Morphobank

Project 4765: O. R. Regalado Fernandez, F. Sour-Tovar. 2023. The agnostid trilobite assemblage from San José de Gracia, Sonora, México, and the implications of phylogenetics in stratigraphic calibration. Historical Biology. 35 (3):436-456.

Specimen: † Acidiscus birdi (unvouchered)
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Abstract

The outcrop of San José de Gracia yields an agnostid assemblage that has been assigned to the middle Cambrian. Although agnostid trilobites are important biostratigraphic markers, the mosaic evolution of this group makes species-level determination difficult, and thus biostratigraphic correlations become cumbersome. The assemblage is described here using morphogroups of the ‘peronopsid’-like agnostids, and specimen-level phylogenetics is used to assess the statistical significance of assigning different ages to the final topology (stratigraphic fit), including the morphogroups as operational taxonomic units. A proper phylogenetic analysis requires considering modularity and ontogenetic development in agnostids, but the overall topology can respond to different stratigraphic calibrations even if the individual relationships are not resolved. Using the distribution of several stratigraphic fit metrics, namely SCI, GER and MSM*, we find that the biozone Plagiura-Poliella (509–598.4 Mya) is the one that offers the best stratigraphic fit for the outcrop of San José de Gracia, and it is consistent with the current biostratigraphic framework that has been built for the middle Cambrian of Sonora.

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Article DOI: 10.1080/08912963.2022.2044323

Project DOI: 10.7934/P4765, http://dx.doi.org/10.7934/P4765

This project contains	Matrices
8 Media 1 Matrix 1 Document 124 Taxa 9 Specimens 129 Characters Total size of project's media files: 50.24M Download Project SDD File	Total scored cells: 14458 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 129 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 311 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:129/0

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MorphoBank Project 4765

Creation Date:
20 July 2023
Publication Date:
27 July 2023
Media downloads: 2

Authors' Institutions

Universidad Nacional Autonoma de Mexico (UNAM)

University College London

Members

Shreya Jariwala
Project Administrator

124

129

14458
(11820, 0, 2638)

Brooke Long-Fox
Full membership

0
(0, 0, 0)

Taxonomic Overview for Matrix 'M29374' (124 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Tsunyidiscus Taxon name last Modified on 07/21/23	8	152	115	0	6	0	0	2
[2] † Neocobboldia dentata Taxon name last Modified on 07/21/23	4	99	99	0	26	0	0	2
[3] † Abakolia minutus Taxon name last Modified on 07/21/23	2	104	102	0	25	0	0	2
[4] † Abakolia pauca Taxon name last Modified on 07/21/23	10	97	94	0	25	0	0	2
[5] † Acidiscus birdi Taxon name last Modified on 07/21/23	12	94	93	0	24	0	0	2
[6] † Acidiscus theristes Taxon name last Modified on 07/21/23	3	109	105	0	21	0	0	2
[7] † Acimetopus bilobatus Taxon name last Modified on 07/21/23	8	96	95	0	26	0	0	2
[8] † Alaskadiscus spinosus Taxon name last Modified on 07/21/23	2	110	108	0	19	0	0	2
[9] † Analox bipunctata Taxon name last Modified on 07/21/23	12	89	87	0	30	0	0	2
[10] † Bathydiscus dolichometopus Taxon name last Modified on 07/21/23	9	92	90	0	30	0	0	2
[11] † Bolboparia superba Taxon name last Modified on 07/21/23	13	97	96	0	20	0	0	2
[12] † Calodiscus lobatus Taxon name last Modified on 07/21/23	9	104	98	0	22	0	0	2
[13] † Moroxxonus notabilis Taxon name last Modified on 07/21/23	9	90	88	0	32	0	0	2
[14] † Chelediscus acifer Taxon name last Modified on 07/21/23	0	103	101	0	28	0	0	2
[15] † Chelediscus chathamensis Taxon name last Modified on 07/21/23	14	90	88	0	27	0	0	2
[16] † Cobboldites comleyensis Taxon name last Modified on 07/21/23	9	89	88	0	32	0	0	2
[17] † Cobboldites itsariensis Taxon name last Modified on 07/21/23	10	90	86	0	33	0	0	2
[18] † Dawsonia bohemicus Taxon name last Modified on 07/21/23	6	96	96	0	27	0	0	2
[19] † Dawsonia dawsoni Taxon name last Modified on 07/21/23	2	100	99	0	28	0	0	2
[20] † Delgadella amouslekensis Taxon name last Modified on 07/21/23	16	89	88	0	25	0	0	2
[21] † Delgadella caudatus Taxon name last Modified on 07/21/23	9	96	95	0	25	0	0	2
[22] † Delgadella lenaicus Taxon name last Modified on 07/21/23	3	101	100	0	26	0	0	2
[23] † Dicerodiscus tsunyiensis Taxon name last Modified on 07/21/23	19	89	88	0	22	0	0	2
[24] † Egyngolia obtusa Taxon name last Modified on 07/21/23	4	109	106	0	19	0	0	2
[25] † Egyngolia willochra Taxon name last Modified on 07/21/23	9	107	104	0	16	0	0	2
[26] † Egyngolia zaicevi Taxon name last Modified on 07/21/23	46	61	61	0	22	0	0	2
[27] † Ekwipagetia marginata Taxon name last Modified on 07/21/23	9	108	104	0	16	0	0	2
[28] † Ekwipagetia plicofimbria Taxon name last Modified on 07/21/23	8	108	104	0	17	0	0	2
[29] † Eodiscus borealis Taxon name last Modified on 07/21/23	3	102	101	0	25	0	0	2
[30] † Eodiscus scanicus Taxon name last Modified on 07/21/23	2	106	103	0	24	0	0	2
[31] † Hebediscina blagonravovi Taxon name last Modified on 07/21/23	17	97	94	0	18	0	0	2
[32] † Hebediscina sardoa Taxon name last Modified on 07/21/23	8	107	103	0	18	0	0	2
[33] † Hebediscina yuqingensis Taxon name last Modified on 07/21/23	12	106	103	0	14	0	0	2
[34] † Hebediscus attleborensis Taxon name last Modified on 07/21/23	16	89	88	0	25	0	0	2
[35] † Helepagetia bitruncula Taxon name last Modified on 07/21/23	58	68	66	0	5	0	0	2
[36] † Jinghediscus nummularius Taxon name last Modified on 07/21/23	60	59	58	0	11	0	0	2
[37] † Kiskinella cristata Taxon name last Modified on 07/21/23	11	102	102	0	16	0	0	2
[38] † Korobovia ocellata Taxon name last Modified on 07/21/23	8	100	98	0	23	0	0	2
[39] † Lenadiscus unicus Taxon name last Modified on 07/21/23	16	98	95	0	18	0	0	2
[40] † Leptochilodiscus punculatus Taxon name last Modified on 07/21/23	8	92	89	0	32	0	0	2
[41] † Leptochilodiscus succinctus Taxon name last Modified on 07/21/23	1	103	99	0	29	0	0	2
[42] † Litometopus longispinus Taxon name last Modified on 07/21/23	9	90	90	0	30	0	0	2
[43] † Luvsanodiscus gammatus Taxon name last Modified on 07/21/23	0	106	105	0	24	0	0	2
[44] † Macannaia maladensis Taxon name last Modified on 07/21/23	1	118	114	0	14	0	0	2
[45] † Mallagnostus desideratus Taxon name last Modified on 07/21/23	47	56	56	0	26	0	0	2
[46] † Mallagnostus limbatus Taxon name last Modified on 07/21/23	9	90	87	0	33	0	0	2
[47] † Mallagnostus llarenai Taxon name last Modified on 07/21/23	6	99	95	0	28	0	0	2
[48] † Meniscuchus menetus Taxon name last Modified on 07/21/23	8	94	92	0	29	0	0	2
[49] † Meniscuchus nanus Taxon name last Modified on 07/21/23	12	88	88	0	29	0	0	2
[50] † Limbadiscus dilata Taxon name last Modified on 07/21/23	20	92	90	0	19	0	0	2
[51] † Limbadiscus incita Taxon name last Modified on 07/21/23	16	98	94	0	19	0	0	2
[52] † Neopagetina rjonsnitzkii Taxon name last Modified on 07/21/23	9	107	105	0	15	0	0	2
[53] † Ninadiscus strobulatus Taxon name last Modified on 07/21/23	13	97	92	0	24	0	0	2
[54] † Oodiscus subgranulatus Taxon name last Modified on 07/21/23	9	92	91	0	29	0	0	2
[55] † Opsidiscus bilobatus Taxon name last Modified on 07/21/23	12	103	100	0	17	0	0	2
[56] † Opsidiscus longispinus Taxon name last Modified on 07/21/23	40	75	74	0	15	0	0	2
[57] † Opsidiscus microspinus Taxon name last Modified on 07/21/23	1	116	112	0	16	0	0	2
[58] † Pagetia bootes Taxon name last Modified on 07/21/23	1	114	113	0	15	0	0	2
[59] † Pagetia prolata Taxon name last Modified on 07/21/23	1	116	114	0	14	0	0	2
[60] † Pagetia aspinosa Taxon name last Modified on 07/21/23	1	114	113	0	15	0	0	2
[61] † Pagetia resseri Taxon name last Modified on 07/21/23	1	114	113	0	15	0	0	2
[62] † Pagetides elegans Taxon name last Modified on 07/21/23	1	116	112	0	16	0	0	2
[63] † Pagetides fragum Taxon name last Modified on 07/21/23	8	107	104	0	17	0	0	2
[64] † Pseudocobboldia pulchra Taxon name last Modified on 07/21/23	10	100	97	0	22	0	0	2
[65] † Runcinodiscus index Taxon name last Modified on 07/21/23	8	96	95	0	26	0	0	2
[66] † Semadiscus sollennis Taxon name last Modified on 07/21/23	45	58	58	0	26	0	0	2
[67] † Serrodiscus ctenoa Taxon name last Modified on 07/21/23	0	105	102	0	27	0	0	2
[68] † Serrodiscus daedalus Taxon name last Modified on 07/21/23	11	98	94	0	24	0	0	2
[69] † Serrodiscus gravestocki Taxon name last Modified on 07/21/23	11	97	95	0	23	0	0	2
[70] † Serrodiscus speciosus Taxon name last Modified on 07/21/23	1	104	102	0	26	0	0	2
[71] † Sinodiscus shipaiensis Taxon name last Modified on 07/21/23	0	106	105	0	24	0	0	2
[72] † Sinodiscus subquadratus Taxon name last Modified on 07/21/23	8	101	98	0	23	0	0	2
[73] † Sinopagetia jinnanensis Taxon name last Modified on 07/21/23	10	101	101	0	18	0	0	2
[74] † Stigmadiscus stenometopus Taxon name last Modified on 07/21/23	16	89	87	0	26	0	0	2
[75] † Tannudiscus altus Taxon name last Modified on 07/21/23	10	91	87	0	32	0	0	2
[76] † Tannudiscus balanus Taxon name last Modified on 07/21/23	0	101	98	0	31	0	0	2
[77] † Tchernyshevioides ninae Taxon name last Modified on 07/21/23	8	101	100	0	21	0	0	2
[78] † Weymouthia nobilis Taxon name last Modified on 07/21/23	27	76	76	0	26	0	0	2
[79] † Yukonia intermedia Taxon name last Modified on 07/21/23	1	115	109	0	19	0	0	2
[80] † Yukonides lacrinus Taxon name last Modified on 07/21/23	9	106	102	0	18	0	0	2
[81] † Acadagnostus syrma Taxon name last Modified on 07/21/23	0	108	108	0	21	0	0	2
[82] † Itagnostus oepiki Taxon name last Modified on 07/21/23	1	104	104	0	24	0	0	2
[83] † Archaeagnostus primigeneus mI Taxon name last Modified on 07/21/23	8	92	92	0	29	0	0	2
[84] † Redeagnostus ferox IIa Taxon name last Modified on 07/21/23	1	103	103	0	25	0	0	2
[85] † Pseudoperonopsis bifidus IIb Taxon name last Modified on 07/21/23	7	97	97	0	25	0	0	2
[86] † Pentagnostus brighamensis mIII Taxon name last Modified on 07/21/23	0	101	101	0	28	0	0	2
[87] † Peronopsis montis Va Taxon name last Modified on 07/21/23	1	101	101	0	27	0	0	2
[88] † Peronopsis amplaxis Va Taxon name last Modified on 07/21/23	0	101	101	0	28	0	0	2
[89] † Itagnostus interstricta Vb Taxon name last Modified on 07/21/23	1	104	104	0	24	0	0	2
[90] † Quadragnostus depressa VI Taxon name last Modified on 07/21/23	7	99	99	0	23	0	0	2
[91] † Diplorrhina cuneifera VII Taxon name last Modified on 07/21/23	7	98	98	0	24	0	0	2
[92] † Gratagnostus latus VIII Taxon name last Modified on 07/21/23	7	99	99	0	23	0	0	2
[93] † P svenax egenus IX Taxon name last Modified on 07/21/23	7	97	97	0	25	0	0	2
[94] † Baltagnostus rakuroensis Xa Taxon name last Modified on 07/21/23	89	39	39	0	1	0	0	2
[95] † P vulgagnostus longinqua Xa Taxon name last Modified on 07/21/23	8	98	98	0	23	0	0	2
[96] † Ammagnostus jegorovae Xb Taxon name last Modified on 07/21/23	7	95	95	0	27	0	0	2
[97] † Pentagnostus bonnerensis mXII Taxon name last Modified on 07/21/23	9	89	89	0	31	0	0	2
[98] † P svenax pusillus XIII Taxon name last Modified on 07/21/23	7	98	98	0	24	0	0	2
[99] † Itagnostus gaspensis XIVa Taxon name last Modified on 07/21/23	6	99	99	0	24	0	0	2
[100] † Acadagnostus acadica XIVb Taxon name last Modified on 07/21/23	7	98	98	0	24	0	0	2
[101] † P proacadagnostus normata XIVb Taxon name last Modified on 07/21/23	7	100	100	0	22	0	0	2
[102] † Peronopsis segmenta mXVI Taxon name last Modified on 07/21/23	90	39	39	0	0	0	0	2
[103] † Peronopsis integer Taxon name last Modified on 07/21/23	4	98	98	0	27	0	0	2
[104] † Peronopsis roddyi Taxon name last Modified on 07/21/23	1	100	100	0	28	0	0	2
[105] † Triplagnostus gibbus Taxon name last Modified on 07/21/23	0	105	105	0	24	0	0	2
[106] † Agnostus agnostus pisiformis Taxon name last Modified on 07/21/23	2	103	103	0	24	0	0	2
[107] † Doryagnostus incertus Taxon name last Modified on 07/21/23	7	99	99	0	23	0	0	2
[108] † Diplagnostus planicauda Taxon name last Modified on 07/21/23	7	101	101	0	21	0	0	2
[109] † Clavagnostus repandus Taxon name last Modified on 07/21/23	7	95	95	0	27	0	0	2
[110] † Glyptagnostus reticulatus Taxon name last Modified on 07/21/23	7	98	98	0	24	0	0	2
[111] † Ptychagnostus praecurrens Taxon name last Modified on 07/21/23	0	103	103	0	26	0	0	2
[112] † Condylopyge amitina Taxon name last Modified on 07/21/23	0	102	102	0	27	0	0	2
[113] † Pleuroctenium granulatum Taxon name last Modified on 07/21/23	0	100	100	0	29	0	0	2
[114] † W interstricta FCMP 1106+1107+1108 Taxon name last Modified on 07/21/23	27	109	102	0	0	0	0	2
[115] † P interstricta FCMP 1105 Taxon name last Modified on 07/21/23	89	38	38	0	2	0	0	2
[116] † P interstricta FCMP 1110+1111 Taxon name last Modified on 07/21/23	87	45	42	0	0	0	0	2
[117] † W interstricta FCMP 1109 Taxon name last Modified on 07/21/23	3	97	97	0	29	0	0	2
[118] † P montis FCMP 1100+1101 Taxon name last Modified on 07/21/23	96	35	33	0	0	0	0	2
[119] † C montis FCMP 1102 Taxon name last Modified on 07/21/23	44	61	61	0	24	0	0	2
[120] † P montis FCMP 1103 Taxon name last Modified on 07/21/23	91	37	37	0	1	0	0	2
[121] † P montis FCMP 1104 Taxon name last Modified on 07/21/23	90	37	37	0	2	0	0	2
[122] † W brighamensis FCMP1126+1125+1127 Taxon name last Modified on 07/21/23	36	97	93	0	0	0	0	2
[123] † P praecurrens FCMP all Taxon name last Modified on 07/21/23	24	114	105	0	0	0	0	2
[124] † W bonnerensis FCMP all Taxon name last Modified on 07/21/23	21	142	108	0	0	0	0	2

Project downloads

type	number of downloads	Individual items downloaded (where applicable)
Total downloads from project	2
Media downloads	2	M890391 (2 downloads);