Project 4775: R. S. Thompson, J. C. Parish, S. C. Maidment, P. M. Barrett. 2012. Phylogeny of the ankylosaurian dinosaurs (Ornithischia: Thyreophora). Journal of Systematic Palaeontology. 10 (2):301-312.
Abstract
Ankylosauria is a diverse clade of quadrupedal ornithischian dinosaurs whose remains are known from Middle Jurassic to latest Cretaceous sediments worldwide. Despite a long history of research, ankylosaur interrelationships remain poorly resolved and existing cladistic analyses suffer from limited character and taxon sampling. Here, we present the most comprehensive phylogenetic analysis of the group attempted to date. The traditional ankylosaurid–nodosaurid dichotomy is maintained. Ankylosauridae forms a well-resolved clade, which includes Zhongyuansaurus, the first ankylosaurid known to lack a tail club. Nodosauridae includes a number of taxa that were resolved either as ‘polacanthids’ or basal ankylosaurids in previous analyses. The use of a broader character sample allows analysis of the interrelationships of all valid ankylosaur species; this has revealed several previously unrecognized relationships. Stegosauria is recovered as the sister taxon to Ankylosauria, while Scelidosaurus is found to be a basal thyreophoran. Dedicated methods for coding continuous characters could be used in future to improve the resolution of ankylosaur phylogeny, particularly in order to explore the relationships within the poorly resolved nodosaurid clade.Read the article »
Article DOI: 10.1080/14772019.2011.569091
Project DOI: 10.7934/P4775, http://dx.doi.org/10.7934/P4775
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 4260 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 170 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 352 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:170/0 |
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MorphoBank Project 4775
MorphoBank Project 4775
- Creation Date:
25 July 2023 - Publication Date:
01 August 2023
Authors' Institutions
- Natural History Museum, London
- University of Oxford
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Shreya Jariwala Project Administrator | 51 | 1 | 1 | 170 | 0 | 0 | 4260 (4260, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29383' (51 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Lesothosaurus diagnotsicus Taxon name last Modified on 07/25/23 | 19 | 151 | 151 | 0 | 0 | 0 | 0 | 1 |
[2] † Emausaurus Taxon name last Modified on 07/25/23 | 103 | 67 | 67 | 0 | 0 | 0 | 0 | 1 |
[3] † Scutellosaurus Taxon name last Modified on 07/25/23 | 89 | 81 | 81 | 0 | 0 | 0 | 0 | 1 |
[4] † Scelidosaurus Taxon name last Modified on 07/25/23 | 27 | 143 | 143 | 0 | 0 | 0 | 0 | 1 |
[5] † Huayangosaurus tabaii Taxon name last Modified on 07/25/23 | 31 | 139 | 139 | 0 | 0 | 0 | 0 | 1 |
[6] † Stegosaurus Taxon name last Modified on 07/25/23 | 15 | 155 | 155 | 0 | 0 | 0 | 0 | 1 |
[7] † Anoplosaurus curtonotus Taxon name last Modified on 07/25/23 | 152 | 18 | 18 | 0 | 0 | 0 | 0 | 1 |
[8] † Animantarx ramaljonsei Taxon name last Modified on 07/25/23 | 67 | 103 | 103 | 0 | 0 | 0 | 0 | 1 |
[9] † Ankylosaurus marginoventris Taxon name last Modified on 07/25/23 | 30 | 140 | 140 | 0 | 0 | 0 | 0 | 1 |
[10] † Antarctopelta oliveroi Taxon name last Modified on 07/25/23 | 154 | 16 | 16 | 0 | 0 | 0 | 0 | 1 |
[11] † Cedarpelta bilbeyhallorum Taxon name last Modified on 07/25/23 | 87 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[12] † Chrichtonsaurus benxiensis Taxon name last Modified on 07/25/23 | 62 | 108 | 108 | 0 | 0 | 0 | 0 | 1 |
[13] † Edmontonia Taxon name last Modified on 07/25/23 | 18 | 152 | 152 | 0 | 0 | 0 | 0 | 1 |
[14] † Euoplocephalus tutus Taxon name last Modified on 07/25/23 | 4 | 166 | 166 | 0 | 0 | 0 | 0 | 1 |
[15] † Gargoyleosaurus parkpinorum Taxon name last Modified on 07/25/23 | 63 | 107 | 107 | 0 | 0 | 0 | 0 | 1 |
[16] † Gastonia burgei Taxon name last Modified on 07/25/23 | 33 | 137 | 137 | 0 | 0 | 0 | 0 | 1 |
[17] † Gobisaurus domoculus Taxon name last Modified on 07/25/23 | 101 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[18] † Hoplitosaurus marshi Taxon name last Modified on 07/25/23 | 145 | 25 | 25 | 0 | 0 | 0 | 0 | 1 |
[19] † Hungarosaurus tormai Taxon name last Modified on 07/25/23 | 74 | 97 | 96 | 0 | 0 | 0 | 0 | 1 |
[20] † Hylaeosaurus armatus Taxon name last Modified on 07/25/23 | 132 | 38 | 38 | 0 | 0 | 0 | 0 | 1 |
[21] † Liaoningosaurus paradoxus Taxon name last Modified on 07/25/23 | 121 | 49 | 49 | 0 | 0 | 0 | 0 | 1 |
[22] † Minmi paravertebra Taxon name last Modified on 07/25/23 | 50 | 122 | 120 | 0 | 0 | 0 | 0 | 1 |
[23] † Minotaurasaurus ramachandrani Taxon name last Modified on 07/25/23 | 94 | 76 | 76 | 0 | 0 | 0 | 0 | 1 |
[24] † Mymoorapelta maysi Taxon name last Modified on 07/25/23 | 117 | 53 | 53 | 0 | 0 | 0 | 0 | 1 |
[25] † Niobrarasaurus coleii Taxon name last Modified on 07/25/23 | 113 | 57 | 57 | 0 | 0 | 0 | 0 | 1 |
[26] † Nodocephalosaurus kirtlandensis Taxon name last Modified on 07/25/23 | 146 | 24 | 24 | 0 | 0 | 0 | 0 | 1 |
[27] † Nodosaurus textillis Taxon name last Modified on 07/25/23 | 135 | 35 | 35 | 0 | 0 | 0 | 0 | 1 |
[28] † Panoplosaurus mirus Taxon name last Modified on 07/25/23 | 40 | 131 | 130 | 0 | 0 | 0 | 0 | 1 |
[29] † Pawpawsaurus cambelli Taxon name last Modified on 07/25/23 | 94 | 76 | 76 | 0 | 0 | 0 | 0 | 1 |
[30] † Peloroplites cedrimontanus Taxon name last Modified on 07/25/23 | 82 | 88 | 88 | 0 | 0 | 0 | 0 | 1 |
[31] † Pinacosaurus grangeri Taxon name last Modified on 07/25/23 | 18 | 152 | 152 | 0 | 0 | 0 | 0 | 1 |
[32] † Pinacosaurus mephistocephalus Taxon name last Modified on 07/25/23 | 101 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[33] † Polacanthus foxii Taxon name last Modified on 07/25/23 | 105 | 65 | 65 | 0 | 0 | 0 | 0 | 1 |
[34] † Polacanthus rudgwickensis Taxon name last Modified on 07/25/23 | 162 | 8 | 8 | 0 | 0 | 0 | 0 | 1 |
[35] † Saichania chulsanensis Taxon name last Modified on 07/25/23 | 39 | 131 | 131 | 0 | 0 | 0 | 0 | 1 |
[36] † Sauropelta edwardsi Taxon name last Modified on 07/25/23 | 33 | 137 | 137 | 0 | 0 | 0 | 0 | 1 |
[37] † Shamosaurus scutatus Taxon name last Modified on 07/25/23 | 65 | 105 | 105 | 0 | 0 | 0 | 0 | 1 |
[38] † Shanxia tianzhenensis Taxon name last Modified on 07/25/23 | 132 | 38 | 38 | 0 | 0 | 0 | 0 | 1 |
[39] † Silvisaurus condrayi Taxon name last Modified on 07/25/23 | 66 | 104 | 104 | 0 | 0 | 0 | 0 | 1 |
[40] † Stegopelta landerensis Taxon name last Modified on 07/25/23 | 152 | 19 | 18 | 0 | 0 | 0 | 0 | 1 |
[41] † Struthiosaurus austriacus Taxon name last Modified on 07/25/23 | 68 | 102 | 102 | 0 | 0 | 0 | 0 | 1 |
[42] † Struthiosaurus languedocensis Taxon name last Modified on 07/25/23 | 144 | 26 | 26 | 0 | 0 | 0 | 0 | 1 |
[43] † Talarurus plicatospineus Taxon name last Modified on 07/25/23 | 69 | 101 | 101 | 0 | 0 | 0 | 0 | 1 |
[44] † Tarchia gigantea Taxon name last Modified on 07/25/23 | 59 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[45] † Tatankocephalus cooneyorum Taxon name last Modified on 07/25/23 | 128 | 42 | 42 | 0 | 0 | 0 | 0 | 1 |
[46] † Texasetes pleurohalio Taxon name last Modified on 07/25/23 | 137 | 33 | 33 | 0 | 0 | 0 | 0 | 1 |
[47] † Tianzhenosaurus youngi Taxon name last Modified on 07/25/23 | 42 | 128 | 128 | 0 | 0 | 0 | 0 | 1 |
[48] † Tsagantegia longicranialis Taxon name last Modified on 07/25/23 | 102 | 68 | 68 | 0 | 0 | 0 | 0 | 1 |
[49] † Zhejiangosaurus lishuiensis Taxon name last Modified on 07/25/23 | 135 | 35 | 35 | 0 | 0 | 0 | 0 | 1 |
[50] † Zhonguansaurus lauyangensis Taxon name last Modified on 07/25/23 | 120 | 50 | 50 | 0 | 0 | 0 | 0 | 1 |
[51] † Dyoplosaurus acutosquameus Taxon name last Modified on 07/25/23 | 140 | 30 | 30 | 0 | 0 | 0 | 0 | 1 |
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