Morphobank

Project 4469: A. Laboury, R. F. Bennion, B. Thuy, R. Weis, V. Fischer. 2022. Anatomy and phylogenetic relationships of Temnodontosaurus zetlandicus (Reptilia: Ichthyosauria). Zoological Journal of the Linnean Society. 195 (1):172-194.

Specimen: † Ophthalmosaurus icenicus (unvouchered)
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Abstract

Parvipelvia is a major clade of ichthyosaurians that diversified during the Triassic-Jurassic transition. The interrelationships of early parvipelvians remain unclear and many genera are loosely diagnosed, such as Temnodontosaurus, an ecologically important genus from the Early Jurassic of Western Europe. One taxon concentrates many taxonomic issues: ‘Ichthyosaurus’ acutirostris was previously assigned to Temnodontosaurus and for which ‘Ichthyosaurus’ zetlandicus represents a junior synonym. We redescribe the holotype of ‘Ichthyosaurus’ zetlandicus (CAMSM J35176) and a new specimen probably attributable to this taxon (MNHNL TU885) from the Toarcian of Luxembourg. We find that Temnodontosaurus zetlandicus comb. nov. is a valid species that should be referred to the genus Temnodontosaurus, sharing a number of traits with Temnodontosaurus nuertingensis and Temnodontosaurus trigonodon, despite having a distinct cranial architecture. Our phylogenetic analyses under both implied weighting maximum parsimony and Bayesian inference recover T. zetlandicus as closely related to several species currently assigned to Temnodontosaurus. Species included in Temnodontosaurus form a polyphyletic yet well-clustered group among basal neoichthyosaurians, demonstrating that the monophyly of this genus needs to be thoroughly investigated.

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Article DOI: 10.1093/zoolinnean/zlab118

Project DOI: 10.7934/P4469, http://dx.doi.org/10.7934/P4469

This project contains	Matrices
4 Media 1 Matrix 1 Document 110 Taxa 4 Specimens 287 Characters Total size of project's media files: 1000.63k Download Project SDD File	Total scored cells: 16900 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 287 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 622 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:282/5

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MorphoBank Project 4469

Creation Date:
04 November 2022
Publication Date:
13 November 2022
Media downloads: 3
Matrix downloads: 2

Authors' Institutions

Université de Liège

Royal Belgian Institute of Natural Sciences

National museum for Natural History Luxembourg

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Taxonomic Overview for Matrix 'M29064' (110 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Acamptonectes densus Taxon name last Modified on 11/13/22	166	116	116	0	5	0	0	1
[2] † Aegirosaurus leptospondylus Taxon name last Modified on 11/13/22	94	181	181	0	12	0	0	1
[3] † Arthropterygius chrisorum Taxon name last Modified on 11/13/22	223	64	64	0	0	0	0	1
[4] † Athabascasaurus bitumineus Taxon name last Modified on 11/13/22	182	95	95	0	10	0	0	1
[5] † Barracudasauroides panxianensis Taxon name last Modified on 11/13/22	78	202	202	0	7	0	0	1
[6] † Besanosaurus leptorhynchus Taxon name last Modified on 11/13/22	55	226	226	0	6	0	0	1
[7] † Brachypterygius extremus Taxon name last Modified on 11/13/22	118	162	162	0	7	0	0	1
[8] † Brachypterygius pseudoscythica Taxon name last Modified on 11/13/22	201	81	81	0	5	0	0	1
[9] † Californosaurus perrini Taxon name last Modified on 11/13/22	184	101	101	0	2	0	0	1
[10] † Callawayia neoscapularis Taxon name last Modified on 11/13/22	60	218	218	0	9	0	0	1
[11] † Cartorhynchus lenticarpus Taxon name last Modified on 11/13/22	123	141	141	0	23	0	0	1
[12] † Caypullisaurus bonapartei Taxon name last Modified on 11/13/22	101	174	174	0	12	0	0	1
[13] † Chacaicosaurus cayi Taxon name last Modified on 11/13/22	225	57	57	0	5	0	0	1
[14] † Chaohusaurus chaoxianensis Taxon name last Modified on 11/13/22	63	215	215	0	9	0	0	1
[15] † Chaohusaurus geishanensis Taxon name last Modified on 11/13/22	179	105	105	0	3	0	0	1
[16] † Chaohusaurus zhangjiawanensis Taxon name last Modified on 11/13/22	113	159	159	0	15	0	0	1
[17] † Contectopalatus atavus Taxon name last Modified on 11/13/22	54	220	220	0	13	0	0	1
[18] † Cryopterygius kielanae Taxon name last Modified on 11/13/22	220	65	65	0	2	0	0	1
[19] † Cryopterygius kristiansenae Taxon name last Modified on 11/13/22	111	174	174	0	2	0	0	1
[20] † Cymbospondylus buchseri Taxon name last Modified on 11/13/22	162	118	118	0	7	0	0	1
[21] † Cymbospondylus nichollsi Taxon name last Modified on 11/13/22	174	104	104	0	9	0	0	1
[22] † Cymbospondylus petrinus Taxon name last Modified on 11/13/22	51	224	224	0	12	0	0	1
[23] † Eurhinosaurus longirostris Taxon name last Modified on 11/13/22	15	258	258	0	14	0	0	1
[24] † Excalibosaurus costini Taxon name last Modified on 11/13/22	82	195	195	0	10	0	0	1
[25] † Gengasaurus nicosiai Taxon name last Modified on 11/13/22	212	72	72	0	3	0	0	1
[26] † Grendelius alekseevi Taxon name last Modified on 11/13/22	146	134	134	0	7	0	0	1
[27] † Grendelius zhuravlevi Taxon name last Modified on 11/13/22	240	45	45	0	2	0	0	1
[28] † Grippia longirostris Taxon name last Modified on 11/13/22	55	220	220	0	12	0	0	1
[29] † Guizhouichthyosaurus tangae Taxon name last Modified on 11/13/22	28	246	246	0	13	0	0	1
[30] † Guizhouichthyosaurus wolonggangense Taxon name last Modified on 11/13/22	116	167	167	0	4	0	0	1
[31] † Gulosaurus helmi Taxon name last Modified on 11/13/22	127	151	151	0	9	0	0	1
[32] † Hauffiopteryx altera Taxon name last Modified on 11/13/22	127	154	154	0	6	0	0	1
[33] † Hauffiopteryx typicus Taxon name last Modified on 11/13/22	6	281	269	0	12	0	0	1
[34] † Himalayasaurus tibetensis Taxon name last Modified on 11/13/22	278	9	9	0	0	0	0	1
[35] † Hudsonelpidia brevirostris Taxon name last Modified on 11/13/22	194	90	90	0	3	0	0	1
[36] † Hupehsuchus nanchangensis Taxon name last Modified on 11/13/22	83	179	179	0	25	0	0	1
[37] † Ichthyosaurus anningae Taxon name last Modified on 11/13/22	111	169	169	0	7	0	0	1
[38] † Ichthyosaurus breviceps Taxon name last Modified on 11/13/22	59	218	218	0	10	0	0	1
[39] † Ichthyosaurus communis Taxon name last Modified on 11/13/22	0	276	276	0	11	0	0	1
[40] † Ichthyosaurus conybeari Taxon name last Modified on 11/13/22	60	219	219	0	8	0	0	1
[41] † Ichthyosaurus larkini Taxon name last Modified on 11/13/22	70	208	208	0	9	0	0	1
[42] † Ichthyosaurus somersetensis Taxon name last Modified on 11/13/22	41	235	235	0	11	0	0	1
[43] † Janusaurus lundi Taxon name last Modified on 11/13/22	128	148	148	0	11	0	0	1
[44] † Keilhauia nui Taxon name last Modified on 11/13/22	218	66	66	0	3	0	0	1
[45] † Leninia stellans Taxon name last Modified on 11/13/22	205	78	78	0	4	0	0	1
[46] † Leptonectes moorei Taxon name last Modified on 11/13/22	154	128	128	0	5	0	0	1
[47] † Leptonectes solei Taxon name last Modified on 11/13/22	159	126	126	0	2	0	0	1
[48] † Leptonectes tenuirostris Taxon name last Modified on 11/13/22	30	244	244	0	13	0	0	1
[49] † Macgowania janiceps Taxon name last Modified on 11/13/22	131	152	152	0	4	0	0	1
[50] † Maiaspondylus lindoei Taxon name last Modified on 11/13/22	195	88	88	0	4	0	0	1
[51] † Malawania anachronus Taxon name last Modified on 11/13/22	221	62	62	0	4	0	0	1
[52] † Mixosaurus cornalianus Taxon name last Modified on 11/13/22	24	255	255	0	8	0	0	1
[53] † Mixosaurus kuhnschneyderi Taxon name last Modified on 11/13/22	103	177	177	0	7	0	0	1
[54] † Mixosaurus xindianensis Taxon name last Modified on 11/13/22	91	191	191	0	5	0	0	1
[55] † Mollesaurus periallus Taxon name last Modified on 11/13/22	188	97	97	0	2	0	0	1
[56] † Muiscasaurus catheti Taxon name last Modified on 11/13/22	205	79	79	0	3	0	0	1
[57] † Nannopterygius enthekiodon Taxon name last Modified on 11/13/22	199	86	86	0	2	0	0	1
[58] † Ophthalmosaurus icenicus Taxon name last Modified on 11/13/22	1	273	273	0	13	0	0	1
[59] † Ophthalmosaurus natans Taxon name last Modified on 11/13/22	51	223	223	0	13	0	0	1
[60] † Ophthalmosaurus yasykovi Taxon name last Modified on 11/13/22	153	128	128	0	6	0	0	1
[61] † Palvennia hoybergeti Taxon name last Modified on 11/13/22	172	107	107	0	8	0	0	1
[62] † Paraophthalmosaurus kabanovi Taxon name last Modified on 11/13/22	213	70	70	0	4	0	0	1
[63] † Paraophthalmosaurus saveljeviensis Taxon name last Modified on 11/13/22	200	83	83	0	4	0	0	1
[64] † Parvinatator wapitiensis Taxon name last Modified on 11/13/22	185	100	100	0	2	0	0	1
[65] † Pervushovisaurus bannovkensis Taxon name last Modified on 11/13/22	260	27	27	0	0	0	0	1
[66] † Pervushovisaurus campylodon Taxon name last Modified on 11/13/22	265	20	20	0	2	0	0	1
[67] † Pessopteryx nisseri Taxon name last Modified on 11/13/22	215	68	68	0	4	0	0	1
[68] † Phalarodon callawayi Taxon name last Modified on 11/13/22	150	132	132	0	5	0	0	1
[69] † Phalarodon fraasi Taxon name last Modified on 11/13/22	111	167	167	0	9	0	0	1
[70] † Phantomosaurus neubigi Taxon name last Modified on 11/13/22	212	73	73	0	2	0	0	1
[71] † Platypterygius americanus Taxon name last Modified on 11/13/22	108	164	164	0	15	0	0	1
[72] † Platypterygius australis Taxon name last Modified on 11/13/22	13	250	250	0	24	0	0	1
[73] † Platypterygius hauthali Taxon name last Modified on 11/13/22	233	51	51	0	3	0	0	1
[74] † Platypterygius hercynicus Taxon name last Modified on 11/13/22	83	190	190	0	14	0	0	1
[75] † Platypterygius platydactylus Taxon name last Modified on 11/13/22	167	114	114	0	6	0	0	1
[76] † Platypterygius sachicarum Taxon name last Modified on 11/13/22	140	138	138	0	9	0	0	1
[77] † Protoichthyosaurus prostaxalis Taxon name last Modified on 11/13/22	79	198	197	0	11	0	0	1
[78] † Protoichthyosaurus applebyi Taxon name last Modified on 11/13/22	145	131	131	0	11	0	0	1
[79] † Qianichthyosaurus xingyiensis Taxon name last Modified on 11/13/22	126	150	150	0	11	0	0	1
[80] † Qianichthyosaurus zhoui Taxon name last Modified on 11/13/22	70	205	205	0	12	0	0	1
[81] † Quasianosteosaurus vikinghoegdai Taxon name last Modified on 11/13/22	235	50	50	0	2	0	0	1
[82] † Sclerocormus parviceps Taxon name last Modified on 11/13/22	148	120	120	0	19	0	0	1
[83] † Shastasaurus liangae Taxon name last Modified on 11/13/22	63	191	191	0	33	0	0	1
[84] † Shastasaurus pacificus Taxon name last Modified on 11/13/22	99	166	166	0	22	0	0	1
[85] † Shastasaurus sikkaniensis Taxon name last Modified on 11/13/22	176	92	92	0	19	0	0	1
[86] † Shonisaurus popularis Taxon name last Modified on 11/13/22	74	204	204	0	9	0	0	1
[87] † Simbirskiasaurus birjukovi Taxon name last Modified on 11/13/22	232	52	52	0	3	0	0	1
[88] † Sisteronia seeleyi Taxon name last Modified on 11/13/22	237	49	49	0	1	0	0	1
[89] † Stenopterygius aaleniensis Taxon name last Modified on 11/13/22	112	167	167	0	8	0	0	1
[90] † Stenopterygius quadriscissus Taxon name last Modified on 11/13/22	9	267	266	0	12	0	0	1
[91] † Stenopterygius triscissus Taxon name last Modified on 11/13/22	8	267	267	0	12	0	0	1
[92] † Stenopterygius uniter Taxon name last Modified on 11/13/22	45	233	233	0	9	0	0	1
[93] † Suevoleviathan integer Taxon name last Modified on 11/13/22	23	252	252	0	12	0	0	1
[94] † Sveltonectes insolitus Taxon name last Modified on 11/13/22	77	199	199	0	11	0	0	1
[95] † Temnodontosaurus azerguensis Taxon name last Modified on 11/13/22	147	114	114	0	26	0	0	1
[96] † Temnodontosaurus crassimanus Taxon name last Modified on 11/13/22	130	149	149	0	8	0	0	1
[97] † Temnodontosaurus eurycephalus Taxon name last Modified on 11/13/22	218	68	68	0	1	0	0	1
[98] † Temnodontosaurus platyodon Taxon name last Modified on 11/13/22	8	263	263	0	16	0	0	1
[99] † Temnodontosaurus nuertingensis Taxon name last Modified on 11/13/22	207	78	78	0	2	0	0	1
[100] † Temnodontosaurus trigonodon Taxon name last Modified on 11/13/22	8	265	265	0	14	0	0	1
[101] † Ichthyosaurus zetlandicus Taxon name last Modified on 11/13/22	173	111	111	0	3	0	0	1
[102] † Thalattoarchon saurophagis Taxon name last Modified on 11/13/22	209	76	76	0	2	0	0	1
[103] † Toretocnemus californicus Taxon name last Modified on 11/13/22	220	63	63	0	4	0	0	1
[104] † Toretocnemus zitteli Taxon name last Modified on 11/13/22	162	115	115	0	10	0	0	1
[105] † Undorosaurus gorodischensis Taxon name last Modified on 11/13/22	165	117	117	0	5	0	0	1
[106] † Undorosaurus trautscholdi Taxon name last Modified on 11/13/22	235	49	49	0	3	0	0	1
[107] † Utatsusaurus hataii Taxon name last Modified on 11/13/22	88	191	191	0	8	0	0	1
[108] † Wahlisaurus massarae Taxon name last Modified on 11/13/22	173	112	112	0	2	0	0	1
[109] † Wimanius odontopalatus Taxon name last Modified on 11/13/22	231	54	54	0	2	0	0	1
[110] † Xinminosaurus catactes Taxon name last Modified on 11/13/22	182	100	100	0	5	0	0	1

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	9
Document downloads	1	Original data matrix, NEXUS (1 download);
Project downloads	3
Matrix downloads	2	Data matrix from author with minor modifications (2 downloads);
Media downloads	3	M848046 (2 downloads); M848043 (1 download);