Project 4531: R. O. Gómez, G. F. Turazzini. 2021. The fossil record and phylogeny of South American horned frogs (Anura, Ceratophryidae). Journal of Systematic Palaeontology. 19 (2):91-130.
Abstract
South American horned frogs (Ceratophryidae), with their large heads, wide gapes and fang-like teeth, are among the most charismatic, best-known and well-studied neobatrachian anurans. The family comprises 12 extant species with hyperossified skulls and has a relatively rich fossil record, particularly in the Pampas, which dates back to the late Miocene. However, several records have been overlooked in recent summaries, and many taxonomic assignments remain indeterminate or are questionable and have yet to be tested within a quantitative phylogenetic framework. Here we provide a complete up-to-date survey of the palaeontological record of Ceratophryidae, including some remarkable new records. We also tested their systematic position through comprehensive phylogenetic analyses based on osteological data, providing several synapomorphies for all relevant nodes. Finally, we discuss these integrated data in relation to divergence time estimates, and propose a set of fossil calibrations that provide hard minimum bounds for crown-group Ceratophryidae and the subclades within it, and illuminate the acquisition of polyploidy within the group.Read the article »
Article DOI: 10.1080/14772019.2021.1892845
Project DOI: 10.7934/P4531, http://dx.doi.org/10.7934/P4531
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 10973 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 256 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 747 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:256/0 |
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MorphoBank Project 4531
MorphoBank Project 4531
- Creation Date:
20 January 2023 - Publication Date:
14 March 2023 - Project views: 3691
- Matrix downloads: 1
Authors' Institutions
- Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)
- Universidad de Buenos Aires
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Raul Gomez Project Administrator | 73 | 1 | 1 | 256 | 0 | 0 | 10973 (10973, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29193' (71 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Alsodes nodosus Last Modified in 03/14/23 | 18 | 238 | 238 | 0 | 0 | 0 | 0 | 1 |
[2] Atelognathus patagonicus Last Modified in 03/14/23 | 26 | 237 | 230 | 0 | 0 | 0 | 0 | 1 |
[3] Batrachyla taeniata Last Modified in 03/14/23 | 21 | 238 | 235 | 0 | 0 | 0 | 0 | 1 |
[4] † C ameghinorum 14322 25 Last Modified in 03/14/23 | 163 | 93 | 93 | 0 | 0 | 0 | 0 | 1 |
[5] † C ameghinorum 14324 Last Modified in 03/14/23 | 199 | 57 | 57 | 0 | 0 | 0 | 0 | 1 |
[6] † C ameghinorum hol14318 Last Modified in 03/14/23 | 142 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[7] † C ameghinorum par 19731 Last Modified in 03/14/23 | 92 | 164 | 164 | 0 | 0 | 0 | 0 | 1 |
[8] † C ameghinorum par14317 Last Modified in 03/14/23 | 142 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[9] † C ameghinorum par14319 Last Modified in 03/14/23 | 142 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[10] † Ceratophrys ”ensenadensis” PVL616 Last Modified in 03/14/23 | 214 | 43 | 42 | 0 | 0 | 0 | 0 | 1 |
[11] † Ceratophrys ”rusconii” Last Modified in 03/14/23 | 148 | 109 | 108 | 0 | 0 | 0 | 0 | 1 |
[12] Ceratophrys aurita Last Modified in 03/14/23 | 7 | 265 | 249 | 0 | 0 | 0 | 0 | 1 |
[13] Ceratophrys calcarata Last Modified in 03/14/23 | 7 | 256 | 249 | 0 | 0 | 0 | 0 | 1 |
[14] Ceratophrys cornuta Last Modified in 03/14/23 | 4 | 262 | 252 | 0 | 0 | 0 | 0 | 1 |
[15] Ceratophrys cranwelli Last Modified in 03/14/23 | 1 | 279 | 255 | 0 | 0 | 0 | 0 | 1 |
[16] Ceratophrys joazeirensis Last Modified in 03/14/23 | 110 | 148 | 146 | 0 | 0 | 0 | 0 | 1 |
[17] Ce ornata FCEN1882(1) Last Modified in 03/14/23 | 2 | 254 | 254 | 0 | 0 | 0 | 0 | 1 |
[18] Ce ornata FCEN1882(9) Last Modified in 03/14/23 | 3 | 253 | 253 | 0 | 0 | 0 | 0 | 1 |
[19] Ceratophrys ornata Hudson Last Modified in 03/14/23 | 13 | 243 | 243 | 0 | 0 | 0 | 0 | 1 |
[20] Ceratophrys stolzmanni Last Modified in 03/14/23 | 20 | 240 | 236 | 0 | 0 | 0 | 0 | 1 |
[21] Chacophrys pierottii Last Modified in 03/14/23 | 4 | 263 | 252 | 0 | 0 | 0 | 0 | 1 |
[22] Crossodactylus trachystomus Last Modified in 03/14/23 | 30 | 226 | 226 | 0 | 0 | 0 | 0 | 1 |
[23] Cycloramphus dubius Last Modified in 03/14/23 | 21 | 238 | 235 | 0 | 0 | 0 | 0 | 1 |
[24] Eupsophus roseus Last Modified in 03/14/23 | 18 | 246 | 238 | 0 | 0 | 0 | 0 | 1 |
[25] † FCS.PA1.E1.P4.1 paso Alsina Last Modified in 03/14/23 | 236 | 20 | 20 | 0 | 0 | 0 | 0 | 1 |
[26] † FCS.SA1.S1.525.1 san Antonio Last Modified in 03/14/23 | 238 | 19 | 18 | 0 | 0 | 0 | 0 | 1 |
[27] † GHUNLPam8633 cerro Azul Fm Last Modified in 03/14/23 | 240 | 17 | 16 | 0 | 0 | 0 | 0 | 1 |
[28] Hylodes asper Last Modified in 03/14/23 | 21 | 236 | 235 | 0 | 0 | 0 | 0 | 1 |
[29] Hylorina sylvatica Last Modified in 03/14/23 | 26 | 233 | 230 | 0 | 0 | 0 | 0 | 1 |
[30] † ianigla.pv112 Last Modified in 03/14/23 | 107 | 156 | 149 | 0 | 0 | 0 | 0 | 1 |
[31] Lepidobatrachus asper Last Modified in 03/14/23 | 2 | 262 | 254 | 0 | 0 | 0 | 0 | 1 |
[32] † Lepidobatrachus australis MMH85.12.2a Last Modified in 03/14/23 | 155 | 102 | 101 | 0 | 0 | 0 | 0 | 1 |
[33] Lepidobatrachus laevis Last Modified in 03/14/23 | 2 | 267 | 254 | 0 | 0 | 0 | 0 | 1 |
[34] Lepidobatrachus llanensis Last Modified in 03/14/23 | 2 | 267 | 254 | 0 | 0 | 0 | 0 | 1 |
[35] Leptodactylus latrans Last Modified in 03/14/23 | 19 | 238 | 237 | 0 | 0 | 0 | 0 | 1 |
[36] Limnomedusa macroglossa Last Modified in 03/14/23 | 21 | 238 | 235 | 0 | 0 | 0 | 0 | 1 |
[37] † MACN a10434 Colhué Huapi Last Modified in 03/14/23 | 242 | 15 | 14 | 0 | 0 | 0 | 0 | 1 |
[38] † MACNPv17911 p Loberia Last Modified in 03/14/23 | 227 | 30 | 29 | 0 | 0 | 0 | 0 | 1 |
[39] † MACNPv18074 chapadmalal Last Modified in 03/14/23 | 224 | 33 | 32 | 0 | 0 | 0 | 0 | 1 |
[40] † MACNPv19659 punta San Andres Last Modified in 03/14/23 | 179 | 77 | 77 | 0 | 0 | 0 | 0 | 1 |
[41] † MD.CH.06.165 chasico Last Modified in 03/14/23 | 214 | 47 | 42 | 0 | 0 | 0 | 0 | 1 |
[42] † MLP 86.iii.25.150 Tapalqué Last Modified in 03/14/23 | 214 | 43 | 42 | 0 | 0 | 0 | 0 | 1 |
[43] † MLP 86.iii.25.151 Tapalqué Last Modified in 03/14/23 | 238 | 19 | 18 | 0 | 0 | 0 | 0 | 1 |
[44] † MLP04.V.2.701 centinela Last Modified in 03/14/23 | 74 | 183 | 182 | 0 | 0 | 0 | 0 | 1 |
[45] † MLP04.V.2.702 centinela Last Modified in 03/14/23 | 225 | 32 | 31 | 0 | 0 | 0 | 0 | 1 |
[46] † MLP86.VIII.1.4 tres Reyes Last Modified in 03/14/23 | 146 | 110 | 110 | 0 | 0 | 0 | 0 | 1 |
[47] † MLP86.VIII.1.5 tres Reyes Last Modified in 03/14/23 | 145 | 112 | 111 | 0 | 0 | 0 | 0 | 1 |
[48] † MLP86.VIII.1.6 tres Reyes Last Modified in 03/14/23 | 167 | 89 | 89 | 0 | 0 | 0 | 0 | 1 |
[49] † MLP88.V.20.1 mar del Plata Last Modified in 03/14/23 | 141 | 116 | 115 | 0 | 0 | 0 | 0 | 1 |
[50] † MLP89.XII.35.25 bca Parodi Last Modified in 03/14/23 | 208 | 51 | 48 | 0 | 0 | 0 | 0 | 1 |
[51] † MLP94.II.1.171 q Salado Last Modified in 03/14/23 | 153 | 104 | 103 | 0 | 0 | 0 | 0 | 1 |
[52] † MLP97.V.1.19 la Estafeta Last Modified in 03/14/23 | 165 | 91 | 91 | 0 | 0 | 0 | 0 | 1 |
[53] † MMP 4680 Last Modified in 03/14/23 | 193 | 67 | 63 | 0 | 0 | 0 | 0 | 1 |
[54] † MMP733 bajada de Los Lobos Last Modified in 03/14/23 | 189 | 68 | 67 | 0 | 0 | 0 | 0 | 1 |
[55] † MMPM4846 daireaux Last Modified in 03/14/23 | 220 | 38 | 36 | 0 | 0 | 0 | 0 | 1 |
[56] † MNHN un Ñuapua Last Modified in 03/14/23 | 173 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[57] † MPH.P0124 las Brusquitas Last Modified in 03/14/23 | 199 | 61 | 57 | 0 | 0 | 0 | 0 | 1 |
[58] † MPS127 san Pedro Last Modified in 03/14/23 | 168 | 88 | 88 | 0 | 0 | 0 | 0 | 1 |
[59] † NHMUK 18896 Lagoa Santa Last Modified in 03/14/23 | 179 | 85 | 77 | 0 | 0 | 0 | 0 | 1 |
[60] Odontophrynus americanus Last Modified in 03/14/23 | 14 | 257 | 242 | 0 | 0 | 0 | 0 | 1 |
[61] Proceratophrys boiei Last Modified in 03/14/23 | 5 | 254 | 251 | 0 | 0 | 0 | 0 | 1 |
[62] † PVSJ284 ullum Last Modified in 03/14/23 | 216 | 45 | 40 | 0 | 0 | 0 | 0 | 1 |
[63] Rhinoderma darwinii Last Modified in 03/14/23 | 43 | 215 | 213 | 0 | 0 | 0 | 0 | 1 |
[64] Telmatobius hauthali Last Modified in 03/14/23 | 23 | 237 | 233 | 0 | 0 | 0 | 0 | 1 |
[65] Telmatobius marmoratus Last Modified in 03/14/23 | 60 | 200 | 196 | 0 | 0 | 0 | 0 | 1 |
[66] Telmatobius oxycephalus Last Modified in 03/14/23 | 24 | 238 | 232 | 0 | 0 | 0 | 0 | 1 |
[67] Telmatobius scrocchii Last Modified in 03/14/23 | 23 | 240 | 233 | 0 | 0 | 0 | 0 | 1 |
[68] Telmatobius thompsoni Last Modified in 03/14/23 | 24 | 234 | 232 | 0 | 0 | 0 | 0 | 1 |
[69] Telmatobius verrucosus Last Modified in 03/14/23 | 75 | 183 | 181 | 0 | 0 | 0 | 0 | 1 |
[70] Thoropa taophora Last Modified in 03/14/23 | 16 | 244 | 240 | 0 | 0 | 0 | 0 | 1 |
[71] Zachaenus parvulus Last Modified in 03/14/23 | 23 | 235 | 233 | 0 | 0 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 3691 | |
Documents list | 158 | |
Project overview | 329 | |
Matrix views | 180 | Matrix landing page (170 views); The fossil record and phylogeny of South American horned frogs (Anura, Ceratophryidae) (10 views); |
Media views | 685 | Media search (574 views); M851971 (111 views); |
Views for media list | 243 | |
Specimen list | 393 | |
Taxon list | 1565 | |
Bibliography | 138 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 2 | |
Project downloads | 1 | |
Matrix downloads | 1 | The fossil record and phylogeny of South American horned frogs (Anura, Ceratophryidae) (1 download); |