206	206	0	0	0	0	2
[139] † Dilong paradoxus Last Modified in 12/12/22	279	581	581	0	0	0	0	2
[140] † Eotyrannus lengi Last Modified in 12/12/22	584	276	276	0	0	0	0	2
[141] † Juratyrant Last Modified in 12/12/22	702	158	158	0	0	0	0	2
[142] † Xiongguanlong Last Modified in 12/12/22	545	315	315	0	0	0	0	2
[143] † Dryptosaurus Last Modified in 12/12/22	713	148	147	0	0	0	0	2
[144] † Appalachiosaurus Last Modified in 12/12/22	612	248	248	0	0	0	0	2
[145] † Bistahieversor Last Modified in 12/12/22	550	310	310	0	0	0	0	2
[146] † Albertosaurus sacrophagus Last Modified in 12/12/22	160	707	700	0	0	0	0	2
[147] † Gorgosaurus libratus Last Modified in 12/12/22	76	787	784	0	0	0	0	2
[148] † Alioramus Last Modified in 12/12/22	305	556	555	0	0	0	0	2
[149] † Teratophoneus Last Modified in 12/12/22	715	145	145	0	0	0	0	2
[150] † Daspletosaurus Last Modified in 12/12/22	180	683	680	0	0	0	0	2
[151] † Tyrannosaurus rex Last Modified in 12/12/22	52	814	808	0	0	0	0	2
[152] † Tarbosaurus baatar Last Modified in 12/12/22	83	777	777	0	0	0	0	2
[153] † Halszkaraptor escuilliei Last Modified in 12/12/22	557	306	303	0	0	0	0	2

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	5208
Project overview	302
Media views	780	Media search (663 views); M848408 (117 views);
Matrix views	220	Matrix landing page (192 views); Cau et al (2017) (1) (13 views); Cau et al (2017) (2) (15 views);
Documents list	184
Views for media list	236
Bibliography	150
Specimen list	400
Taxon list	2936

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	6
Project downloads	3
Matrix downloads	3	Cau et al (2017) (1) (2 downloads); Cau et al (2017) (2) (1 download);