Morphobank

Project 4765: O. R. Regalado Fernandez, F. Sour-Tovar. 2023. The agnostid trilobite assemblage from San José de Gracia, Sonora, México, and the implications of phylogenetics in stratigraphic calibration. Historical Biology. 35 (3):436-456.

Specimen: † Acidiscus birdi (unvouchered)
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Abstract

The outcrop of San José de Gracia yields an agnostid assemblage that has been assigned to the middle Cambrian. Although agnostid trilobites are important biostratigraphic markers, the mosaic evolution of this group makes species-level determination difficult, and thus biostratigraphic correlations become cumbersome. The assemblage is described here using morphogroups of the ‘peronopsid’-like agnostids, and specimen-level phylogenetics is used to assess the statistical significance of assigning different ages to the final topology (stratigraphic fit), including the morphogroups as operational taxonomic units. A proper phylogenetic analysis requires considering modularity and ontogenetic development in agnostids, but the overall topology can respond to different stratigraphic calibrations even if the individual relationships are not resolved. Using the distribution of several stratigraphic fit metrics, namely SCI, GER and MSM*, we find that the biozone Plagiura-Poliella (509–598.4 Mya) is the one that offers the best stratigraphic fit for the outcrop of San José de Gracia, and it is consistent with the current biostratigraphic framework that has been built for the middle Cambrian of Sonora.

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Article DOI: 10.1080/08912963.2022.2044323

Project DOI: 10.7934/P4765, http://dx.doi.org/10.7934/P4765

This project contains	Matrices
8 Media 1 Matrix 1 Document 124 Taxa 9 Specimens 129 Characters Total size of project's media files: 50.24M Download Project SDD File	Total scored cells: 14458 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 129 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 311 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:129/0

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MorphoBank Project 4765

Creation Date:
20 July 2023
Publication Date:
27 July 2023
Project views: 4570
Media downloads: 2

Authors' Institutions

Universidad Nacional Autonoma de Mexico (UNAM)

University College London

Members

Shreya Jariwala
Project Administrator

124

129

14458
(11820, 0, 2638)

Brooke Long-Fox
Full membership

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Taxonomic Overview for Matrix 'M29374' (124 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Tsunyidiscus Last Modified in 07/21/23	8	152	115	0	6	0	0	2
[2] † Neocobboldia dentata Last Modified in 07/21/23	4	99	99	0	26	0	0	2
[3] † Abakolia minutus Last Modified in 07/21/23	2	104	102	0	25	0	0	2
[4] † Abakolia pauca Last Modified in 07/21/23	10	97	94	0	25	0	0	2
[5] † Acidiscus birdi Last Modified in 07/21/23	12	94	93	0	24	0	0	2
[6] † Acidiscus theristes Last Modified in 07/21/23	3	109	105	0	21	0	0	2
[7] † Acimetopus bilobatus Last Modified in 07/21/23	8	96	95	0	26	0	0	2
[8] † Alaskadiscus spinosus Last Modified in 07/21/23	2	110	108	0	19	0	0	2
[9] † Analox bipunctata Last Modified in 07/21/23	12	89	87	0	30	0	0	2
[10] † Bathydiscus dolichometopus Last Modified in 07/21/23	9	92	90	0	30	0	0	2
[11] † Bolboparia superba Last Modified in 07/21/23	13	97	96	0	20	0	0	2
[12] † Calodiscus lobatus Last Modified in 07/21/23	9	104	98	0	22	0	0	2
[13] † Moroxxonus notabilis Last Modified in 07/21/23	9	90	88	0	32	0	0	2
[14] † Chelediscus acifer Last Modified in 07/21/23	0	103	101	0	28	0	0	2
[15] † Chelediscus chathamensis Last Modified in 07/21/23	14	90	88	0	27	0	0	2
[16] † Cobboldites comleyensis Last Modified in 07/21/23	9	89	88	0	32	0	0	2
[17] † Cobboldites itsariensis Last Modified in 07/21/23	10	90	86	0	33	0	0	2
[18] † Dawsonia bohemicus Last Modified in 07/21/23	6	96	96	0	27	0	0	2
[19] † Dawsonia dawsoni Last Modified in 07/21/23	2	100	99	0	28	0	0	2
[20] † Delgadella amouslekensis Last Modified in 07/21/23	16	89	88	0	25	0	0	2
[21] † Delgadella caudatus Last Modified in 07/21/23	9	96	95	0	25	0	0	2
[22] † Delgadella lenaicus Last Modified in 07/21/23	3	101	100	0	26	0	0	2
[23] † Dicerodiscus tsunyiensis Last Modified in 07/21/23	19	89	88	0	22	0	0	2
[24] † Egyngolia obtusa Last Modified in 07/21/23	4	109	106	0	19	0	0	2
[25] † Egyngolia willochra Last Modified in 07/21/23	9	107	104	0	16	0	0	2
[26] † Egyngolia zaicevi Last Modified in 07/21/23	46	61	61	0	22	0	0	2
[27] † Ekwipagetia marginata Last Modified in 07/21/23	9	108	104	0	16	0	0	2
[28] † Ekwipagetia plicofimbria Last Modified in 07/21/23	8	108	104	0	17	0	0	2
[29] † Eodiscus borealis Last Modified in 07/21/23	3	102	101	0	25	0	0	2
[30] † Eodiscus scanicus Last Modified in 07/21/23	2	106	103	0	24	0	0	2
[31] † Hebediscina blagonravovi Last Modified in 07/21/23	17	97	94	0	18	0	0	2
[32] † Hebediscina sardoa Last Modified in 07/21/23	8	107	103	0	18	0	0	2
[33] † Hebediscina yuqingensis Last Modified in 07/21/23	12	106	103	0	14	0	0	2
[34] † Hebediscus attleborensis Last Modified in 07/21/23	16	89	88	0	25	0	0	2
[35] † Helepagetia bitruncula Last Modified in 07/21/23	58	68	66	0	5	0	0	2
[36] † Jinghediscus nummularius Last Modified in 07/21/23	60	59	58	0	11	0	0	2
[37] † Kiskinella cristata Last Modified in 07/21/23	11	102	102	0	16	0	0	2
[38] † Korobovia ocellata Last Modified in 07/21/23	8	100	98	0	23	0	0	2
[39] † Lenadiscus unicus Last Modified in 07/21/23	16	98	95	0	18	0	0	2
[40] † Leptochilodiscus punculatus Last Modified in 07/21/23	8	92	89	0	32	0	0	2
[41] † Leptochilodiscus succinctus Last Modified in 07/21/23	1	103	99	0	29	0	0	2
[42] † Litometopus longispinus Last Modified in 07/21/23	9	90	90	0	30	0	0	2
[43] † Luvsanodiscus gammatus Last Modified in 07/21/23	0	106	105	0	24	0	0	2
[44] † Macannaia maladensis Last Modified in 07/21/23	1	118	114	0	14	0	0	2
[45] † Mallagnostus desideratus Last Modified in 07/21/23	47	56	56	0	26	0	0	2
[46] † Mallagnostus limbatus Last Modified in 07/21/23	9	90	87	0	33	0	0	2
[47] † Mallagnostus llarenai Last Modified in 07/21/23	6	99	95	0	28	0	0	2
[48] † Meniscuchus menetus Last Modified in 07/21/23	8	94	92	0	29	0	0	2
[49] † Meniscuchus nanus Last Modified in 07/21/23	12	88	88	0	29	0	0	2
[50] † Limbadiscus dilata Last Modified in 07/21/23	20	92	90	0	19	0	0	2
[51] † Limbadiscus incita Last Modified in 07/21/23	16	98	94	0	19	0	0	2
[52] † Neopagetina rjonsnitzkii Last Modified in 07/21/23	9	107	105	0	15	0	0	2
[53] † Ninadiscus strobulatus Last Modified in 07/21/23	13	97	92	0	24	0	0	2
[54] † Oodiscus subgranulatus Last Modified in 07/21/23	9	92	91	0	29	0	0	2
[55] † Opsidiscus bilobatus Last Modified in 07/21/23	12	103	100	0	17	0	0	2
[56] † Opsidiscus longispinus Last Modified in 07/21/23	40	75	74	0	15	0	0	2
[57] † Opsidiscus microspinus Last Modified in 07/21/23	1	116	112	0	16	0	0	2
[58] † Pagetia bootes Last Modified in 07/21/23	1	114	113	0	15	0	0	2
[59] † Pagetia prolata Last Modified in 07/21/23	1	116	114	0	14	0	0	2
[60] † Pagetia aspinosa Last Modified in 07/21/23	1	114	113	0	15	0	0	2
[61] † Pagetia resseri Last Modified in 07/21/23	1	114	113	0	15	0	0	2
[62] † Pagetides elegans Last Modified in 07/21/23	1	116	112	0	16	0	0	2
[63] † Pagetides fragum Last Modified in 07/21/23	8	107	104	0	17	0	0	2
[64] † Pseudocobboldia pulchra Last Modified in 07/21/23	10	100	97	0	22	0	0	2
[65] † Runcinodiscus index Last Modified in 07/21/23	8	96	95	0	26	0	0	2
[66] † Semadiscus sollennis Last Modified in 07/21/23	45	58	58	0	26	0	0	2
[67] † Serrodiscus ctenoa Last Modified in 07/21/23	0	105	102	0	27	0	0	2
[68] † Serrodiscus daedalus Last Modified in 07/21/23	11	98	94	0	24	0	0	2
[69] † Serrodiscus gravestocki Last Modified in 07/21/23	11	97	95	0	23	0	0	2
[70] † Serrodiscus speciosus Last Modified in 07/21/23	1	104	102	0	26	0	0	2
[71] † Sinodiscus shipaiensis Last Modified in 07/21/23	0	106	105	0	24	0	0	2
[72] † Sinodiscus subquadratus Last Modified in 07/21/23	8	101	98	0	23	0	0	2
[73] † Sinopagetia jinnanensis Last Modified in 07/21/23	10	101	101	0	18	0	0	2
[74] † Stigmadiscus stenometopus Last Modified in 07/21/23	16	89	87	0	26	0	0	2
[75] † Tannudiscus altus Last Modified in 07/21/23	10	91	87	0	32	0	0	2
[76] † Tannudiscus balanus Last Modified in 07/21/23	0	101	98	0	31	0	0	2
[77] † Tchernyshevioides ninae Last Modified in 07/21/23	8	101	100	0	21	0	0	2
[78] † Weymouthia nobilis Last Modified in 07/21/23	27	76	76	0	26	0	0	2
[79] † Yukonia intermedia Last Modified in 07/21/23	1	115	109	0	19	0	0	2
[80] † Yukonides lacrinus Last Modified in 07/21/23	9	106	102	0	18	0	0	2
[81] † Acadagnostus syrma Last Modified in 07/21/23	0	108	108	0	21	0	0	2
[82] † Itagnostus oepiki Last Modified in 07/21/23	1	104	104	0	24	0	0	2
[83] † Archaeagnostus primigeneus mI Last Modified in 07/21/23	8	92	92	0	29	0	0	2
[84] † Redeagnostus ferox IIa Last Modified in 07/21/23	1	103	103	0	25	0	0	2
[85] † Pseudoperonopsis bifidus IIb Last Modified in 07/21/23	7	97	97	0	25	0	0	2
[86] † Pentagnostus brighamensis mIII Last Modified in 07/21/23	0	101	101	0	28	0	0	2
[87] † Peronopsis montis Va Last Modified in 07/21/23	1	101	101	0	27	0	0	2
[88] † Peronopsis amplaxis Va Last Modified in 07/21/23	0	101	101	0	28	0	0	2
[89] † Itagnostus interstricta Vb Last Modified in 07/21/23	1	104	104	0	24	0	0	2
[90] † Quadragnostus depressa VI Last Modified in 07/21/23	7	99	99	0	23	0	0	2
[91] † Diplorrhina cuneifera VII Last Modified in 07/21/23	7	98	98	0	24	0	0	2
[92] † Gratagnostus latus VIII Last Modified in 07/21/23	7	99	99	0	23	0	0	2
[93] † P svenax egenus IX Last Modified in 07/21/23	7	97	97	0	25	0	0	2
[94] † Baltagnostus rakuroensis Xa Last Modified in 07/21/23	89	39	39	0	1	0	0	2
[95] † P vulgagnostus longinqua Xa Last Modified in 07/21/23	8	98	98	0	23	0	0	2
[96] † Ammagnostus jegorovae Xb Last Modified in 07/21/23	7	95	95	0	27	0	0	2
[97] † Pentagnostus bonnerensis mXII Last Modified in 07/21/23	9	89	89	0	31	0	0	2
[98] † P svenax pusillus XIII Last Modified in 07/21/23	7	98	98	0	24	0	0	2
[99] † Itagnostus gaspensis XIVa Last Modified in 07/21/23	6	99	99	0	24	0	0	2
[100] † Acadagnostus acadica XIVb Last Modified in 07/21/23	7	98	98	0	24	0	0	2
[101] † P proacadagnostus normata XIVb Last Modified in 07/21/23	7	100	100	0	22	0	0	2
[102] † Peronopsis segmenta mXVI Last Modified in 07/21/23	90	39	39	0	0	0	0	2
[103] † Peronopsis integer Last Modified in 07/21/23	4	98	98	0	27	0	0	2
[104] † Peronopsis roddyi Last Modified in 07/21/23	1	100	100	0	28	0	0	2
[105] † Triplagnostus gibbus Last Modified in 07/21/23	0	105	105	0	24	0	0	2
[106] † Agnostus agnostus pisiformis Last Modified in 07/21/23	2	103	103	0	24	0	0	2
[107] † Doryagnostus incertus Last Modified in 07/21/23	7	99	99	0	23	0	0	2
[108] † Diplagnostus planicauda Last Modified in 07/21/23	7	101	101	0	21	0	0	2
[109] † Clavagnostus repandus Last Modified in 07/21/23	7	95	95	0	27	0	0	2
[110] † Glyptagnostus reticulatus Last Modified in 07/21/23	7	98	98	0	24	0	0	2
[111] † Ptychagnostus praecurrens Last Modified in 07/21/23	0	103	103	0	26	0	0	2
[112] † Condylopyge amitina Last Modified in 07/21/23	0	102	102	0	27	0	0	2
[113] † Pleuroctenium granulatum Last Modified in 07/21/23	0	100	100	0	29	0	0	2
[114] † W interstricta FCMP 1106+1107+1108 Last Modified in 07/21/23	27	109	102	0	0	0	0	2
[115] † P interstricta FCMP 1105 Last Modified in 07/21/23	89	38	38	0	2	0	0	2
[116] † P interstricta FCMP 1110+1111 Last Modified in 07/21/23	87	45	42	0	0	0	0	2
[117] † W interstricta FCMP 1109 Last Modified in 07/21/23	3	97	97	0	29	0	0	2
[118] † P montis FCMP 1100+1101 Last Modified in 07/21/23	96	35	33	0	0	0	0	2
[119] † C montis FCMP 1102 Last Modified in 07/21/23	44	61	61	0	24	0	0	2
[120] † P montis FCMP 1103 Last Modified in 07/21/23	91	37	37	0	1	0	0	2
[121] † P montis FCMP 1104 Last Modified in 07/21/23	90	37	37	0	2	0	0	2
[122] † W brighamensis FCMP1126+1125+1127 Last Modified in 07/21/23	36	97	93	0	0	0	0	2
[123] † P praecurrens FCMP all Last Modified in 07/21/23	24	114	105	0	0	0	0	2
[124] † W bonnerensis FCMP all Last Modified in 07/21/23	21	142	108	0	0	0	0	2

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type	number of views	Individual items viewed (where applicable)
Total project views	4570
Project overview	210
Matrix views	129	Matrix landing page (124 views); Fernandez et al 2023 (5 views);
Media views	1401	Media search (438 views); M890389 (136 views); M890390 (80 views); M890391 (224 views); M890392 (77 views); M890393 (66 views); M890394 (129 views); M890395 (125 views); M890396 (126 views);
Documents list	118
Bibliography	92
Views for media list	156
Specimen list	924
Taxon list	1540

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type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	2
Media downloads	2	M890391 (2 downloads);