Morphobank

Project 4823: D. De Simão-Oliveira, F. L. Pinheiro, M. B. De Andrade, F. A. Pretto. 2022. Redescription, taxonomic revaluation and phylogenetic affinities of Proterochampsa nodosa (Archosauriformes: Proterochampsidae) from the early Late Triassic of the Candelaria Sequence (Santa Maria Supersequence). Zoological Journal of the Linnean Society. 196 (4):1310-1332.

Specimen: † Chanaresuchus bonapartei (unvouchered)
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Abstract

Proterochampsidae are a group of predatory, putatively semi-aquatic, archosauriforms endemic to the Middle and Upper Triassic strata of Argentina and Brazil (Ischigualasto Formation and Santa Maria Supersequence). Here, the Brazilian species Proterochampsa nodosa from the Candelária Sequence (Santa Maria Supersequence of the Paraná Basin) is revisited with a detailed description, phylogenetic analysis and taxonomic revaluation. Through the application of computed tomographic scans, we were able to reveal part of the previously hidden morphology of its mandible and palate. We also discuss further differences between P. nodosa and P. barrionuevoi. The new phylogenetic analysis bolsters the monophyly of Proterochampsidae, the genus Proterochampsa and its position as the basalmost proterochampsid taxon in the clade.

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Article DOI: 10.1093/zoolinnean/zlac048

Project DOI: 10.7934/P4823, http://dx.doi.org/10.7934/P4823

This project contains	Matrices
1 Media 1 Matrix 2 Documents 151 Taxa 1 Specimen 697 Characters Total size of project's media files: 31.73k Download Project SDD File	Total scored cells: 44165 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 697 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1622 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:591/106

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MorphoBank Project 4823

Creation Date:
08 August 2023
Publication Date:
10 August 2023
Project views: 2947

Authors' Institutions

Pontificia Universidade Catolica do Rio Grande do Sul (PUCRS)

Universidade Federal de Santa Maria (UFSM)

Universidade Federal do Pampa (UNIPAMPA)

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MorphoBank Curator
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Taxonomic Overview for Matrix 'M29431' (151 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Petrolacosaurus kansensis Last Modified in 08/10/23	74	543	525	0	82	0	0	1
[2] † Acerosodontosaurus piveteaui Last Modified in 08/10/23	454	196	190	0	47	0	0	1
[3] † Youngina capensis Last Modified in 08/10/23	71	562	514	0	70	0	0	1
[4] † Paliguana whitei Last Modified in 08/10/23	585	91	89	0	21	0	0	1
[5] † Planocephalosaurus robinsonae Last Modified in 08/10/23	218	406	394	0	74	0	0	1
[6] † Gephyrosaurus bridensis Last Modified in 08/10/23	194	447	420	0	60	0	0	1
[7] † Cteniogenys sp. Last Modified in 08/10/23	376	284	277	0	42	0	0	1
[8] † Simoedosaurus lemoinei Last Modified in 08/10/23	139	497	492	0	65	0	0	1
[9] † Aenigmastropheus parringtoni Last Modified in 08/10/23	656	42	40	0	0	0	0	1
[10] † Protorosaurus speneri Last Modified in 08/10/23	179	466	418	0	65	0	0	1
[11] † Dinocephalosaurus orientalis Last Modified in 08/10/23	337	283	279	0	81	0	0	1
[12] † Amotosaurus rotfeldensis Last Modified in 08/10/23	354	300	293	0	48	0	0	1
[13] † Macrocnemus bassanii Last Modified in 08/10/23	221	416	384	0	64	0	0	1
[14] † Macrocnemus obristi Last Modified in 08/10/23	618	63	62	0	17	0	0	1
[15] † Macrocnemus fuyuanensis Last Modified in 08/10/23	283	355	348	0	66	0	0	1
[16] † Fuyuansaurus acutirostris Last Modified in 08/10/23	553	124	120	0	23	0	0	1
[17] † Pectodens zhenyuensis Last Modified in 08/10/23	432	219	210	0	55	0	0	1
[18] † Augustaburiania vatagini Last Modified in 08/10/23	623	69	69	0	5	0	0	1
[19] † Protanystropheus antiquus Last Modified in 08/10/23	679	20	18	0	0	0	0	1
[20] † Trachelosaurus fischeri Last Modified in 08/10/23	609	83	73	0	15	0	0	1
[21] † Tanystropheus longobardicus Last Modified in 08/10/23	77	550	492	0	87	0	0	1
[22] † Tanystropheus haasi Last Modified in 08/10/23	688	9	9	0	0	0	0	1
[23] † Jesairosaurus lehmani Last Modified in 08/10/23	328	316	299	0	64	0	0	1
[24] † Pamelaria dolichotrachela Last Modified in 08/10/23	148	501	496	0	53	0	0	1
[25] † Azendohsaurus madagaskarensis Last Modified in 08/10/23	59	585	575	0	58	0	0	1
[26] † Azendohsaurus laaroussi Last Modified in 08/10/23	640	51	50	0	7	0	0	1
[27] † Shringasaurus indicus Last Modified in 08/10/23	361	310	304	0	32	0	0	1
[28] † Trilophosaurus buettneri Last Modified in 08/10/23	80	545	512	0	77	0	0	1
[29] † Spinosuchus combined Last Modified in 08/10/23	400	257	255	0	42	0	0	1
[30] † Teraterpeton hrynewichorum Last Modified in 08/10/23	429	224	220	0	48	0	0	1
[31] † Noteosuchus colletti Last Modified in 08/10/23	519	165	153	0	16	0	0	1
[32] † Mesosuchus browni Last Modified in 08/10/23	70	573	542	0	56	0	0	1
[33] † Howesia browni Last Modified in 08/10/23	313	357	330	0	33	0	0	1
[34] † Eohyosaurus wolvaardti Last Modified in 08/10/23	542	146	140	0	14	0	0	1
[35] † Rhynchosaurus articeps Last Modified in 08/10/23	167	477	461	0	57	0	0	1
[36] † Bentonyx sidensis Last Modified in 08/10/23	440	219	218	0	39	0	0	1
[37] † Mesodapedon kuttyi Last Modified in 08/10/23	659	35	35	0	3	0	0	1
[38] † Ammorhynchus navajoi Last Modified in 08/10/23	639	44	43	0	15	0	0	1
[39] † Stenaulorhynchus stockleyi Last Modified in 08/10/23	114	528	526	0	57	0	0	1
[40] † Brasinorhynchus mariantensis Last Modified in 08/10/23	401	263	261	0	35	0	0	1
[41] † Chanares rhynchosaur Last Modified in 08/10/23	654	43	40	0	3	0	0	1
[42] † Langeronyx brodiei Last Modified in 08/10/23	541	129	128	0	28	0	0	1
[43] † Fodonyx spenceri Last Modified in 08/10/23	559	125	125	0	13	0	0	1
[44] † Isalorhynchus genovefae Last Modified in 08/10/23	192	456	452	0	51	0	0	1
[45] † Eorasaurus olsoni Last Modified in 08/10/23	672	27	24	0	0	0	0	1
[46] † Prolacertoides jimusarensis Last Modified in 08/10/23	608	68	66	0	22	0	0	1
[47] † Prolacerta broomi Last Modified in 08/10/23	31	622	576	0	57	0	0	1
[48] † K australiensis combined Last Modified in 08/10/23	579	98	93	0	25	0	0	1
[49] † Boreopricea funerea Last Modified in 08/10/23	377	289	261	0	39	0	0	1
[50] † Teyujagua paradoxa Last Modified in 08/10/23	423	229	227	0	47	0	0	1
[51] † Archosaurus rossicus holotype Last Modified in 08/10/23	678	19	19	0	0	0	0	1
[52] † Archosaurus complete Last Modified in 08/10/23	611	87	84	0	2	0	0	1
[53] † Proterosuchus fergusi Last Modified in 08/10/23	151	526	511	0	33	0	0	1
[54] † Proterosuchus goweri Last Modified in 08/10/23	431	256	252	0	14	0	0	1
[55] † Proterosuchus alexanderi Last Modified in 08/10/23	202	463	459	0	35	0	0	1
[56] † "Chasmatosaurus" yuani Last Modified in 08/10/23	158	510	496	0	42	0	0	1
[57] † Panchet proterosuchid Last Modified in 08/10/23	537	157	148	0	12	0	0	1
[58] † Tasmaniosaurus triassicus Last Modified in 08/10/23	585	107	100	0	12	0	0	1
[59] † Vonhuenia fredericki Last Modified in 08/10/23	676	19	19	0	2	0	0	1
[60] † C rossicus combined Last Modified in 08/10/23	652	44	43	0	2	0	0	1
[61] † C magnus combined Last Modified in 08/10/23	670	26	26	0	1	0	0	1
[62] † Chasmatosuchus vjushkovi Last Modified in 08/10/23	675	24	22	0	0	0	0	1
[63] † Koilamasuchus gonzalezdiazi Last Modified in 08/10/23	667	31	28	0	2	0	0	1
[64] † Kalisuchus rewanensis holotype Last Modified in 08/10/23	652	40	35	0	10	0	0	1
[65] † NMQR 3570 Last Modified in 08/10/23	661	27	27	0	9	0	0	1
[66] † Fugusuchus hejiapanensis Last Modified in 08/10/23	479	201	198	0	19	0	0	1
[67] † Sarmatosuchus otschevi Last Modified in 08/10/23	459	238	231	0	7	0	0	1
[68] † Guchengosuchus shiguaiensis Last Modified in 08/10/23	496	184	176	0	25	0	0	1
[69] † Cuyosuchus huenei Last Modified in 08/10/23	511	190	180	0	6	0	0	1
[70] † Garjainia prima Last Modified in 08/10/23	112	553	548	0	36	0	0	1
[71] † Garjainia madiba combined Last Modified in 08/10/23	362	329	313	0	16	0	0	1
[72] † Erythrosuchus africanus Last Modified in 08/10/23	67	591	580	0	43	0	0	1
[73] † Shansisuchus shansisuchus Last Modified in 08/10/23	169	492	486	0	41	0	0	1
[74] † Shansisuchus kuyeheensis Last Modified in 08/10/23	610	82	77	0	10	0	0	1
[75] † Chalishevia cothurnata Last Modified in 08/10/23	599	90	88	0	10	0	0	1
[76] † Youngosuchus sinensis Last Modified in 08/10/23	365	320	313	0	19	0	0	1
[77] † Uralosaurus combined Last Modified in 08/10/23	664	27	27	0	6	0	0	1
[78] † Vancleavea campi Last Modified in 08/10/23	217	426	421	0	57	0	0	1
[79] † Litorosuchus somnii Last Modified in 08/10/23	280	396	390	0	27	0	0	1
[80] † Asperoris mnyama Last Modified in 08/10/23	583	104	92	0	16	0	0	1
[81] † Euparkeria capensis Last Modified in 08/10/23	41	642	630	0	26	0	0	1
[82] † Dorosuchus neoetus Last Modified in 08/10/23	594	102	102	0	1	0	0	1
[83] † Halazhaisuchus qiaoensis Last Modified in 08/10/23	600	92	91	0	6	0	0	1
[84] † Osmolskina czatkoviensis Last Modified in 08/10/23	673	19	19	0	5	0	0	1
[85] † Osmolskina complete Last Modified in 08/10/23	266	419	409	0	22	0	0	1
[86] † Yarasuchus deccanensis Last Modified in 08/10/23	499	191	188	0	9	0	0	1
[87] † Dongusuchus efremovi Last Modified in 08/10/23	666	31	31	0	0	0	0	1
[88] † Teleocrater rhadinus Last Modified in 08/10/23	417	264	263	0	17	0	0	1
[89] † Spondylosoma absconditum Last Modified in 08/10/23	626	69	68	0	3	0	0	1
[90] † Proterochampsa barrionuevoi Last Modified in 08/10/23	215	441	433	0	49	0	0	1
[91] † Proterochampsa nodosa Last Modified in 08/10/23	447	231	225	0	20	0	0	1
[92] † Rhadinosuchus gracilis Last Modified in 08/10/23	587	104	98	0	12	0	0	1
[93] † Tropidosuchus romeri Last Modified in 08/10/23	166	509	499	0	31	0	0	1
[94] † Cerritosaurus binsfeldi Last Modified in 08/10/23	469	211	206	0	22	0	0	1
[95] † Gualosuchus reigi Last Modified in 08/10/23	209	459	449	0	37	0	0	1
[96] † Chanaresuchus bonapartei Last Modified in 08/10/23	60	610	592	0	41	0	0	1
[97] † Pseudochampsa ischigualastensis Last Modified in 08/10/23	291	383	377	0	29	0	0	1
[98] † Archeopelta arborensis Last Modified in 08/10/23	592	104	104	0	1	0	0	1
[99] † Tarjadia ruthae Last Modified in 08/10/23	235	442	438	0	24	0	0	1
[100] † Erpetosuchus granti Last Modified in 08/10/23	374	301	297	0	24	0	0	1
[101] † Erpetosuchus sp Last Modified in 08/10/23	608	85	80	0	9	0	0	1
[102] † Dyoplax arenaceus Last Modified in 08/10/23	646	51	49	0	2	0	0	1
[103] † Jaxtasuchus salomoni Last Modified in 08/10/23	500	189	182	0	15	0	0	1
[104] † Doswellia kaltenbachi Last Modified in 08/10/23	319	336	330	0	48	0	0	1
[105] † Parasuchus angustifrons Last Modified in 08/10/23	443	225	222	0	32	0	0	1
[106] † Parasuchus hislopi Last Modified in 08/10/23	90	578	571	0	34	0	0	1
[107] † Nicrosaurus kapffi Last Modified in 08/10/23	227	437	428	0	40	0	0	1
[108] † Smilosuchus spp. Last Modified in 08/10/23	88	569	559	0	45	0	0	1
[109] † Diandongosuchus fuyuanensis Last Modified in 08/10/23	289	390	388	0	20	0	0	1
[110] † Ornithosuchus woodwardi Last Modified in 08/10/23	212	460	453	0	28	0	0	1
[111] † Riojasuchus tenuisceps Last Modified in 08/10/23	98	571	559	0	34	0	0	1
[112] † Venaticosuchus rusconii Last Modified in 08/10/23	527	155	153	0	17	0	0	1
[113] † Nundasuchus songeaensis Last Modified in 08/10/23	449	241	239	0	9	0	0	1
[114] † Turfanosuchus dabanensis Last Modified in 08/10/23	215	463	456	0	24	0	0	1
[115] † Gracilisuchus stipanicicorum Last Modified in 08/10/23	143	532	520	0	34	0	0	1
[116] † Yonghesuchus sangbiensis Last Modified in 08/10/23	511	171	168	0	18	0	0	1
[117] † Parringtonia gracilis Last Modified in 08/10/23	607	85	80	0	10	0	0	1
[118] † Otter sandstone archosaur Last Modified in 08/10/23	660	38	35	0	2	0	0	1
[119] † Aetosauroides scagliai Last Modified in 08/10/23	269	418	403	0	24	0	0	1
[120] † Aetosaurus ferratus Last Modified in 08/10/23	167	506	501	0	29	0	0	1
[121] † Stagonolepis robertsoni Last Modified in 08/10/23	60	606	606	0	31	0	0	1
[122] † Ticinosuchus ferox Last Modified in 08/10/23	413	274	267	0	16	0	0	1
[123] † Stagonosuchus nyassicus Last Modified in 08/10/23	548	150	144	0	5	0	0	1
[124] † Dagasuchus santacruzensis Last Modified in 08/10/23	672	25	25	0	0	0	0	1
[125] † Qianosuchus mixtus Last Modified in 08/10/23	262	410	408	0	27	0	0	1
[126] † Arizonasaurus babbitti Last Modified in 08/10/23	314	352	347	0	36	0	0	1
[127] † Lotosaurus adentus Last Modified in 08/10/23	267	373	367	0	63	0	0	1
[128] † Ctenosauriscus koeneni Last Modified in 08/10/23	650	31	31	0	16	0	0	1
[129] † Hypselorhachis mirabilis Last Modified in 08/10/23	672	25	24	0	1	0	0	1
[130] † Waldhaus poposauroid Last Modified in 08/10/23	665	33	32	0	0	0	0	1
[131] † Vytshegdosuchus zbeshartensis Last Modified in 08/10/23	690	7	7	0	0	0	0	1
[132] † Bystrowisuchus flerovi Last Modified in 08/10/23	680	19	16	0	1	0	0	1
[133] † Bromsgroveia walkeri Last Modified in 08/10/23	660	37	37	0	0	0	0	1
[134] † Xilousuchus sapingensis Last Modified in 08/10/23	492	188	187	0	17	0	0	1
[135] † Moenkopi poposauroid Last Modified in 08/10/23	659	38	34	0	4	0	0	1
[136] † Mandasuchus tanyauchen Last Modified in 08/10/23	442	246	241	0	14	0	0	1
[137] † Luperosuchus fractus Last Modified in 08/10/23	583	113	111	0	3	0	0	1
[138] † Decuriasuchus quartacolonia Last Modified in 08/10/23	316	358	353	0	28	0	0	1
[139] † Batrachotomus kupferzellensis Last Modified in 08/10/23	128	544	535	0	30	0	0	1
[140] † Prestosuchus chiniquensis Last Modified in 08/10/23	71	602	596	0	30	0	0	1
[141] † Scleromochlus taylori Last Modified in 08/10/23	524	155	144	0	29	0	0	1
[142] † Dimorphodon macronyx Last Modified in 08/10/23	329	318	312	0	56	0	0	1
[143] † Lagerpeton chanarensis Last Modified in 08/10/23	538	143	139	0	18	0	0	1
[144] † Marasuchus lilloensis Last Modified in 08/10/23	345	327	321	0	31	0	0	1
[145] † Lutungutali sitwensis Last Modified in 08/10/23	666	35	30	0	1	0	0	1
[146] † Asilisaurus kongwe Last Modified in 08/10/23	459	222	220	0	18	0	0	1
[147] † Nyasasaurus parringtoni Last Modified in 08/10/23	652	44	43	0	2	0	0	1
[148] † Lewisuchus admixtus Last Modified in 08/10/23	273	408	395	0	25	0	0	1
[149] † Silesaurus opolensis Last Modified in 08/10/23	127	529	519	0	50	0	0	1
[150] † Heterodontosaurus tucki Last Modified in 08/10/23	74	548	542	0	77	0	0	1
[151] † Herrerasaurus ischigualastensis Last Modified in 08/10/23	68	587	575	0	54	0	0	1

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	2947
Project overview	163
Matrix views	112	Matrix landing page (106 views); Data matrix provided by author (6 views);
Documents list	107
Media views	458	Media search (386 views); M890604 (72 views);
Bibliography	86
Views for media list	137
Specimen list	238
Taxon list	1646