Project 4798: C. F. Kammerer. 2019. Revision of the Tanzanian dicynodont Dicynodon huenei (Therapsida: Anomodontia) from the Permian Usili Formation. PeerJ. 7:e7420.
Abstract
A single species of the dicynodontoid dicynodont Dicynodon is currently recognized from the late Permian Usili Formation of Tanzania: Dicynodon huenei Haughton, 1932. Restudy of the known Tanzanian materials of D . huenei demonstrates that they represent two distinct morphotypes, here considered separate taxa. The holotype of D . huenei is not referable to Dicynodon and instead is transferred to the genus Daptocephalus (but retained as a valid species, Daptocephalus huenei comb. nov.). A number of published dicynodontoid specimens from the Usili Formation, however, are referable to Dicynodon , and are here recognized as a new species ( Dicynodon angielczyki sp. nov.) Dicynodon angielczyki can be distinguished from its South African congener Dicynodon lacerticeps by the presence of an expansion of the squamosal and jugal beneath the postorbital bar and a curved, posterolateral expansion of the squamosal behind the temporal fenestra. Inclusion of Dicynodon angielczyki and D . huenei in a phylogenetic analysis supports their referral to Dicynodon and Daptocephalus (respectively). These results indicate higher basinal endemism in large late Permian dicynodonts than previously thought, a sharp contrast to the cosmopolitanism in the group in the earliest Triassic.Read the article »
Article DOI: 10.7717/peerj.7420
Project DOI: 10.7934/P4798, http://dx.doi.org/10.7934/P4798
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 13162 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 174 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 418 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:174/0 |
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MorphoBank Project 4798
MorphoBank Project 4798
- Creation Date:
01 August 2023 - Publication Date:
15 August 2023
Authors' Institutions
- North Carolina Museum of Natural Sciences, Raleigh
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Shreya Jariwala Project Administrator | 107 | 1 | 1 | 174 | 0 | 0 | 13162 (13162, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29406' (107 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Biarmosuchus tener Taxon name last Modified on 08/01/23 | 22 | 152 | 152 | 0 | 0 | 0 | 0 | 1 |
[2] † Hipposaurus boonstrai Taxon name last Modified on 08/01/23 | 24 | 150 | 150 | 0 | 0 | 0 | 0 | 1 |
[3] † Archaeosyodon praeventor Taxon name last Modified on 08/01/23 | 67 | 107 | 107 | 0 | 0 | 0 | 0 | 1 |
[4] † Titanophoneus potens Taxon name last Modified on 08/01/23 | 13 | 162 | 161 | 0 | 0 | 0 | 0 | 1 |
[5] † Gorgonops torvus Taxon name last Modified on 08/01/23 | 11 | 163 | 163 | 0 | 0 | 0 | 0 | 1 |
[6] † Lycosuchus vanderrieti Taxon name last Modified on 08/01/23 | 19 | 155 | 155 | 0 | 0 | 0 | 0 | 1 |
[7] † Glanosuchus macrops Taxon name last Modified on 08/01/23 | 11 | 163 | 163 | 0 | 0 | 0 | 0 | 1 |
[8] † Biseridens qilianicus Taxon name last Modified on 08/01/23 | 59 | 115 | 115 | 0 | 0 | 0 | 0 | 1 |
[9] † Anomocephalus africanus Taxon name last Modified on 08/01/23 | 146 | 28 | 28 | 0 | 0 | 0 | 0 | 1 |
[10] † Tiarajudens eccentricus Taxon name last Modified on 08/01/23 | 120 | 54 | 54 | 0 | 0 | 0 | 0 | 1 |
[11] † Otsheria netzvetajevi Taxon name last Modified on 08/01/23 | 88 | 86 | 86 | 0 | 0 | 0 | 0 | 1 |
[12] † Ulemica Taxon name last Modified on 08/01/23 | 47 | 128 | 127 | 0 | 0 | 0 | 0 | 1 |
[13] † Suminia getmanovi Taxon name last Modified on 08/01/23 | 19 | 155 | 155 | 0 | 0 | 0 | 0 | 1 |
[14] † Patranomodon nyaphulii Taxon name last Modified on 08/01/23 | 44 | 130 | 130 | 0 | 0 | 0 | 0 | 1 |
[15] † Galeops whaitsi Taxon name last Modified on 08/01/23 | 56 | 118 | 118 | 0 | 0 | 0 | 0 | 1 |
[16] † Galepus jouberti Taxon name last Modified on 08/01/23 | 134 | 40 | 40 | 0 | 0 | 0 | 0 | 1 |
[17] † Galechirus scholtzi Taxon name last Modified on 08/01/23 | 117 | 57 | 57 | 0 | 0 | 0 | 0 | 1 |
[18] † Eodicynodon oelofseni Taxon name last Modified on 08/01/23 | 80 | 94 | 94 | 0 | 0 | 0 | 0 | 1 |
[19] † Eodicynodon oosthuizeni Taxon name last Modified on 08/01/23 | 6 | 168 | 168 | 0 | 0 | 0 | 0 | 1 |
[20] † Colobodectes cluveri Taxon name last Modified on 08/01/23 | 70 | 104 | 104 | 0 | 0 | 0 | 0 | 1 |
[21] † Lanthanostegus mohoii Taxon name last Modified on 08/01/23 | 115 | 59 | 59 | 0 | 0 | 0 | 0 | 1 |
[22] † Eosimops newtoni Taxon name last Modified on 08/01/23 | 15 | 161 | 159 | 0 | 0 | 0 | 0 | 1 |
[23] † Diictodon feliceps Taxon name last Modified on 08/01/23 | 7 | 170 | 167 | 0 | 0 | 0 | 0 | 1 |
[24] † Robertia broomiana Taxon name last Modified on 08/01/23 | 12 | 162 | 162 | 0 | 0 | 0 | 0 | 1 |
[25] † Prosictodon dubei Taxon name last Modified on 08/01/23 | 48 | 126 | 126 | 0 | 0 | 0 | 0 | 1 |
[26] † Endothiodon bathystoma Taxon name last Modified on 08/01/23 | 13 | 164 | 161 | 0 | 0 | 0 | 0 | 1 |
[27] † Endothiodon tolani Taxon name last Modified on 08/01/23 | 46 | 129 | 128 | 0 | 0 | 0 | 0 | 1 |
[28] † Niassodon mfumukasi Taxon name last Modified on 08/01/23 | 43 | 131 | 131 | 0 | 0 | 0 | 0 | 1 |
[29] † Brachyprosopus broomi Taxon name last Modified on 08/01/23 | 21 | 153 | 153 | 0 | 0 | 0 | 0 | 1 |
[30] † Pristerodon mackayi Taxon name last Modified on 08/01/23 | 7 | 167 | 167 | 0 | 0 | 0 | 0 | 1 |
[31] † Emydops Taxon name last Modified on 08/01/23 | 7 | 168 | 167 | 0 | 0 | 0 | 0 | 1 |
[32] † Digalodon rubidgei Taxon name last Modified on 08/01/23 | 53 | 121 | 121 | 0 | 0 | 0 | 0 | 1 |
[33] † Dicynodontoides Taxon name last Modified on 08/01/23 | 8 | 166 | 166 | 0 | 0 | 0 | 0 | 1 |
[34] † Kombuisia frerensis Taxon name last Modified on 08/01/23 | 54 | 120 | 120 | 0 | 0 | 0 | 0 | 1 |
[35] † Myosaurus gracilis Taxon name last Modified on 08/01/23 | 27 | 147 | 147 | 0 | 0 | 0 | 0 | 1 |
[36] † Cistecephalus microrhinus Taxon name last Modified on 08/01/23 | 11 | 165 | 163 | 0 | 0 | 0 | 0 | 1 |
[37] † Cistecephaloides boonstrai Taxon name last Modified on 08/01/23 | 52 | 122 | 122 | 0 | 0 | 0 | 0 | 1 |
[38] † Kawingasaurus fossilis Taxon name last Modified on 08/01/23 | 35 | 139 | 139 | 0 | 0 | 0 | 0 | 1 |
[39] † Daqingshanodon limbus Taxon name last Modified on 08/01/23 | 52 | 122 | 122 | 0 | 0 | 0 | 0 | 1 |
[40] † Keyseria benjamini Taxon name last Modified on 08/01/23 | 74 | 100 | 100 | 0 | 0 | 0 | 0 | 1 |
[41] † Rhachiocephalus magnus Taxon name last Modified on 08/01/23 | 18 | 157 | 156 | 0 | 0 | 0 | 0 | 1 |
[42] † Kitchinganomodon crassus Taxon name last Modified on 08/01/23 | 39 | 135 | 135 | 0 | 0 | 0 | 0 | 1 |
[43] † Oudenodon bainii Taxon name last Modified on 08/01/23 | 6 | 169 | 168 | 0 | 0 | 0 | 0 | 1 |
[44] † Tropidostoma dubium Taxon name last Modified on 08/01/23 | 17 | 157 | 157 | 0 | 0 | 0 | 0 | 1 |
[45] † Australobarbarus Taxon name last Modified on 08/01/23 | 41 | 133 | 133 | 0 | 0 | 0 | 0 | 1 |
[46] † Syops vanhoepeni Taxon name last Modified on 08/01/23 | 86 | 88 | 88 | 0 | 0 | 0 | 0 | 1 |
[47] † Odontocyclops whaitsi Taxon name last Modified on 08/01/23 | 21 | 153 | 153 | 0 | 0 | 0 | 0 | 1 |
[48] † Idelesaurus tataricus Taxon name last Modified on 08/01/23 | 41 | 133 | 133 | 0 | 0 | 0 | 0 | 1 |
[49] † Aulacephalodon bainii Taxon name last Modified on 08/01/23 | 6 | 173 | 168 | 0 | 0 | 0 | 0 | 1 |
[50] † Pelanomodon moschops Taxon name last Modified on 08/01/23 | 62 | 113 | 112 | 0 | 0 | 0 | 0 | 1 |
[51] † Geikia locusticeps Taxon name last Modified on 08/01/23 | 35 | 139 | 139 | 0 | 0 | 0 | 0 | 1 |
[52] † Geikia elginensis Taxon name last Modified on 08/01/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 1 |
[53] † Elph borealis Taxon name last Modified on 08/01/23 | 60 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[54] † Interpresosaurus blomi Taxon name last Modified on 08/01/23 | 130 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[55] † Katumbia parringtoni Taxon name last Modified on 08/01/23 | 53 | 121 | 121 | 0 | 0 | 0 | 0 | 1 |
[56] † Delectosaurus arefjevi Taxon name last Modified on 08/01/23 | 66 | 109 | 108 | 0 | 0 | 0 | 0 | 1 |
[57] † Dicynodon lacerticeps Taxon name last Modified on 08/01/23 | 35 | 140 | 139 | 0 | 0 | 0 | 0 | 1 |
[58] † Daptocephalus leoniceps Taxon name last Modified on 08/01/23 | 26 | 148 | 148 | 0 | 0 | 0 | 0 | 1 |
[59] † Dinanomodon gilli Taxon name last Modified on 08/01/23 | 50 | 124 | 124 | 0 | 0 | 0 | 0 | 1 |
[60] † Peramodon amalitzkii Taxon name last Modified on 08/01/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 1 |
[61] † Vivaxosaurus trautscholdi Taxon name last Modified on 08/01/23 | 32 | 144 | 142 | 0 | 0 | 0 | 0 | 1 |
[62] † Jimusaria sinkiangensis Taxon name last Modified on 08/01/23 | 53 | 121 | 121 | 0 | 0 | 0 | 0 | 1 |
[63] † Sintocephalus alticeps Taxon name last Modified on 08/01/23 | 94 | 80 | 80 | 0 | 0 | 0 | 0 | 1 |
[64] † Basilodon woodwardi Taxon name last Modified on 08/01/23 | 63 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[65] † Turfanodon bogdaensis Taxon name last Modified on 08/01/23 | 88 | 86 | 86 | 0 | 0 | 0 | 0 | 1 |
[66] † Gordonia traquairi Taxon name last Modified on 08/01/23 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 1 |
[67] † Euptychognathus bathyrhynchus Taxon name last Modified on 08/01/23 | 60 | 114 | 114 | 0 | 0 | 0 | 0 | 1 |
[68] † Lystrosaurus hedini Taxon name last Modified on 08/01/23 | 41 | 133 | 133 | 0 | 0 | 0 | 0 | 1 |
[69] † Lystrosaurus maccaigi Taxon name last Modified on 08/01/23 | 34 | 140 | 140 | 0 | 0 | 0 | 0 | 1 |
[70] † Lystrosaurus curvatus Taxon name last Modified on 08/01/23 | 34 | 140 | 140 | 0 | 0 | 0 | 0 | 1 |
[71] † Lystrosaurus declivis Taxon name last Modified on 08/01/23 | 13 | 161 | 161 | 0 | 0 | 0 | 0 | 1 |
[72] † Lystrosaurus murrayi Taxon name last Modified on 08/01/23 | 13 | 161 | 161 | 0 | 0 | 0 | 0 | 1 |
[73] † Shansiodon Taxon name last Modified on 08/01/23 | 33 | 141 | 141 | 0 | 0 | 0 | 0 | 1 |
[74] † Tetragonias njalilus Taxon name last Modified on 08/01/23 | 9 | 166 | 165 | 0 | 0 | 0 | 0 | 1 |
[75] † Vinceria andina Taxon name last Modified on 08/01/23 | 78 | 96 | 96 | 0 | 0 | 0 | 0 | 1 |
[76] † Rhinodicynodon gracile Taxon name last Modified on 08/01/23 | 41 | 133 | 133 | 0 | 0 | 0 | 0 | 1 |
[77] † Dinodontosaurus Taxon name last Modified on 08/01/23 | 16 | 158 | 158 | 0 | 0 | 0 | 0 | 1 |
[78] † Angonisaurus cruickshanki Taxon name last Modified on 08/01/23 | 45 | 129 | 129 | 0 | 0 | 0 | 0 | 1 |
[79] † Kannemeyeria simocephalus Taxon name last Modified on 08/01/23 | 10 | 164 | 164 | 0 | 0 | 0 | 0 | 1 |
[80] † Kannemeyeria lophorhinus Taxon name last Modified on 08/01/23 | 45 | 129 | 129 | 0 | 0 | 0 | 0 | 1 |
[81] † Dolichuranus primaevus Taxon name last Modified on 08/01/23 | 30 | 144 | 144 | 0 | 0 | 0 | 0 | 1 |
[82] † Sinokannemeyeria Taxon name last Modified on 08/01/23 | 26 | 148 | 148 | 0 | 0 | 0 | 0 | 1 |
[83] † Parakannemeyeria Taxon name last Modified on 08/01/23 | 23 | 152 | 151 | 0 | 0 | 0 | 0 | 1 |
[84] † Xiyukannemeyeria brevirostris Taxon name last Modified on 08/01/23 | 46 | 128 | 128 | 0 | 0 | 0 | 0 | 1 |
[85] † Rabidosaurus cristatus Taxon name last Modified on 08/01/23 | 101 | 73 | 73 | 0 | 0 | 0 | 0 | 1 |
[86] † Rhadiodromus Taxon name last Modified on 08/01/23 | 79 | 96 | 95 | 0 | 0 | 0 | 0 | 1 |
[87] † Wadiasaurus indicus Taxon name last Modified on 08/01/23 | 64 | 111 | 110 | 0 | 0 | 0 | 0 | 1 |
[88] † Shaanbeikannemeyeria Taxon name last Modified on 08/01/23 | 53 | 121 | 121 | 0 | 0 | 0 | 0 | 1 |
[89] † Rechnisaurus cristarhynchus Taxon name last Modified on 08/01/23 | 99 | 75 | 75 | 0 | 0 | 0 | 0 | 1 |
[90] † Uralokannemeyeria vjuschkovi Taxon name last Modified on 08/01/23 | 95 | 79 | 79 | 0 | 0 | 0 | 0 | 1 |
[91] † Zambiasaurus submersus Taxon name last Modified on 08/01/23 | 111 | 63 | 63 | 0 | 0 | 0 | 0 | 1 |
[92] † Placerias hesternus Taxon name last Modified on 08/01/23 | 51 | 123 | 123 | 0 | 0 | 0 | 0 | 1 |
[93] † Moghreberia nmachouensis Taxon name last Modified on 08/01/23 | 76 | 98 | 98 | 0 | 0 | 0 | 0 | 1 |
[94] † Stahleckeria potens Taxon name last Modified on 08/01/23 | 10 | 164 | 164 | 0 | 0 | 0 | 0 | 1 |
[95] † Sangusaurus parringtonii Taxon name last Modified on 08/01/23 | 40 | 134 | 134 | 0 | 0 | 0 | 0 | 1 |
[96] † Eubrachiosaurus browni Taxon name last Modified on 08/01/23 | 163 | 11 | 11 | 0 | 0 | 0 | 0 | 1 |
[97] † Jachaleria Taxon name last Modified on 08/01/23 | 43 | 131 | 131 | 0 | 0 | 0 | 0 | 1 |
[98] † Ischigualastia jenseni Taxon name last Modified on 08/01/23 | 21 | 153 | 153 | 0 | 0 | 0 | 0 | 1 |
[99] † Rastodon procurvidens Taxon name last Modified on 08/01/23 | 71 | 103 | 103 | 0 | 0 | 0 | 0 | 1 |
[100] † Sauroscaptor tharavati Taxon name last Modified on 08/01/23 | 79 | 95 | 95 | 0 | 0 | 0 | 0 | 1 |
[101] † Bulbasaurus phylloxyron Taxon name last Modified on 08/01/23 | 51 | 127 | 123 | 0 | 0 | 0 | 0 | 1 |
[102] † Compsodon helmoedi Taxon name last Modified on 08/01/23 | 63 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[103] † Pentasaurus goggai Taxon name last Modified on 08/01/23 | 150 | 24 | 24 | 0 | 0 | 0 | 0 | 1 |
[104] † Thliptosaurus imperforatus Taxon name last Modified on 08/01/23 | 106 | 68 | 68 | 0 | 0 | 0 | 0 | 1 |
[105] † Ufudocyclops mukanelai Taxon name last Modified on 08/01/23 | 63 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[106] † Dicynodon angielczyki Taxon name last Modified on 08/01/23 | 38 | 136 | 136 | 0 | 0 | 0 | 0 | 1 |
[107] † Daptocephalus huenei Taxon name last Modified on 08/01/23 | 64 | 110 | 110 | 0 | 0 | 0 | 0 | 1 |