Morphobank

Project 4550: M. D. Ezcurra, H. Sues. 2021. A re-assessment of the osteology and phylogenetic relationships of the enigmatic, large-headed reptile Sphodrosaurus pennsylvanicus (Late Triassic, Pennsylvania, USA) indicates archosauriform affinities. Journal of Systematic Palaeontology. 19 (24):1643-1677.

Specimen: † Heterodontosaurus tucki (unvouchered)
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Abstract

Most Triassic terrestrial diapsids belong to two clades, Lepidosauromorpha or (the more diverse) Archosauromorpha. Nevertheless, the phylogenetic relationships of some Triassic diapsids have remained uncertain for decades because of the lack of preservation of phylogenetically relevant anatomical regions or because of unusual combinations of features. One of these enigmatic forms is the small-sized Sphodrosaurus pennsylvanicus from the Upper Triassic Hammer Creek Formation of the Newark Supergroup in Pennsylvania (USA). It was first identified as a procolophonid parareptile, later as a probable rhynchosaur archosauromorph, and more recently as an indeterminate neodiapsid. Here we revise the anatomy of Sphodrosaurus pennsylvanicus in order to include it for the first time in a quantitative phylogenetic analysis, which is focused on Permo–Triassic neodiapsids. Sphodrosaurus pennsylvanicus is recovered in this analysis as a doswelliid proterochampsian within Archosauromorpha. As a result, this taxon is added to the list of doswelliids known from the Carnian–Norian of the eastern and south-western USA. Previous authors recognized that the most unusual feature of Sphodrosaurus pennsylvanicus is its proportionally very large skull. Phylogenetic generalized least squares regressions confirmed that Sphodrosaurus pennsylvanicus has a larger skull than the vast majority of Permo–Triassic diapsids. Optimization in the phylogeny of the skull width to presacral length ratio shows the most likely scenario is that the extremely broad skull of Sphodrosaurus pennsylvanicus is autapomorphic, but it is not unique among archosauromorphs, being paralleled by hyperodapedontine rhynchosaurs and the proterochampsian Proterochampsa barrionuevoi. Exploration of a morphospace of linear measurements shows that Sphodrosaurus pennsylvanicus shares strong similarities with the probably semi-aquatic Proterochampsa barrionuevoi, suggesting that the former species may have had a similar mode of life. A linear discriminant analysis of ungual functional categories found that the only preserved ungual of Sphodrosaurus pennsylvanicus was suitable for digging or some other way of substrate processing.

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Article DOI: 10.1080/14772019.2022.2057820

Project DOI: 10.7934/P4550, http://dx.doi.org/10.7934/P4550

This project contains	Matrices
1 Media 1 Matrix 1 Document 230 Taxa 1 Specimen 889 Characters Total size of project's media files: 30.66k Download Project SDD File	Total scored cells: 82712 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 889 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 2098 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:745/144

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MorphoBank Project 4550

Creation Date:
27 January 2023
Publication Date:
31 January 2023
Matrix downloads: 3

Authors' Institutions

National Museum of Natural History, Smithsonian Institution

Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)

Museo Argentino de Ciencias Naturales Bernardino Rivadavia

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Taxonomic Overview for Matrix 'M29138' (230 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Petrolacosaurus kansensis Taxon name last Modified on 01/31/23	113	675	656	0	104	0	0	1
[2] † Orovenator mayorum Taxon name last Modified on 01/31/23	541	296	293	0	55	0	0	1
[3] † Acerosodontosaurus piveteaui Taxon name last Modified on 01/31/23	565	264	253	0	64	0	0	1
[4] † Hovasaurus boulei Taxon name last Modified on 01/31/23	378	469	456	0	55	0	0	1
[5] † Youngina capensis Taxon name last Modified on 01/31/23	100	705	655	0	91	0	0	1
[6] † Claudiosaurus germaini Taxon name last Modified on 01/31/23	239	565	554	0	95	0	0	1
[7] † Elachistosuchus huenei Taxon name last Modified on 01/31/23	652	227	216	0	20	0	0	1
[8] † Feralisaurus corami Taxon name last Modified on 01/31/23	719	141	137	0	33	0	0	1
[9] † Paliguana whitei Taxon name last Modified on 01/31/23	665	191	183	0	40	0	0	1
[10] † Fraxinisaura rozynekae Taxon name last Modified on 01/31/23	618	238	233	0	38	0	0	1
[11] † Marmoretta oxoniensis Taxon name last Modified on 01/31/23	587	265	259	0	43	0	0	1
[12] † Huehuecuetzpalli mixtecus Taxon name last Modified on 01/31/23	370	429	425	0	94	0	0	1
[13] † Megachirella wachtleri Taxon name last Modified on 01/31/23	587	255	252	0	50	0	0	1
[14] † Salvator rufescens Taxon name last Modified on 01/31/23	19	746	742	0	127	0	0	1
[15] † Planocephalosaurus robinsonae Taxon name last Modified on 01/31/23	297	502	487	0	94	0	0	1
[16] † Gephyrosaurus bridensis Taxon name last Modified on 01/31/23	271	541	512	0	82	0	0	1
[17] † Clevosaurus hudsoni Taxon name last Modified on 01/31/23	106	673	669	0	114	0	0	1
[18] † Sphenodon punctatus Taxon name last Modified on 01/31/23	10	764	756	0	122	0	0	1
[19] † Colobops noviportensis Taxon name last Modified on 01/31/23	768	90	89	0	32	0	0	1
[20] † Cteniogenys sp Taxon name last Modified on 01/31/23	496	341	334	0	57	0	0	1
[21] † Simoedosaurus lemoinei Taxon name last Modified on 01/31/23	199	606	600	0	89	0	0	1
[22] † ISIR 1132 Taxon name last Modified on 01/31/23	872	14	14	0	3	0	0	1
[23] † Aenigmastropheus parringtoni Taxon name last Modified on 01/31/23	840	48	46	0	2	0	0	1
[24] † Protorosaurus speneri Taxon name last Modified on 01/31/23	248	578	526	0	78	0	0	1
[25] † Dinocephalosaurus orientalis Taxon name last Modified on 01/31/23	439	354	349	0	100	0	0	1
[26] † Amotosaurus rotfeldensis Taxon name last Modified on 01/31/23	464	373	361	0	62	0	0	1
[27] † Macrocnemus bassanii Taxon name last Modified on 01/31/23	159	644	606	0	96	0	0	1
[28] † Macrocnemus obristi Taxon name last Modified on 01/31/23	783	87	85	0	21	0	0	1
[29] † Macrocnemus fuyuanensis Taxon name last Modified on 01/31/23	364	455	444	0	81	0	0	1
[30] † Fuyuansaurus acutirostris Taxon name last Modified on 01/31/23	718	147	143	0	27	0	0	1
[31] † Pectodens zhenyuensis Taxon name last Modified on 01/31/23	543	283	274	0	72	0	0	1
[32] † Augustaburiania vatagini Taxon name last Modified on 01/31/23	799	84	83	0	7	0	0	1
[33] † Protanystropheus antiquus Taxon name last Modified on 01/31/23	866	24	22	0	1	0	0	1
[34] † Trachelosaurus fischeri Taxon name last Modified on 01/31/23	786	101	89	0	14	0	0	1
[35] † Tanystropheus spp Taxon name last Modified on 01/31/23	119	681	619	0	109	0	0	1
[36] † Tanystropheus haasi Taxon name last Modified on 01/31/23	877	11	11	0	1	0	0	1
[37] † Raibliania calligarisi Taxon name last Modified on 01/31/23	805	67	62	0	22	0	0	1
[38] † Langobardisaurus pandolfii Taxon name last Modified on 01/31/23	523	311	295	0	70	0	0	1
[39] † Tanytrachelos ahynis Taxon name last Modified on 01/31/23	492	358	334	0	63	0	0	1
[40] † Jesairosaurus lehmani Taxon name last Modified on 01/31/23	444	375	358	0	81	0	0	1
[41] † UFRGS pv 492 T Taxon name last Modified on 01/31/23	873	16	15	0	1	0	0	1
[42] † Elessaurus gondwanoccidens Taxon name last Modified on 01/31/23	788	100	96	0	3	0	0	1
[43] † Pamelaria dolichotrachela Taxon name last Modified on 01/31/23	209	622	616	0	64	0	0	1
[44] † Azendohsaurus madagaskarensis Taxon name last Modified on 01/31/23	97	728	715	0	72	0	0	1
[45] † Azendohsaurus laaroussii Taxon name last Modified on 01/31/23	809	71	69	0	11	0	0	1
[46] † Shringasaurus indicus Taxon name last Modified on 01/31/23	408	435	426	0	55	0	0	1
[47] † Malerisaurus robinsonae Taxon name last Modified on 01/31/23	537	333	318	0	34	0	0	1
[48] † Malerisaurus langstoni Taxon name last Modified on 01/31/23	680	203	200	0	9	0	0	1
[49] † Malerisaurus all NA Taxon name last Modified on 01/31/23	426	441	417	0	45	0	0	1
[50] † Trilophosaurus buettneri Taxon name last Modified on 01/31/23	111	695	658	0	93	0	0	1
[51] † Spinosuchus caseanus Taxon name last Modified on 01/31/23	457	370	365	0	67	0	0	1
[52] † Teraterpeton hrynewichorum Taxon name last Modified on 01/31/23	420	397	391	0	78	0	0	1
[53] † Arctosaurus osborni Taxon name last Modified on 01/31/23	875	13	13	0	1	0	0	1
[54] † Eifelosaurus triadicus Taxon name last Modified on 01/31/23	840	45	41	0	6	0	0	1
[55] † Noteosuchus colletti Taxon name last Modified on 01/31/23	653	218	206	0	21	0	0	1
[56] † Mesosuchus browni Taxon name last Modified on 01/31/23	100	716	682	0	78	0	0	1
[57] † Howesia browni Taxon name last Modified on 01/31/23	435	417	389	0	44	0	0	1
[58] † Eohyosaurus wolvaardti Taxon name last Modified on 01/31/23	702	172	165	0	21	0	0	1
[59] † Rhynchosaurus articeps Taxon name last Modified on 01/31/23	235	578	560	0	82	0	0	1
[60] † Bentonyx sidensis Taxon name last Modified on 01/31/23	582	253	251	0	56	0	0	1
[61] † Mesodapedon kuttyi Taxon name last Modified on 01/31/23	839	39	39	0	11	0	0	1
[62] † Ammorhynchus navajoi Taxon name last Modified on 01/31/23	818	47	46	0	25	0	0	1
[63] † Stenaulorhynchus stockleyi Taxon name last Modified on 01/31/23	161	646	644	0	84	0	0	1
[64] † Brasinorhynchus mariantensis Taxon name last Modified on 01/31/23	536	301	299	0	54	0	0	1
[65] † CRILAR pv 461 Taxon name last Modified on 01/31/23	873	13	11	0	5	0	0	1
[66] † CRILAR pv 462 Taxon name last Modified on 01/31/23	873	12	11	0	5	0	0	1
[67] † Elorhynchus carrolli Taxon name last Modified on 01/31/23	828	50	47	0	14	0	0	1
[68] † CRILAR pv 497 Taxon name last Modified on 01/31/23	855	30	27	0	7	0	0	1
[69] † Chanares rhynchosaur Taxon name last Modified on 01/31/23	805	71	67	0	17	0	0	1
[70] † Langeronyx brodiei Taxon name last Modified on 01/31/23	691	155	154	0	44	0	0	1
[71] † Fodonyx spenceri Taxon name last Modified on 01/31/23	708	158	157	0	24	0	0	1
[72] † Isalorhynchus genovefae Taxon name last Modified on 01/31/23	250	567	561	0	76	0	0	1
[73] † Hyperodapedon sanjuanensis Taxon name last Modified on 01/31/23	459	374	368	0	61	0	0	1
[74] † Hyperodapedon mariensis Taxon name last Modified on 01/31/23	431	387	382	0	76	0	0	1
[75] † Hyperodapedon huenei Taxon name last Modified on 01/31/23	506	325	324	0	59	0	0	1
[76] † PVSJ 2728 Taxon name last Modified on 01/31/23	856	22	22	0	11	0	0	1
[77] † Hyperodapedon gordoni Taxon name last Modified on 01/31/23	146	655	648	0	95	0	0	1
[78] † Hyperodapedon huxleyi Taxon name last Modified on 01/31/23	142	652	650	0	97	0	0	1
[79] † Teyumbaita sulcognathus Taxon name last Modified on 01/31/23	282	530	524	0	83	0	0	1
[80] † Eorasaurus olsoni Taxon name last Modified on 01/31/23	859	30	27	0	2	0	0	1
[81] † Prolacertoides jimusarensis Taxon name last Modified on 01/31/23	783	76	74	0	31	0	0	1
[82] † Prolacerta broomi Taxon name last Modified on 01/31/23	44	789	740	0	72	0	0	1
[83] † K australiensis combined Taxon name last Modified on 01/31/23	746	116	111	0	32	0	0	1
[84] † Boreopricea funerea Taxon name last Modified on 01/31/23	494	353	326	0	51	0	0	1
[85] † Teyujagua paradoxa Taxon name last Modified on 01/31/23	506	320	319	0	63	0	0	1
[86] † Antarctanax shackletoni Taxon name last Modified on 01/31/23	810	74	68	0	7	0	0	1
[87] † Archosaurus rossicus holotype Taxon name last Modified on 01/31/23	865	24	23	0	1	0	0	1
[88] † Archosaurus complete Taxon name last Modified on 01/31/23	781	111	106	0	2	0	0	1
[89] † Proterosuchus fergusi Taxon name last Modified on 01/31/23	188	675	658	0	41	0	0	1
[90] † Proterosuchus goweri Taxon name last Modified on 01/31/23	578	299	294	0	17	0	0	1
[91] † Proterosuchus alexanderi Taxon name last Modified on 01/31/23	268	580	574	0	46	0	0	1
[92] † FC dpv 2641 Taxon name last Modified on 01/31/23	881	7	7	0	1	0	0	1
[93] † "Chasmatosaurus" yuani Taxon name last Modified on 01/31/23	211	639	624	0	53	0	0	1
[94] † UFSM 11444 Taxon name last Modified on 01/31/23	869	19	19	0	1	0	0	1
[95] † UFSM 11394 Taxon name last Modified on 01/31/23	860	29	27	0	1	0	0	1
[96] † Tasmaniosaurus triassicus Taxon name last Modified on 01/31/23	746	138	129	0	14	0	0	1
[97] † Vonhuenia fredericki Taxon name last Modified on 01/31/23	868	19	19	0	2	0	0	1
[98] † C rossicus combined Taxon name last Modified on 01/31/23	836	54	52	0	1	0	0	1
[99] † C magnus combined Taxon name last Modified on 01/31/23	855	35	33	0	1	0	0	1
[100] † Chasmatosuchus vjushkovi Taxon name last Modified on 01/31/23	860	31	28	0	1	0	0	1
[101] † Koilamasuchus gonzalezdiazi Taxon name last Modified on 01/31/23	852	35	33	0	4	0	0	1
[102] † Kalisuchus rewanensis holotype Taxon name last Modified on 01/31/23	834	48	43	0	12	0	0	1
[103] † NMQR 3570 Taxon name last Modified on 01/31/23	843	35	35	0	11	0	0	1
[104] † Fugusuchus hejiapanensis Taxon name last Modified on 01/31/23	630	240	237	0	21	0	0	1
[105] † Sarmatosuchus otschevi Taxon name last Modified on 01/31/23	599	289	279	0	11	0	0	1
[106] † Guchengosuchus shiguaiensis Taxon name last Modified on 01/31/23	655	218	208	0	25	0	0	1
[107] † CRILAR pv 499 Taxon name last Modified on 01/31/23	882	6	6	0	1	0	0	1
[108] † Cuyosuchus huenei Taxon name last Modified on 01/31/23	665	224	211	0	13	0	0	1
[109] † Garjainia prima Taxon name last Modified on 01/31/23	127	731	715	0	42	0	0	1
[110] † Garjainia madiba combined Taxon name last Modified on 01/31/23	470	413	391	0	22	0	0	1
[111] † Erythrosuchus africanus Taxon name last Modified on 01/31/23	113	732	718	0	50	0	0	1
[112] † Shansisuchus shansisuchus Taxon name last Modified on 01/31/23	235	614	603	0	49	0	0	1
[113] † Shansisuchus kuyeheensis Taxon name last Modified on 01/31/23	784	100	94	0	11	0	0	1
[114] † Chalishevia cothurnata Taxon name last Modified on 01/31/23	797	84	81	0	11	0	0	1
[115] † Bharitalasuchus tapani Taxon name last Modified on 01/31/23	770	120	109	0	7	0	0	1
[116] † Youngosuchus sinensis Taxon name last Modified on 01/31/23	490	383	377	0	22	0	0	1
[117] † Uralosaurus combined Taxon name last Modified on 01/31/23	847	37	35	0	7	0	0	1
[118] † Vancleavea campi Taxon name last Modified on 01/31/23	291	528	521	0	75	0	0	1
[119] † Litorosuchus somnii Taxon name last Modified on 01/31/23	380	482	475	0	34	0	0	1
[120] † Asperoris mnyama Taxon name last Modified on 01/31/23	758	127	108	0	17	0	0	1
[121] † Euparkeria capensis Taxon name last Modified on 01/31/23	53	810	798	0	38	0	0	1
[122] † Dorosuchus neoetus Taxon name last Modified on 01/31/23	753	132	132	0	4	0	0	1
[123] † Halazhaisuchus qiaoensis Taxon name last Modified on 01/31/23	775	106	104	0	10	0	0	1
[124] † Osmolskina czatkoviensis Taxon name last Modified on 01/31/23	858	24	24	0	7	0	0	1
[125] † Osmolskina complete Taxon name last Modified on 01/31/23	364	507	496	0	29	0	0	1
[126] † Polymorphodon adorfi Taxon name last Modified on 01/31/23	651	226	219	0	17	0	0	1
[127] † Yarasuchus deccanensis Taxon name last Modified on 01/31/23	637	247	240	0	11	0	0	1
[128] † Dongusuchus efremovi Taxon name last Modified on 01/31/23	845	41	41	0	3	0	0	1
[129] † Teleocrater rhadinus Taxon name last Modified on 01/31/23	531	342	339	0	19	0	0	1
[130] † Spondylosoma absconditum Taxon name last Modified on 01/31/23	801	87	85	0	3	0	0	1
[131] † Proterochampsa barrionuevoi Taxon name last Modified on 01/31/23	265	580	565	0	59	0	0	1
[132] † Proterochampsa nodosa Taxon name last Modified on 01/31/23	618	250	242	0	24	0	0	1
[133] † Tropidosuchus romeri Taxon name last Modified on 01/31/23	198	657	640	0	44	0	0	1
[134] † Cerritosaurus binsfeldi Taxon name last Modified on 01/31/23	619	248	243	0	27	0	0	1
[135] † Pseudochampsa ischigualastensis Taxon name last Modified on 01/31/23	321	533	519	0	44	0	0	1
[136] † Rhadinosuchus gracilis Taxon name last Modified on 01/31/23	751	132	125	0	13	0	0	1
[137] † Gualosuchus reigi Taxon name last Modified on 01/31/23	288	562	551	0	48	0	0	1
[138] † Chanaresuchus bonapartei Taxon name last Modified on 01/31/23	93	760	738	0	54	0	0	1
[139] † Archeopelta arborensis Taxon name last Modified on 01/31/23	762	121	121	0	6	0	0	1
[140] † Tarjadia ruthae Taxon name last Modified on 01/31/23	331	532	527	0	31	0	0	1
[141] † Pagosvenator candelariensis Taxon name last Modified on 01/31/23	706	182	180	0	3	0	0	1
[142] † Erpetosuchus granti Taxon name last Modified on 01/31/23	457	409	396	0	27	0	0	1
[143] † Erpetosuchus sp Taxon name last Modified on 01/31/23	761	118	117	0	11	0	0	1
[144] † Parringtonia gracilis Taxon name last Modified on 01/31/23	383	485	477	0	29	0	0	1
[145] † Dyoplax arenaceus Taxon name last Modified on 01/31/23	825	63	61	0	3	0	0	1
[146] † Jaxtasuchus salomoni Taxon name last Modified on 01/31/23	652	224	217	0	20	0	0	1
[147] † Doswellia kaltenbachi Taxon name last Modified on 01/31/23	435	398	391	0	63	0	0	1
[148] † Rugarhynchos sixmilensis Taxon name last Modified on 01/31/23	663	180	175	0	51	0	0	1
[149] † Sphodrosaurus pennsylvanicus Taxon name last Modified on 01/31/23	772	121	96	0	11	0	0	1
[150] † Triopticus primus Taxon name last Modified on 01/31/23	779	107	98	0	7	0	0	1
[151] † Parasuchus angustifrons Taxon name last Modified on 01/31/23	597	262	259	0	33	0	0	1
[152] † Parasuchus hislopi Taxon name last Modified on 01/31/23	125	731	720	0	42	0	0	1
[153] † Nicrosaurus kapffi Taxon name last Modified on 01/31/23	288	562	553	0	47	0	0	1
[154] † Smilosuchus spp Taxon name last Modified on 01/31/23	151	692	677	0	55	0	0	1
[155] † Diandongosuchus fuyuanensis Taxon name last Modified on 01/31/23	374	496	491	0	24	0	0	1
[156] † Wannia scurriensis Taxon name last Modified on 01/31/23	671	194	181	0	32	0	0	1
[157] † Ebrachosuchus neukami Taxon name last Modified on 01/31/23	556	301	297	0	35	0	0	1
[158] † Paleorhinus sawini Taxon name last Modified on 01/31/23	595	269	265	0	29	0	0	1
[159] † Angistorhinus talainti Taxon name last Modified on 01/31/23	517	338	335	0	37	0	0	1
[160] † Mystriosuchus planirostris Taxon name last Modified on 01/31/23	343	522	502	0	44	0	0	1
[161] † Machaeroprosopus pristinus Taxon name last Modified on 01/31/23	394	468	448	0	47	0	0	1
[162] † Redondasaurus spp Taxon name last Modified on 01/31/23	525	337	325	0	39	0	0	1
[163] † Ornithosuchus woodwardi Taxon name last Modified on 01/31/23	286	573	564	0	35	0	0	1
[164] † Riojasuchus tenuisceps Taxon name last Modified on 01/31/23	146	707	694	0	43	0	0	1
[165] † Venaticosuchus rusconii Taxon name last Modified on 01/31/23	685	188	185	0	19	0	0	1
[166] † Dynamosuchus collisensis Taxon name last Modified on 01/31/23	586	302	299	0	4	0	0	1
[167] † Nundasuchus songeaensis Taxon name last Modified on 01/31/23	578	303	297	0	14	0	0	1
[168] † Turfanosuchus dabanensis Taxon name last Modified on 01/31/23	288	579	569	0	30	0	0	1
[169] † Gracilisuchus stipanicicorum Taxon name last Modified on 01/31/23	169	691	677	0	43	0	0	1
[170] † Yonghesuchus sangbiensis Taxon name last Modified on 01/31/23	662	209	205	0	22	0	0	1
[171] † Otter sandstone archosaur Taxon name last Modified on 01/31/23	835	57	51	0	3	0	0	1
[172] † Revueltosaurus callenderi Taxon name last Modified on 01/31/23	151	705	698	0	40	0	0	1
[173] † Aetosauroides scagliai Taxon name last Modified on 01/31/23	132	729	715	0	41	0	0	1
[174] † Aetosaurus ferratus Taxon name last Modified on 01/31/23	252	607	600	0	37	0	0	1
[175] † Stagonolepis robertsoni Taxon name last Modified on 01/31/23	112	738	736	0	41	0	0	1
[176] † Ticinosuchus ferox Taxon name last Modified on 01/31/23	535	338	330	0	23	0	0	1
[177] † Prestosuchus nyassicus Taxon name last Modified on 01/31/23	707	182	174	0	8	0	0	1
[178] † Dagasuchus santacruzensis Taxon name last Modified on 01/31/23	854	33	33	0	2	0	0	1
[179] † Qianosuchus mixtus Taxon name last Modified on 01/31/23	352	506	504	0	33	0	0	1
[180] † Arizonasaurus babbitti Taxon name last Modified on 01/31/23	413	440	435	0	41	0	0	1
[181] † Lotosaurus adentus Taxon name last Modified on 01/31/23	356	459	450	0	83	0	0	1
[182] † Ctenosauriscus koeneni Taxon name last Modified on 01/31/23	836	36	36	0	17	0	0	1
[183] † Hypselorhachis mirabilis Taxon name last Modified on 01/31/23	862	26	25	0	2	0	0	1
[184] † Waldhaus poposauroid Taxon name last Modified on 01/31/23	849	41	39	0	1	0	0	1
[185] † Vytshegdosuchus zbeshartensis Taxon name last Modified on 01/31/23	875	13	13	0	1	0	0	1
[186] † Bystrowisuchus flerovi Taxon name last Modified on 01/31/23	868	21	19	0	2	0	0	1
[187] † Bromsgroveia walkeri Taxon name last Modified on 01/31/23	844	44	44	0	1	0	0	1
[188] † Xilousuchus sapingensis Taxon name last Modified on 01/31/23	633	239	237	0	18	0	0	1
[189] † Moenkopi poposauroid Taxon name last Modified on 01/31/23	838	48	44	0	7	0	0	1
[190] † Mandasuchus tanyauchen Taxon name last Modified on 01/31/23	572	304	297	0	20	0	0	1
[191] † Luperosuchus fractus Taxon name last Modified on 01/31/23	756	131	129	0	4	0	0	1
[192] † Decuriasuchus quartacolonia Taxon name last Modified on 01/31/23	410	450	444	0	35	0	0	1
[193] † Batrachotomus kupferzellensis Taxon name last Modified on 01/31/23	191	668	658	0	36	0	0	1
[194] † Prestosuchus chiniquensis Taxon name last Modified on 01/31/23	97	757	752	0	40	0	0	1
[195] † Scleromochlus taylori Taxon name last Modified on 01/31/23	687	183	171	0	31	0	0	1
[196] † Austriadraco dallavecchiai Taxon name last Modified on 01/31/23	697	196	180	0	12	0	0	1
[197] † Preondactylus buffarinii Taxon name last Modified on 01/31/23	615	237	228	0	44	0	0	1
[198] † Austriadactylus cristatus Taxon name last Modified on 01/31/23	640	223	218	0	31	0	0	1
[199] † Seazzadactylus venieri Taxon name last Modified on 01/31/23	612	240	231	0	46	0	0	1
[200] † Raeticodactylus filisurensis Taxon name last Modified on 01/31/23	628	240	232	0	29	0	0	1
[201] † Carniadactylus rosenfeldi Taxon name last Modified on 01/31/23	526	302	296	0	67	0	0	1
[202] † Peteinosaurus zambellii Taxon name last Modified on 01/31/23	631	225	219	0	39	0	0	1
[203] † Caelestiventus hanseni Taxon name last Modified on 01/31/23	699	159	153	0	37	0	0	1
[204] † Dimorphodon macronyx Taxon name last Modified on 01/31/23	387	435	431	0	71	0	0	1
[205] † Eudimorphodon ranzii Taxon name last Modified on 01/31/23	521	327	324	0	44	0	0	1
[206] † Rhamphorhynchus muensteri Taxon name last Modified on 01/31/23	196	571	563	0	130	0	0	1
[207] † Cacibupteryx caribensis Taxon name last Modified on 01/31/23	717	147	143	0	27	0	0	1
[208] † Allkaruen koi Taxon name last Modified on 01/31/23	751	128	126	0	12	0	0	1
[209] † Lagerpeton chanarensis Taxon name last Modified on 01/31/23	577	279	273	0	37	0	0	1
[210] † Ixalerpeton polesinensis Taxon name last Modified on 01/31/23	490	368	362	0	37	0	0	1
[211] † Kongonaphon kely Taxon name last Modified on 01/31/23	803	77	73	0	13	0	0	1
[212] † PVSJ 883 Taxon name last Modified on 01/31/23	878	10	10	0	1	0	0	1
[213] † Dromomeron romeri Taxon name last Modified on 01/31/23	711	155	147	0	31	0	0	1
[214] † Dromomeron gregorii Taxon name last Modified on 01/31/23	744	140	140	0	5	0	0	1
[215] † Dromomeron gigas Taxon name last Modified on 01/31/23	851	32	32	0	6	0	0	1
[216] † Lagosuchus talampayensis Taxon name last Modified on 01/31/23	445	413	406	0	38	0	0	1
[217] † Lutungutali sitwensis Taxon name last Modified on 01/31/23	847	44	39	0	3	0	0	1
[218] † Asilisaurus kongwe Taxon name last Modified on 01/31/23	340	520	511	0	38	0	0	1
[219] † Nyasasaurus parringtoni Taxon name last Modified on 01/31/23	835	51	51	0	3	0	0	1
[220] † Lewisuchus admixtus Taxon name last Modified on 01/31/23	306	560	547	0	31	0	0	1
[221] † Silesaurus opolensis Taxon name last Modified on 01/31/23	188	661	650	0	50	0	0	1
[222] † Heterodontosaurus tucki Taxon name last Modified on 01/31/23	86	718	711	0	89	0	0	1
[223] † Lesothosaurus diagnosticus Taxon name last Modified on 01/31/23	110	703	696	0	82	0	0	1
[224] † Herrerasaurus ischigualastensis Taxon name last Modified on 01/31/23	81	764	750	0	58	0	0	1
[225] † Eoraptor lunensis Taxon name last Modified on 01/31/23	147	697	689	0	52	0	0	1
[226] † Buriolestes schultzi Taxon name last Modified on 01/31/23	116	721	714	0	57	0	0	1
[227] † Saturnalia tupiniquim Taxon name last Modified on 01/31/23	328	524	512	0	48	0	0	1
[228] † Tawa hallae Taxon name last Modified on 01/31/23	239	593	593	0	57	0	0	1
[229] † Coelophysis bauri Taxon name last Modified on 01/31/23	80	740	732	0	77	0	0	1
[230] † Megapnosaurus rhodesiensis Taxon name last Modified on 01/31/23	89	728	720	0	80	0	0	1

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	4
Matrix downloads	3	Data matrix from supplemental materials (3 downloads);
Project downloads	1