Morphobank

Project 4550: M. D. Ezcurra, H. Sues. 2021. A re-assessment of the osteology and phylogenetic relationships of the enigmatic, large-headed reptile Sphodrosaurus pennsylvanicus (Late Triassic, Pennsylvania, USA) indicates archosauriform affinities. Journal of Systematic Palaeontology. 19 (24):1643-1677.

Specimen: † Heterodontosaurus tucki (unvouchered)
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Abstract

Most Triassic terrestrial diapsids belong to two clades, Lepidosauromorpha or (the more diverse) Archosauromorpha. Nevertheless, the phylogenetic relationships of some Triassic diapsids have remained uncertain for decades because of the lack of preservation of phylogenetically relevant anatomical regions or because of unusual combinations of features. One of these enigmatic forms is the small-sized Sphodrosaurus pennsylvanicus from the Upper Triassic Hammer Creek Formation of the Newark Supergroup in Pennsylvania (USA). It was first identified as a procolophonid parareptile, later as a probable rhynchosaur archosauromorph, and more recently as an indeterminate neodiapsid. Here we revise the anatomy of Sphodrosaurus pennsylvanicus in order to include it for the first time in a quantitative phylogenetic analysis, which is focused on Permo–Triassic neodiapsids. Sphodrosaurus pennsylvanicus is recovered in this analysis as a doswelliid proterochampsian within Archosauromorpha. As a result, this taxon is added to the list of doswelliids known from the Carnian–Norian of the eastern and south-western USA. Previous authors recognized that the most unusual feature of Sphodrosaurus pennsylvanicus is its proportionally very large skull. Phylogenetic generalized least squares regressions confirmed that Sphodrosaurus pennsylvanicus has a larger skull than the vast majority of Permo–Triassic diapsids. Optimization in the phylogeny of the skull width to presacral length ratio shows the most likely scenario is that the extremely broad skull of Sphodrosaurus pennsylvanicus is autapomorphic, but it is not unique among archosauromorphs, being paralleled by hyperodapedontine rhynchosaurs and the proterochampsian Proterochampsa barrionuevoi. Exploration of a morphospace of linear measurements shows that Sphodrosaurus pennsylvanicus shares strong similarities with the probably semi-aquatic Proterochampsa barrionuevoi, suggesting that the former species may have had a similar mode of life. A linear discriminant analysis of ungual functional categories found that the only preserved ungual of Sphodrosaurus pennsylvanicus was suitable for digging or some other way of substrate processing.

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Article DOI: 10.1080/14772019.2022.2057820

Project DOI: 10.7934/P4550, http://dx.doi.org/10.7934/P4550

This project contains	Matrices
1 Media 1 Matrix 1 Document 230 Taxa 1 Specimen 889 Characters Total size of project's media files: 30.66k Download Project SDD File	Total scored cells: 82712 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 889 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 2098 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:745/144

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MorphoBank Project 4550

Creation Date:
27 January 2023
Publication Date:
31 January 2023
Project views: 4840
Matrix downloads: 3

Authors' Institutions

National Museum of Natural History, Smithsonian Institution

Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)

Museo Argentino de Ciencias Naturales Bernardino Rivadavia

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Taxonomic Overview for Matrix 'M29138' (230 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Petrolacosaurus kansensis Last Modified in 01/31/23	113	675	656	0	104	0	0	1
[2] † Orovenator mayorum Last Modified in 01/31/23	541	296	293	0	55	0	0	1
[3] † Acerosodontosaurus piveteaui Last Modified in 01/31/23	565	264	253	0	64	0	0	1
[4] † Hovasaurus boulei Last Modified in 01/31/23	378	469	456	0	55	0	0	1
[5] † Youngina capensis Last Modified in 01/31/23	100	705	655	0	91	0	0	1
[6] † Claudiosaurus germaini Last Modified in 01/31/23	239	565	554	0	95	0	0	1
[7] † Elachistosuchus huenei Last Modified in 01/31/23	652	227	216	0	20	0	0	1
[8] † Feralisaurus corami Last Modified in 01/31/23	719	141	137	0	33	0	0	1
[9] † Paliguana whitei Last Modified in 01/31/23	665	191	183	0	40	0	0	1
[10] † Fraxinisaura rozynekae Last Modified in 01/31/23	618	238	233	0	38	0	0	1
[11] † Marmoretta oxoniensis Last Modified in 01/31/23	587	265	259	0	43	0	0	1
[12] † Huehuecuetzpalli mixtecus Last Modified in 01/31/23	370	429	425	0	94	0	0	1
[13] † Megachirella wachtleri Last Modified in 01/31/23	587	255	252	0	50	0	0	1
[14] † Salvator rufescens Last Modified in 01/31/23	19	746	742	0	127	0	0	1
[15] † Planocephalosaurus robinsonae Last Modified in 01/31/23	297	502	487	0	94	0	0	1
[16] † Gephyrosaurus bridensis Last Modified in 01/31/23	271	541	512	0	82	0	0	1
[17] † Clevosaurus hudsoni Last Modified in 01/31/23	106	673	669	0	114	0	0	1
[18] † Sphenodon punctatus Last Modified in 01/31/23	10	764	756	0	122	0	0	1
[19] † Colobops noviportensis Last Modified in 01/31/23	768	90	89	0	32	0	0	1
[20] † Cteniogenys sp Last Modified in 01/31/23	496	341	334	0	57	0	0	1
[21] † Simoedosaurus lemoinei Last Modified in 01/31/23	199	606	600	0	89	0	0	1
[22] † ISIR 1132 Last Modified in 01/31/23	872	14	14	0	3	0	0	1
[23] † Aenigmastropheus parringtoni Last Modified in 01/31/23	840	48	46	0	2	0	0	1
[24] † Protorosaurus speneri Last Modified in 01/31/23	248	578	526	0	78	0	0	1
[25] † Dinocephalosaurus orientalis Last Modified in 01/31/23	439	354	349	0	100	0	0	1
[26] † Amotosaurus rotfeldensis Last Modified in 01/31/23	464	373	361	0	62	0	0	1
[27] † Macrocnemus bassanii Last Modified in 01/31/23	159	644	606	0	96	0	0	1
[28] † Macrocnemus obristi Last Modified in 01/31/23	783	87	85	0	21	0	0	1
[29] † Macrocnemus fuyuanensis Last Modified in 01/31/23	364	455	444	0	81	0	0	1
[30] † Fuyuansaurus acutirostris Last Modified in 01/31/23	718	147	143	0	27	0	0	1
[31] † Pectodens zhenyuensis Last Modified in 01/31/23	543	283	274	0	72	0	0	1
[32] † Augustaburiania vatagini Last Modified in 01/31/23	799	84	83	0	7	0	0	1
[33] † Protanystropheus antiquus Last Modified in 01/31/23	866	24	22	0	1	0	0	1
[34] † Trachelosaurus fischeri Last Modified in 01/31/23	786	101	89	0	14	0	0	1
[35] † Tanystropheus spp Last Modified in 01/31/23	119	681	619	0	109	0	0	1
[36] † Tanystropheus haasi Last Modified in 01/31/23	877	11	11	0	1	0	0	1
[37] † Raibliania calligarisi Last Modified in 01/31/23	805	67	62	0	22	0	0	1
[38] † Langobardisaurus pandolfii Last Modified in 01/31/23	523	311	295	0	70	0	0	1
[39] † Tanytrachelos ahynis Last Modified in 01/31/23	492	358	334	0	63	0	0	1
[40] † Jesairosaurus lehmani Last Modified in 01/31/23	444	375	358	0	81	0	0	1
[41] † UFRGS pv 492 T Last Modified in 01/31/23	873	16	15	0	1	0	0	1
[42] † Elessaurus gondwanoccidens Last Modified in 01/31/23	788	100	96	0	3	0	0	1
[43] † Pamelaria dolichotrachela Last Modified in 01/31/23	209	622	616	0	64	0	0	1
[44] † Azendohsaurus madagaskarensis Last Modified in 01/31/23	97	728	715	0	72	0	0	1
[45] † Azendohsaurus laaroussii Last Modified in 01/31/23	809	71	69	0	11	0	0	1
[46] † Shringasaurus indicus Last Modified in 01/31/23	408	435	426	0	55	0	0	1
[47] † Malerisaurus robinsonae Last Modified in 01/31/23	537	333	318	0	34	0	0	1
[48] † Malerisaurus langstoni Last Modified in 01/31/23	680	203	200	0	9	0	0	1
[49] † Malerisaurus all NA Last Modified in 01/31/23	426	441	417	0	45	0	0	1
[50] † Trilophosaurus buettneri Last Modified in 01/31/23	111	695	658	0	93	0	0	1
[51] † Spinosuchus caseanus Last Modified in 01/31/23	457	370	365	0	67	0	0	1
[52] † Teraterpeton hrynewichorum Last Modified in 01/31/23	420	397	391	0	78	0	0	1
[53] † Arctosaurus osborni Last Modified in 01/31/23	875	13	13	0	1	0	0	1
[54] † Eifelosaurus triadicus Last Modified in 01/31/23	840	45	41	0	6	0	0	1
[55] † Noteosuchus colletti Last Modified in 01/31/23	653	218	206	0	21	0	0	1
[56] † Mesosuchus browni Last Modified in 01/31/23	100	716	682	0	78	0	0	1
[57] † Howesia browni Last Modified in 01/31/23	435	417	389	0	44	0	0	1
[58] † Eohyosaurus wolvaardti Last Modified in 01/31/23	702	172	165	0	21	0	0	1
[59] † Rhynchosaurus articeps Last Modified in 01/31/23	235	578	560	0	82	0	0	1
[60] † Bentonyx sidensis Last Modified in 01/31/23	582	253	251	0	56	0	0	1
[61] † Mesodapedon kuttyi Last Modified in 01/31/23	839	39	39	0	11	0	0	1
[62] † Ammorhynchus navajoi Last Modified in 01/31/23	818	47	46	0	25	0	0	1
[63] † Stenaulorhynchus stockleyi Last Modified in 01/31/23	161	646	644	0	84	0	0	1
[64] † Brasinorhynchus mariantensis Last Modified in 01/31/23	536	301	299	0	54	0	0	1
[65] † CRILAR pv 461 Last Modified in 01/31/23	873	13	11	0	5	0	0	1
[66] † CRILAR pv 462 Last Modified in 01/31/23	873	12	11	0	5	0	0	1
[67] † Elorhynchus carrolli Last Modified in 01/31/23	828	50	47	0	14	0	0	1
[68] † CRILAR pv 497 Last Modified in 01/31/23	855	30	27	0	7	0	0	1
[69] † Chanares rhynchosaur Last Modified in 01/31/23	805	71	67	0	17	0	0	1
[70] † Langeronyx brodiei Last Modified in 01/31/23	691	155	154	0	44	0	0	1
[71] † Fodonyx spenceri Last Modified in 01/31/23	708	158	157	0	24	0	0	1
[72] † Isalorhynchus genovefae Last Modified in 01/31/23	250	567	561	0	76	0	0	1
[73] † Hyperodapedon sanjuanensis Last Modified in 01/31/23	459	374	368	0	61	0	0	1
[74] † Hyperodapedon mariensis Last Modified in 01/31/23	431	387	382	0	76	0	0	1
[75] † Hyperodapedon huenei Last Modified in 01/31/23	506	325	324	0	59	0	0	1
[76] † PVSJ 2728 Last Modified in 01/31/23	856	22	22	0	11	0	0	1
[77] † Hyperodapedon gordoni Last Modified in 01/31/23	146	655	648	0	95	0	0	1
[78] † Hyperodapedon huxleyi Last Modified in 01/31/23	142	652	650	0	97	0	0	1
[79] † Teyumbaita sulcognathus Last Modified in 01/31/23	282	530	524	0	83	0	0	1
[80] † Eorasaurus olsoni Last Modified in 01/31/23	859	30	27	0	2	0	0	1
[81] † Prolacertoides jimusarensis Last Modified in 01/31/23	783	76	74	0	31	0	0	1
[82] † Prolacerta broomi Last Modified in 01/31/23	44	789	740	0	72	0	0	1
[83] † K australiensis combined Last Modified in 01/31/23	746	116	111	0	32	0	0	1
[84] † Boreopricea funerea Last Modified in 01/31/23	494	353	326	0	51	0	0	1
[85] † Teyujagua paradoxa Last Modified in 01/31/23	506	320	319	0	63	0	0	1
[86] † Antarctanax shackletoni Last Modified in 01/31/23	810	74	68	0	7	0	0	1
[87] † Archosaurus rossicus holotype Last Modified in 01/31/23	865	24	23	0	1	0	0	1
[88] † Archosaurus complete Last Modified in 01/31/23	781	111	106	0	2	0	0	1
[89] † Proterosuchus fergusi Last Modified in 01/31/23	188	675	658	0	41	0	0	1
[90] † Proterosuchus goweri Last Modified in 01/31/23	578	299	294	0	17	0	0	1
[91] † Proterosuchus alexanderi Last Modified in 01/31/23	268	580	574	0	46	0	0	1
[92] † FC dpv 2641 Last Modified in 01/31/23	881	7	7	0	1	0	0	1
[93] † "Chasmatosaurus" yuani Last Modified in 01/31/23	211	639	624	0	53	0	0	1
[94] † UFSM 11444 Last Modified in 01/31/23	869	19	19	0	1	0	0	1
[95] † UFSM 11394 Last Modified in 01/31/23	860	29	27	0	1	0	0	1
[96] † Tasmaniosaurus triassicus Last Modified in 01/31/23	746	138	129	0	14	0	0	1
[97] † Vonhuenia fredericki Last Modified in 01/31/23	868	19	19	0	2	0	0	1
[98] † C rossicus combined Last Modified in 01/31/23	836	54	52	0	1	0	0	1
[99] † C magnus combined Last Modified in 01/31/23	855	35	33	0	1	0	0	1
[100] † Chasmatosuchus vjushkovi Last Modified in 01/31/23	860	31	28	0	1	0	0	1
[101] † Koilamasuchus gonzalezdiazi Last Modified in 01/31/23	852	35	33	0	4	0	0	1
[102] † Kalisuchus rewanensis holotype Last Modified in 01/31/23	834	48	43	0	12	0	0	1
[103] † NMQR 3570 Last Modified in 01/31/23	843	35	35	0	11	0	0	1
[104] † Fugusuchus hejiapanensis Last Modified in 01/31/23	630	240	237	0	21	0	0	1
[105] † Sarmatosuchus otschevi Last Modified in 01/31/23	599	289	279	0	11	0	0	1
[106] † Guchengosuchus shiguaiensis Last Modified in 01/31/23	655	218	208	0	25	0	0	1
[107] † CRILAR pv 499 Last Modified in 01/31/23	882	6	6	0	1	0	0	1
[108] † Cuyosuchus huenei Last Modified in 01/31/23	665	224	211	0	13	0	0	1
[109] † Garjainia prima Last Modified in 01/31/23	127	731	715	0	42	0	0	1
[110] † Garjainia madiba combined Last Modified in 01/31/23	470	413	391	0	22	0	0	1
[111] † Erythrosuchus africanus Last Modified in 01/31/23	113	732	718	0	50	0	0	1
[112] † Shansisuchus shansisuchus Last Modified in 01/31/23	235	614	603	0	49	0	0	1
[113] † Shansisuchus kuyeheensis Last Modified in 01/31/23	784	100	94	0	11	0	0	1
[114] † Chalishevia cothurnata Last Modified in 01/31/23	797	84	81	0	11	0	0	1
[115] † Bharitalasuchus tapani Last Modified in 01/31/23	770	120	109	0	7	0	0	1
[116] † Youngosuchus sinensis Last Modified in 01/31/23	490	383	377	0	22	0	0	1
[117] † Uralosaurus combined Last Modified in 01/31/23	847	37	35	0	7	0	0	1
[118] † Vancleavea campi Last Modified in 01/31/23	291	528	521	0	75	0	0	1
[119] † Litorosuchus somnii Last Modified in 01/31/23	380	482	475	0	34	0	0	1
[120] † Asperoris mnyama Last Modified in 01/31/23	758	127	108	0	17	0	0	1
[121] † Euparkeria capensis Last Modified in 01/31/23	53	810	798	0	38	0	0	1
[122] † Dorosuchus neoetus Last Modified in 01/31/23	753	132	132	0	4	0	0	1
[123] † Halazhaisuchus qiaoensis Last Modified in 01/31/23	775	106	104	0	10	0	0	1
[124] † Osmolskina czatkoviensis Last Modified in 01/31/23	858	24	24	0	7	0	0	1
[125] † Osmolskina complete Last Modified in 01/31/23	364	507	496	0	29	0	0	1
[126] † Polymorphodon adorfi Last Modified in 01/31/23	651	226	219	0	17	0	0	1
[127] † Yarasuchus deccanensis Last Modified in 01/31/23	637	247	240	0	11	0	0	1
[128] † Dongusuchus efremovi Last Modified in 01/31/23	845	41	41	0	3	0	0	1
[129] † Teleocrater rhadinus Last Modified in 01/31/23	531	342	339	0	19	0	0	1
[130] † Spondylosoma absconditum Last Modified in 01/31/23	801	87	85	0	3	0	0	1
[131] † Proterochampsa barrionuevoi Last Modified in 01/31/23	265	580	565	0	59	0	0	1
[132] † Proterochampsa nodosa Last Modified in 01/31/23	618	250	242	0	24	0	0	1
[133] † Tropidosuchus romeri Last Modified in 01/31/23	198	657	640	0	44	0	0	1
[134] † Cerritosaurus binsfeldi Last Modified in 01/31/23	619	248	243	0	27	0	0	1
[135] † Pseudochampsa ischigualastensis Last Modified in 01/31/23	321	533	519	0	44	0	0	1
[136] † Rhadinosuchus gracilis Last Modified in 01/31/23	751	132	125	0	13	0	0	1
[137] † Gualosuchus reigi Last Modified in 01/31/23	288	562	551	0	48	0	0	1
[138] † Chanaresuchus bonapartei Last Modified in 01/31/23	93	760	738	0	54	0	0	1
[139] † Archeopelta arborensis Last Modified in 01/31/23	762	121	121	0	6	0	0	1
[140] † Tarjadia ruthae Last Modified in 01/31/23	331	532	527	0	31	0	0	1
[141] † Pagosvenator candelariensis Last Modified in 01/31/23	706	182	180	0	3	0	0	1
[142] † Erpetosuchus granti Last Modified in 01/31/23	457	409	396	0	27	0	0	1
[143] † Erpetosuchus sp Last Modified in 01/31/23	761	118	117	0	11	0	0	1
[144] † Parringtonia gracilis Last Modified in 01/31/23	383	485	477	0	29	0	0	1
[145] † Dyoplax arenaceus Last Modified in 01/31/23	825	63	61	0	3	0	0	1
[146] † Jaxtasuchus salomoni Last Modified in 01/31/23	652	224	217	0	20	0	0	1
[147] † Doswellia kaltenbachi Last Modified in 01/31/23	435	398	391	0	63	0	0	1
[148] † Rugarhynchos sixmilensis Last Modified in 01/31/23	663	180	175	0	51	0	0	1
[149] † Sphodrosaurus pennsylvanicus Last Modified in 01/31/23	772	121	96	0	11	0	0	1
[150] † Triopticus primus Last Modified in 01/31/23	779	107	98	0	7	0	0	1
[151] † Parasuchus angustifrons Last Modified in 01/31/23	597	262	259	0	33	0	0	1
[152] † Parasuchus hislopi Last Modified in 01/31/23	125	731	720	0	42	0	0	1
[153] † Nicrosaurus kapffi Last Modified in 01/31/23	288	562	553	0	47	0	0	1
[154] † Smilosuchus spp Last Modified in 01/31/23	151	692	677	0	55	0	0	1
[155] † Diandongosuchus fuyuanensis Last Modified in 01/31/23	374	496	491	0	24	0	0	1
[156] † Wannia scurriensis Last Modified in 01/31/23	671	194	181	0	32	0	0	1
[157] † Ebrachosuchus neukami Last Modified in 01/31/23	556	301	297	0	35	0	0	1
[158] † Paleorhinus sawini Last Modified in 01/31/23	595	269	265	0	29	0	0	1
[159] † Angistorhinus talainti Last Modified in 01/31/23	517	338	335	0	37	0	0	1
[160] † Mystriosuchus planirostris Last Modified in 01/31/23	343	522	502	0	44	0	0	1
[161] † Machaeroprosopus pristinus Last Modified in 01/31/23	394	468	448	0	47	0	0	1
[162] † Redondasaurus spp Last Modified in 01/31/23	525	337	325	0	39	0	0	1
[163] † Ornithosuchus woodwardi Last Modified in 01/31/23	286	573	564	0	35	0	0	1
[164] † Riojasuchus tenuisceps Last Modified in 01/31/23	146	707	694	0	43	0	0	1
[165] † Venaticosuchus rusconii Last Modified in 01/31/23	685	188	185	0	19	0	0	1
[166] † Dynamosuchus collisensis Last Modified in 01/31/23	586	302	299	0	4	0	0	1
[167] † Nundasuchus songeaensis Last Modified in 01/31/23	578	303	297	0	14	0	0	1
[168] † Turfanosuchus dabanensis Last Modified in 01/31/23	288	579	569	0	30	0	0	1
[169] † Gracilisuchus stipanicicorum Last Modified in 01/31/23	169	691	677	0	43	0	0	1
[170] † Yonghesuchus sangbiensis Last Modified in 01/31/23	662	209	205	0	22	0	0	1
[171] † Otter sandstone archosaur Last Modified in 01/31/23	835	57	51	0	3	0	0	1
[172] † Revueltosaurus callenderi Last Modified in 01/31/23	151	705	698	0	40	0	0	1
[173] † Aetosauroides scagliai Last Modified in 01/31/23	132	729	715	0	41	0	0	1
[174] † Aetosaurus ferratus Last Modified in 01/31/23	252	607	600	0	37	0	0	1
[175] † Stagonolepis robertsoni Last Modified in 01/31/23	112	738	736	0	41	0	0	1
[176] † Ticinosuchus ferox Last Modified in 01/31/23	535	338	330	0	23	0	0	1
[177] † Prestosuchus nyassicus Last Modified in 01/31/23	707	182	174	0	8	0	0	1
[178] † Dagasuchus santacruzensis Last Modified in 01/31/23	854	33	33	0	2	0	0	1
[179] † Qianosuchus mixtus Last Modified in 01/31/23	352	506	504	0	33	0	0	1
[180] † Arizonasaurus babbitti Last Modified in 01/31/23	413	440	435	0	41	0	0	1
[181] † Lotosaurus adentus Last Modified in 01/31/23	356	459	450	0	83	0	0	1
[182] † Ctenosauriscus koeneni Last Modified in 01/31/23	836	36	36	0	17	0	0	1
[183] † Hypselorhachis mirabilis Last Modified in 01/31/23	862	26	25	0	2	0	0	1
[184] † Waldhaus poposauroid Last Modified in 01/31/23	849	41	39	0	1	0	0	1
[185] † Vytshegdosuchus zbeshartensis Last Modified in 01/31/23	875	13	13	0	1	0	0	1
[186] † Bystrowisuchus flerovi Last Modified in 01/31/23	868	21	19	0	2	0	0	1
[187] † Bromsgroveia walkeri Last Modified in 01/31/23	844	44	44	0	1	0	0	1
[188] † Xilousuchus sapingensis Last Modified in 01/31/23	633	239	237	0	18	0	0	1
[189] † Moenkopi poposauroid Last Modified in 01/31/23	838	48	44	0	7	0	0	1
[190] † Mandasuchus tanyauchen Last Modified in 01/31/23	572	304	297	0	20	0	0	1
[191] † Luperosuchus fractus Last Modified in 01/31/23	756	131	129	0	4	0	0	1
[192] † Decuriasuchus quartacolonia Last Modified in 01/31/23	410	450	444	0	35	0	0	1
[193] † Batrachotomus kupferzellensis Last Modified in 01/31/23	191	668	658	0	36	0	0	1
[194] † Prestosuchus chiniquensis Last Modified in 01/31/23	97	757	752	0	40	0	0	1
[195] † Scleromochlus taylori Last Modified in 01/31/23	687	183	171	0	31	0	0	1
[196] † Austriadraco dallavecchiai Last Modified in 01/31/23	697	196	180	0	12	0	0	1
[197] † Preondactylus buffarinii Last Modified in 01/31/23	615	237	228	0	44	0	0	1
[198] † Austriadactylus cristatus Last Modified in 01/31/23	640	223	218	0	31	0	0	1
[199] † Seazzadactylus venieri Last Modified in 01/31/23	612	240	231	0	46	0	0	1
[200] † Raeticodactylus filisurensis Last Modified in 01/31/23	628	240	232	0	29	0	0	1
[201] † Carniadactylus rosenfeldi Last Modified in 01/31/23	526	302	296	0	67	0	0	1
[202] † Peteinosaurus zambellii Last Modified in 01/31/23	631	225	219	0	39	0	0	1
[203] † Caelestiventus hanseni Last Modified in 01/31/23	699	159	153	0	37	0	0	1
[204] † Dimorphodon macronyx Last Modified in 01/31/23	387	435	431	0	71	0	0	1
[205] † Eudimorphodon ranzii Last Modified in 01/31/23	521	327	324	0	44	0	0	1
[206] † Rhamphorhynchus muensteri Last Modified in 01/31/23	196	571	563	0	130	0	0	1
[207] † Cacibupteryx caribensis Last Modified in 01/31/23	717	147	143	0	27	0	0	1
[208] † Allkaruen koi Last Modified in 01/31/23	751	128	126	0	12	0	0	1
[209] † Lagerpeton chanarensis Last Modified in 01/31/23	577	279	273	0	37	0	0	1
[210] † Ixalerpeton polesinensis Last Modified in 01/31/23	490	368	362	0	37	0	0	1
[211] † Kongonaphon kely Last Modified in 01/31/23	803	77	73	0	13	0	0	1
[212] † PVSJ 883 Last Modified in 01/31/23	878	10	10	0	1	0	0	1
[213] † Dromomeron romeri Last Modified in 01/31/23	711	155	147	0	31	0	0	1
[214] † Dromomeron gregorii Last Modified in 01/31/23	744	140	140	0	5	0	0	1
[215] † Dromomeron gigas Last Modified in 01/31/23	851	32	32	0	6	0	0	1
[216] † Lagosuchus talampayensis Last Modified in 01/31/23	445	413	406	0	38	0	0	1
[217] † Lutungutali sitwensis Last Modified in 01/31/23	847	44	39	0	3	0	0	1
[218] † Asilisaurus kongwe Last Modified in 01/31/23	340	520	511	0	38	0	0	1
[219] † Nyasasaurus parringtoni Last Modified in 01/31/23	835	51	51	0	3	0	0	1
[220] † Lewisuchus admixtus Last Modified in 01/31/23	306	560	547	0	31	0	0	1
[221] † Silesaurus opolensis Last Modified in 01/31/23	188	661	650	0	50	0	0	1
[222] † Heterodontosaurus tucki Last Modified in 01/31/23	86	718	711	0	89	0	0	1
[223] † Lesothosaurus diagnosticus Last Modified in 01/31/23	110	703	696	0	82	0	0	1
[224] † Herrerasaurus ischigualastensis Last Modified in 01/31/23	81	764	750	0	58	0	0	1
[225] † Eoraptor lunensis Last Modified in 01/31/23	147	697	689	0	52	0	0	1
[226] † Buriolestes schultzi Last Modified in 01/31/23	116	721	714	0	57	0	0	1
[227] † Saturnalia tupiniquim Last Modified in 01/31/23	328	524	512	0	48	0	0	1
[228] † Tawa hallae Last Modified in 01/31/23	239	593	593	0	57	0	0	1
[229] † Coelophysis bauri Last Modified in 01/31/23	80	740	732	0	77	0	0	1
[230] † Megapnosaurus rhodesiensis Last Modified in 01/31/23	89	728	720	0	80	0	0	1

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	4840
Project overview	364
Taxon list	2655
Media views	710	Media search (608 views); M851588 (102 views);
Specimen list	313
Documents list	160
Matrix views	197	Matrix landing page (181 views); Data matrix from supplemental materials (16 views);
Views for media list	298
Bibliography	143

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type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	4
Matrix downloads	3	Data matrix from supplemental materials (3 downloads);
Project downloads	1