Project 4531: R. O. Gómez, G. F. Turazzini. 2021. The fossil record and phylogeny of South American horned frogs (Anura, Ceratophryidae). Journal of Systematic Palaeontology. 19 (2):91-130.
Abstract
South American horned frogs (Ceratophryidae), with their large heads, wide gapes and fang-like teeth, are among the most charismatic, best-known and well-studied neobatrachian anurans. The family comprises 12 extant species with hyperossified skulls and has a relatively rich fossil record, particularly in the Pampas, which dates back to the late Miocene. However, several records have been overlooked in recent summaries, and many taxonomic assignments remain indeterminate or are questionable and have yet to be tested within a quantitative phylogenetic framework. Here we provide a complete up-to-date survey of the palaeontological record of Ceratophryidae, including some remarkable new records. We also tested their systematic position through comprehensive phylogenetic analyses based on osteological data, providing several synapomorphies for all relevant nodes. Finally, we discuss these integrated data in relation to divergence time estimates, and propose a set of fossil calibrations that provide hard minimum bounds for crown-group Ceratophryidae and the subclades within it, and illuminate the acquisition of polyploidy within the group.Read the article »
Article DOI: 10.1080/14772019.2021.1892845
Project DOI: 10.7934/P4531, http://dx.doi.org/10.7934/P4531
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 10973 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 256 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 747 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:256/0 |
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MorphoBank Project 4531
MorphoBank Project 4531
- Creation Date:
20 January 2023 - Publication Date:
14 March 2023 - Matrix downloads: 2
Authors' Institutions
- Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)
- Universidad de Buenos Aires
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Raul Gomez Project Administrator | 73 | 1 | 1 | 256 | 0 | 0 | 10973 (10973, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29193' (71 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Alsodes nodosus Taxon name last Modified on 03/14/23 | 18 | 238 | 238 | 0 | 0 | 0 | 0 | 1 |
[2] Atelognathus patagonicus Taxon name last Modified on 03/14/23 | 26 | 237 | 230 | 0 | 0 | 0 | 0 | 1 |
[3] Batrachyla taeniata Taxon name last Modified on 03/14/23 | 21 | 238 | 235 | 0 | 0 | 0 | 0 | 1 |
[4] † C ameghinorum 14322 25 Taxon name last Modified on 03/14/23 | 163 | 93 | 93 | 0 | 0 | 0 | 0 | 1 |
[5] † C ameghinorum 14324 Taxon name last Modified on 03/14/23 | 199 | 57 | 57 | 0 | 0 | 0 | 0 | 1 |
[6] † C ameghinorum hol14318 Taxon name last Modified on 03/14/23 | 142 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[7] † C ameghinorum par 19731 Taxon name last Modified on 03/14/23 | 92 | 164 | 164 | 0 | 0 | 0 | 0 | 1 |
[8] † C ameghinorum par14317 Taxon name last Modified on 03/14/23 | 142 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[9] † C ameghinorum par14319 Taxon name last Modified on 03/14/23 | 142 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[10] † Ceratophrys ”ensenadensis” PVL616 Taxon name last Modified on 03/14/23 | 214 | 43 | 42 | 0 | 0 | 0 | 0 | 1 |
[11] † Ceratophrys ”rusconii” Taxon name last Modified on 03/14/23 | 148 | 109 | 108 | 0 | 0 | 0 | 0 | 1 |
[12] Ceratophrys aurita Taxon name last Modified on 03/14/23 | 7 | 265 | 249 | 0 | 0 | 0 | 0 | 1 |
[13] Ceratophrys calcarata Taxon name last Modified on 03/14/23 | 7 | 256 | 249 | 0 | 0 | 0 | 0 | 1 |
[14] Ceratophrys cornuta Taxon name last Modified on 03/14/23 | 4 | 262 | 252 | 0 | 0 | 0 | 0 | 1 |
[15] Ceratophrys cranwelli Taxon name last Modified on 03/14/23 | 1 | 279 | 255 | 0 | 0 | 0 | 0 | 1 |
[16] Ceratophrys joazeirensis Taxon name last Modified on 03/14/23 | 110 | 148 | 146 | 0 | 0 | 0 | 0 | 1 |
[17] Ce ornata FCEN1882(1) Taxon name last Modified on 03/14/23 | 2 | 254 | 254 | 0 | 0 | 0 | 0 | 1 |
[18] Ce ornata FCEN1882(9) Taxon name last Modified on 03/14/23 | 3 | 253 | 253 | 0 | 0 | 0 | 0 | 1 |
[19] Ceratophrys ornata Hudson Taxon name last Modified on 03/14/23 | 13 | 243 | 243 | 0 | 0 | 0 | 0 | 1 |
[20] Ceratophrys stolzmanni Taxon name last Modified on 03/14/23 | 20 | 240 | 236 | 0 | 0 | 0 | 0 | 1 |
[21] Chacophrys pierottii Taxon name last Modified on 03/14/23 | 4 | 263 | 252 | 0 | 0 | 0 | 0 | 1 |
[22] Crossodactylus trachystomus Taxon name last Modified on 03/14/23 | 30 | 226 | 226 | 0 | 0 | 0 | 0 | 1 |
[23] Cycloramphus dubius Taxon name last Modified on 03/14/23 | 21 | 238 | 235 | 0 | 0 | 0 | 0 | 1 |
[24] Eupsophus roseus Taxon name last Modified on 03/14/23 | 18 | 246 | 238 | 0 | 0 | 0 | 0 | 1 |
[25] † FCS.PA1.E1.P4.1 paso Alsina Taxon name last Modified on 03/14/23 | 236 | 20 | 20 | 0 | 0 | 0 | 0 | 1 |
[26] † FCS.SA1.S1.525.1 san Antonio Taxon name last Modified on 03/14/23 | 238 | 19 | 18 | 0 | 0 | 0 | 0 | 1 |
[27] † GHUNLPam8633 cerro Azul Fm Taxon name last Modified on 03/14/23 | 240 | 17 | 16 | 0 | 0 | 0 | 0 | 1 |
[28] Hylodes asper Taxon name last Modified on 03/14/23 | 21 | 236 | 235 | 0 | 0 | 0 | 0 | 1 |
[29] Hylorina sylvatica Taxon name last Modified on 03/14/23 | 26 | 233 | 230 | 0 | 0 | 0 | 0 | 1 |
[30] † ianigla.pv112 Taxon name last Modified on 03/14/23 | 107 | 156 | 149 | 0 | 0 | 0 | 0 | 1 |
[31] Lepidobatrachus asper Taxon name last Modified on 03/14/23 | 2 | 262 | 254 | 0 | 0 | 0 | 0 | 1 |
[32] † Lepidobatrachus australis MMH85.12.2a Taxon name last Modified on 03/14/23 | 155 | 102 | 101 | 0 | 0 | 0 | 0 | 1 |
[33] Lepidobatrachus laevis Taxon name last Modified on 03/14/23 | 2 | 267 | 254 | 0 | 0 | 0 | 0 | 1 |
[34] Lepidobatrachus llanensis Taxon name last Modified on 03/14/23 | 2 | 267 | 254 | 0 | 0 | 0 | 0 | 1 |
[35] Leptodactylus latrans Taxon name last Modified on 03/14/23 | 19 | 238 | 237 | 0 | 0 | 0 | 0 | 1 |
[36] Limnomedusa macroglossa Taxon name last Modified on 03/14/23 | 21 | 238 | 235 | 0 | 0 | 0 | 0 | 1 |
[37] † MACN a10434 Colhué Huapi Taxon name last Modified on 03/14/23 | 242 | 15 | 14 | 0 | 0 | 0 | 0 | 1 |
[38] † MACNPv17911 p Loberia Taxon name last Modified on 03/14/23 | 227 | 30 | 29 | 0 | 0 | 0 | 0 | 1 |
[39] † MACNPv18074 chapadmalal Taxon name last Modified on 03/14/23 | 224 | 33 | 32 | 0 | 0 | 0 | 0 | 1 |
[40] † MACNPv19659 punta San Andres Taxon name last Modified on 03/14/23 | 179 | 77 | 77 | 0 | 0 | 0 | 0 | 1 |
[41] † MD.CH.06.165 chasico Taxon name last Modified on 03/14/23 | 214 | 47 | 42 | 0 | 0 | 0 | 0 | 1 |
[42] † MLP 86.iii.25.150 Tapalqué Taxon name last Modified on 03/14/23 | 214 | 43 | 42 | 0 | 0 | 0 | 0 | 1 |
[43] † MLP 86.iii.25.151 Tapalqué Taxon name last Modified on 03/14/23 | 238 | 19 | 18 | 0 | 0 | 0 | 0 | 1 |
[44] † MLP04.V.2.701 centinela Taxon name last Modified on 03/14/23 | 74 | 183 | 182 | 0 | 0 | 0 | 0 | 1 |
[45] † MLP04.V.2.702 centinela Taxon name last Modified on 03/14/23 | 225 | 32 | 31 | 0 | 0 | 0 | 0 | 1 |
[46] † MLP86.VIII.1.4 tres Reyes Taxon name last Modified on 03/14/23 | 146 | 110 | 110 | 0 | 0 | 0 | 0 | 1 |
[47] † MLP86.VIII.1.5 tres Reyes Taxon name last Modified on 03/14/23 | 145 | 112 | 111 | 0 | 0 | 0 | 0 | 1 |
[48] † MLP86.VIII.1.6 tres Reyes Taxon name last Modified on 03/14/23 | 167 | 89 | 89 | 0 | 0 | 0 | 0 | 1 |
[49] † MLP88.V.20.1 mar del Plata Taxon name last Modified on 03/14/23 | 141 | 116 | 115 | 0 | 0 | 0 | 0 | 1 |
[50] † MLP89.XII.35.25 bca Parodi Taxon name last Modified on 03/14/23 | 208 | 51 | 48 | 0 | 0 | 0 | 0 | 1 |
[51] † MLP94.II.1.171 q Salado Taxon name last Modified on 03/14/23 | 153 | 104 | 103 | 0 | 0 | 0 | 0 | 1 |
[52] † MLP97.V.1.19 la Estafeta Taxon name last Modified on 03/14/23 | 165 | 91 | 91 | 0 | 0 | 0 | 0 | 1 |
[53] † MMP 4680 Taxon name last Modified on 03/14/23 | 193 | 67 | 63 | 0 | 0 | 0 | 0 | 1 |
[54] † MMP733 bajada de Los Lobos Taxon name last Modified on 03/14/23 | 189 | 68 | 67 | 0 | 0 | 0 | 0 | 1 |
[55] † MMPM4846 daireaux Taxon name last Modified on 03/14/23 | 220 | 38 | 36 | 0 | 0 | 0 | 0 | 1 |
[56] † MNHN un Ñuapua Taxon name last Modified on 03/14/23 | 173 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[57] † MPH.P0124 las Brusquitas Taxon name last Modified on 03/14/23 | 199 | 61 | 57 | 0 | 0 | 0 | 0 | 1 |
[58] † MPS127 san Pedro Taxon name last Modified on 03/14/23 | 168 | 88 | 88 | 0 | 0 | 0 | 0 | 1 |
[59] † NHMUK 18896 Lagoa Santa Taxon name last Modified on 03/14/23 | 179 | 85 | 77 | 0 | 0 | 0 | 0 | 1 |
[60] Odontophrynus americanus Taxon name last Modified on 03/14/23 | 14 | 257 | 242 | 0 | 0 | 0 | 0 | 1 |
[61] Proceratophrys boiei Taxon name last Modified on 03/14/23 | 5 | 254 | 251 | 0 | 0 | 0 | 0 | 1 |
[62] † PVSJ284 ullum Taxon name last Modified on 03/14/23 | 216 | 45 | 40 | 0 | 0 | 0 | 0 | 1 |
[63] Rhinoderma darwinii Taxon name last Modified on 03/14/23 | 43 | 215 | 213 | 0 | 0 | 0 | 0 | 1 |
[64] Telmatobius hauthali Taxon name last Modified on 03/14/23 | 23 | 237 | 233 | 0 | 0 | 0 | 0 | 1 |
[65] Telmatobius marmoratus Taxon name last Modified on 03/14/23 | 60 | 200 | 196 | 0 | 0 | 0 | 0 | 1 |
[66] Telmatobius oxycephalus Taxon name last Modified on 03/14/23 | 24 | 238 | 232 | 0 | 0 | 0 | 0 | 1 |
[67] Telmatobius scrocchii Taxon name last Modified on 03/14/23 | 23 | 240 | 233 | 0 | 0 | 0 | 0 | 1 |
[68] Telmatobius thompsoni Taxon name last Modified on 03/14/23 | 24 | 234 | 232 | 0 | 0 | 0 | 0 | 1 |
[69] Telmatobius verrucosus Taxon name last Modified on 03/14/23 | 75 | 183 | 181 | 0 | 0 | 0 | 0 | 1 |
[70] Thoropa taophora Taxon name last Modified on 03/14/23 | 16 | 244 | 240 | 0 | 0 | 0 | 0 | 1 |
[71] Zachaenus parvulus Taxon name last Modified on 03/14/23 | 23 | 235 | 233 | 0 | 0 | 0 | 0 | 1 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 6 | |
Project downloads | 3 | |
Matrix downloads | 2 | The fossil record and phylogeny of South American horned frogs (Anura, Ceratophryidae) (2 downloads); |
Document downloads | 1 | The fossil record and phylogeny of South American horned frogs (Anura, Ceratophryidae) - Supplemental online material (1 download); |