Project 4545: E. M. Obraztsova, S. A. Krasnolutskii, V. B. Sukhanov, I. G. Danilov. 2022. Xinjiangchelyid turtles from the Middle Jurassic of the Berezovsk coal mine (Krasnoyarsk Territory, Russia): systematics, skeletal morphology, variation, relationships and palaeobiogeographic implications. Journal of Systematic Palaeontology. 20 (1):1-61.
Abstract
This paper is devoted to the description of a new species of xinjiangchelyid turtle – Annemys variabilis sp. nov. – represented by thousands of isolated bones and several more complete specimens from the Middle Jurassic (Bathonian) Itat Formation of the Berezovsk coal mine, Krasnoyarsk Territory, western Siberia, Russia. The description is based on a sample of the best-preserved specimens. The new species differs from other Annemys spp. by a combination of cranial and shell characters. The basisphenoids (about 200 specimens) of A. variabilis demonstrate variation in 12 characters, some of which are reported in turtles for the first time. About 20 shell characters of A. variabilis are subject to ontogenetic or interindividual variation. Some of these characters are variable in other Annemys spp. and other xinjiangchelyids. The formula of the cervical vertebrae [(2(3()4()5()6()7()8) or (2(3()4()5()6()7()8(] with two opisthocoelous vertebrae (2 and 3), four or five amphicoelous vertebrae (4–7 or 4–8), and one procoelous vertebra is unique for this species among xinjiangchelyids, which were previously known to have only amphicoelous cervicals. Other characters of the non-shell postcranium correspond to those of other xinjiangchelyids. Of the three phylogenetic analyses performed in this study, Analysis 1 does not support the monophyly of either Xinjiangchelyidae or Annemys, Analysis 2 shows paraphyly of Xinjiangchelyidae and monophyly of the Annemys clade (A. latiens, A. levensis, A. variabilis and A. wusu), and Analysis 3 supports monophyly of the Annemys clade with A. latiens, A. levensis and A. variabilis, only in the majority rule consensus tree. The diversity of the turtle assemblage of the Itat Formation is re-assessed as containing 2–3 taxa (A. variabilis [=Testudines indet. 2], Testudines indet. 1 and 3), which is in agreement with similar diversities seen in some other Middle Jurassic Asian turtle assemblages. The known record of Annemys spp. is restricted to the northern part of Asia.Read the article »
Article DOI: 10.1080/14772019.2022.2093662
Project DOI: 10.7934/P4545, http://dx.doi.org/10.7934/P4545
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 711 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 31 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 65 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:31/0 |
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MorphoBank Project 4545
MorphoBank Project 4545
- Creation Date:
27 January 2023 - Publication Date:
14 March 2023 - Project views: 3167
- Matrix downloads: 1
Authors' Institutions
- Zoological Institute of the Russian Academy of Sciences
- Sharypovo Regional Museum
- Borissyak Paleontological Institute of the Russian Academy of Sciences
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 26 | 1 | 1 | 31 | 0 | 0 | 711 (711, 0, 0) | 0 | 0 | 0 | ||||
Igor Danilov Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29194' (26 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Kayentachelys Last Modified in 03/14/23 | 3 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[2] † Bashuchelys youngi Last Modified in 03/14/23 | 11 | 21 | 20 | 0 | 0 | 0 | 0 | 2 |
[3] † Bzigongensis Last Modified in 03/14/23 | 3 | 30 | 28 | 0 | 0 | 0 | 0 | 2 |
[4] † Chuannanchelys dashanpuensis Last Modified in 03/14/23 | 5 | 26 | 26 | 0 | 0 | 0 | 0 | 2 |
[5] † Protoxinjiangchelys salis Last Modified in 03/14/23 | 7 | 24 | 24 | 0 | 0 | 0 | 0 | 2 |
[6] † Baenoides cf. Last Modified in 03/14/23 | 5 | 29 | 26 | 0 | 0 | 0 | 0 | 2 |
[7] † Xinjiangchelys latimarginalis Last Modified in 03/14/23 | 2 | 30 | 29 | 0 | 0 | 0 | 0 | 2 |
[8] † Tienfuchelys chungkingensis Last Modified in 03/14/23 | 9 | 22 | 22 | 0 | 0 | 0 | 0 | 2 |
[9] † Tienfuchelys tzuyangensis Last Modified in 03/14/23 | 4 | 29 | 27 | 0 | 0 | 0 | 0 | 2 |
[10] † Xinjiangchelys junggarensis_merged Last Modified in 03/14/23 | 1 | 35 | 30 | 0 | 0 | 0 | 0 | 2 |
[11] † Tianshanensis Last Modified in 03/14/23 | 2 | 29 | 29 | 0 | 0 | 0 | 0 | 2 |
[12] † Annemys levensis Last Modified in 03/14/23 | 1 | 30 | 30 | 0 | 0 | 0 | 0 | 2 |
[13] † Annemys latiens Last Modified in 03/14/23 | 2 | 31 | 29 | 0 | 0 | 0 | 0 | 2 |
[14] † Pzigongensis Last Modified in 03/14/23 | 2 | 32 | 29 | 0 | 0 | 0 | 0 | 2 |
[15] † Annemys sp Last Modified in 03/14/23 | 0 | 43 | 31 | 0 | 0 | 0 | 0 | 2 |
[16] † Xinjiangchelys wusu Last Modified in 03/14/23 | 0 | 34 | 31 | 0 | 0 | 0 | 0 | 2 |
[17] † Brodiechelys brodiei Last Modified in 03/14/23 | 2 | 32 | 29 | 0 | 0 | 0 | 0 | 2 |
[18] † Brodiechelys royoi Last Modified in 03/14/23 | 4 | 27 | 27 | 0 | 0 | 0 | 0 | 2 |
[19] † Xinjiangchelys radiplicatoides Last Modified in 03/14/23 | 2 | 30 | 29 | 0 | 0 | 0 | 0 | 2 |
[20] † Xinjiangchelys chowi Last Modified in 03/14/23 | 14 | 18 | 17 | 0 | 0 | 0 | 0 | 2 |
[21] † Xinjiangchelys qiguensis Last Modified in 03/14/23 | 3 | 28 | 28 | 0 | 0 | 0 | 0 | 2 |
[22] † Phunoichelys Last Modified in 03/14/23 | 10 | 21 | 21 | 0 | 0 | 0 | 0 | 2 |
[23] † Kalasinemys Last Modified in 03/14/23 | 14 | 17 | 17 | 0 | 0 | 0 | 0 | 2 |
[24] † Shartegemys Last Modified in 03/14/23 | 4 | 27 | 27 | 0 | 0 | 0 | 0 | 2 |
[25] † Yanduchelys cf Last Modified in 03/14/23 | 15 | 16 | 16 | 0 | 0 | 0 | 0 | 2 |
[26] † Larachelus morla Last Modified in 03/14/23 | 9 | 22 | 22 | 0 | 0 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 3167 | |
Documents list | 151 | |
Project overview | 275 | |
Media views | 684 | Media search (578 views); M851977 (106 views); |
Matrix views | 185 | Matrix landing page (173 views); Analysis 3 (12 views); |
Bibliography | 136 | |
Taxon list | 1178 | |
Views for media list | 212 | |
Specimen list | 346 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 2 | |
Project downloads | 1 | |
Matrix downloads | 1 | Analysis 3 (1 download); |