Project 5130: C. Hendrickx, T. H. Trapman, S. Wills, F. M. Holwerda, K. H. Stein, O. W. Rauhut, R. R. Melzer, J. V. Woensel, J. W. Reumer. 2024. A combined approach to identify isolated theropod teeth from the Cenomanian Kem Kem Group of Morocco: cladistic, discriminant, and machine learning analyses. Journal of Vertebrate Paleontology. Not_assigned_yet:1-23.
Specimen: † Coelophysis (unvouchered)
View: dentition

Abstract

The Kem Kem Group of Southeastern Morocco, North Africa, is well known for theropod remains, especially isolated teeth. Here, a collection of isolated theropod teeth is assessed for diversity using a combination of linear discriminant, phylogenetic, and machine learning analyses for the first time. The results confirm earlier studies on Kem Kem theropod diversity, with teeth referred to Abelisauridae, Spinosaurinae, and Carcharodontosauridae. A single tooth is ascribed to a non-abelisauroid ceratosaur or a megaraptoran and may represent the enigmatic averostran Deltadromeus. Spinosaurine teeth are clearly differentiated by all three methodologies, whereas abelisaurid and carcharodontosaurid teeth could only be distinguished by the machine learning and phylogenetic analyses. This study shows that a combination of independent methods is most effective at providing strong evidence on theropod dental diversity in a particular assemblage, and that cladistic and machine learning analyses are the most reliable approaches to identify isolated dinosaur teeth. The methodology used here is likely to yield results in other dinosaur assemblages where isolated teeth are more abundant than body fossils.


Read the article »

Article DOI: 10.1080/02724634.2024.2311791

Project DOI: 10.7934/P5130, http://dx.doi.org/10.7934/P5130
This project containsMatrices
  • 1 Media
  • 2 Matrices
  • 115 Taxa
  • 1 Specimen
  • 146 Characters
Total size of project's media files: 1.7M

Download Project SDD File
Total scored cells: 21835
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 237
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 423
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:215/22
Currently Viewing:
MorphoBank Project 5130
  • Creation Date:
    15 March 2024
  • Publication Date:
    15 March 2024
  • Project views: 921

    Authors' Institutions

    • Paläontologisches Museum München (Palaeontological Museum, Munich)

    • Natuurhistorisch Museum Rotterdam (Natural History Museum, Rotterdam)

    • Natural History Museum, London

    • Royal Tyrrell Museum of Palaeontology, Drumheller

    • Katholieke Universiteit Leuven

    • Universiteit Utrecht

    • Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)

    • Naturalis Biodiversity Center

    • Friedrich-Alexander-Universität Erlangen-Nürnberg

    • Bavarian State Collection of Zoology

    • Vrije Universiteit Brussel

    • Birkbeck College, London



    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    medialabels
    rules
    MorphoBank Curator
    Project Administrator
    115112370021835
    (17656, 0, 4179)
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    Taxonomic Overview for Matrix 'M29710' (110 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] † Herrerasaurus
    Last Modified in 03/15/24
    127164015001
    [2] † Daemonosaurus
    Last Modified in 03/15/24
    15706808001
    [3] † Eodromaeus
    Last Modified in 03/15/24
    36555005001
    [4] † Eoraptor
    Last Modified in 03/15/24
    8877508001
    [5] † Dracovenator
    Last Modified in 03/15/24
    14717106001
    [6] † Coelophysis
    Last Modified in 03/15/24
    88473010001
    [7] † Liliensternus
    Last Modified in 03/15/24
    6847609001
    [8] † Dilophosaurus
    Last Modified in 03/15/24
    5847907001
    [9] † Ceratosaurus
    Last Modified in 03/15/24
    11158703001
    [10] † Genyodectes
    Last Modified in 03/15/24
    7897707001
    [11] † Berberosaurus
    Last Modified in 03/15/24
    41474406001
    [12] † Limusaurus juvenile
    Last Modified in 03/15/24
    152623053001
    [13] † Noasaurus
    Last Modified in 03/15/24
    38474607001
    [14] † Masiakasaurus
    Last Modified in 03/15/24
    01018704001
    [15] † Kryptops
    Last Modified in 03/15/24
    39474507001
    [16] † Rugops
    Last Modified in 03/15/24
    31565307001
    [17] † Abelisaurus
    Last Modified in 03/15/24
    9867804001
    [18] † Aucasaurus
    Last Modified in 03/15/24
    33585206001
    [19] † Arcovenator
    Last Modified in 03/15/24
    34565007001
    [20] † Chenanisaurus
    Last Modified in 03/15/24
    10847605001
    [21] † Indosuchus
    Last Modified in 03/15/24
    27675608001
    [22] † Majungasaurus
    Last Modified in 03/15/24
    31018503001
    [23] † Skorpiovenator
    Last Modified in 03/15/24
    11787505001
    [24] † Chilesaurus
    Last Modified in 03/15/24
    184846027001
    [25] † Piatnitzkysaurus
    Last Modified in 03/15/24
    29736002001
    [26] † Marshosaurus
    Last Modified in 03/15/24
    3918206001
    [27] † Monolophosaurus
    Last Modified in 03/15/24
    11897703001
    [28] † Sciuruminus
    Last Modified in 03/15/24
    175246028001
    [29] † Eustreptospondylus
    Last Modified in 03/15/24
    22646207001
    [30] † Afrovenator
    Last Modified in 03/15/24
    33575404001
    [31] † Dubreuillosaurus
    Last Modified in 03/15/24
    2888108001
    [32] † Duriavenator
    Last Modified in 03/15/24
    11817307001
    [33] † Megalosaurus
    Last Modified in 03/15/24
    35665303001
    [34] † Torvosaurus
    Last Modified in 03/15/24
    10907803001
    [35] † Baryonyx
    Last Modified in 03/15/24
    2948603001
    [36] † Suchomimus
    Last Modified in 03/15/24
    11917703001
    [37] † Irritator angaturama
    Last Modified in 03/15/24
    353230026001
    [38] † Spinosaurus
    Last Modified in 03/15/24
    45750037001
    [39] † Erectopus
    Last Modified in 03/15/24
    35535006001
    [40] † Yangchuanosaurus
    Last Modified in 03/15/24
    63292602001
    [41] † Sinraptor dongi
    Last Modified in 03/15/24
    11108604001
    [42] † Sinraptor hepingensis
    Last Modified in 03/15/24
    36575104001
    [43] † Allosaurus
    Last Modified in 03/15/24
    01288803001
    [44] † Neovenator
    Last Modified in 03/15/24
    29645903001
    [45] † Fukuiraptor
    Last Modified in 03/15/24
    17847103001
    [46] † Australovenator
    Last Modified in 03/15/24
    29675804001
    [47] † Megaraptor
    Last Modified in 03/15/24
    96561021001
    [48] † Orkoraptor
    Last Modified in 03/15/24
    403733018001
    [49] † Acrocanthosaurus
    Last Modified in 03/15/24
    7998103001
    [50] † Eocarcharia
    Last Modified in 03/15/24
    35565006001
    [51] † Carcharodontosaurus
    Last Modified in 03/15/24
    29706101001
    [52] † Giganotosaurus
    Last Modified in 03/15/24
    3998602001
    [53] † Mapusaurus
    Last Modified in 03/15/24
    5998303001
    [54] † Bicentenaria
    Last Modified in 03/15/24
    19676606001
    [55] † Zuolong
    Last Modified in 03/15/24
    20726605001
    [56] † Proceratosaurus
    Last Modified in 03/15/24
    8887607001
    [57] † Guanlong
    Last Modified in 03/15/24
    8917607001
    [58] † Dilong
    Last Modified in 03/15/24
    11108406001
    [59] † Eotyrannus
    Last Modified in 03/15/24
    7817608001
    [60] † Raptorex
    Last Modified in 03/15/24
    6897906001
    [61] † Gorgosaurus
    Last Modified in 03/15/24
    11098406001
    [62] † Alioramus
    Last Modified in 03/15/24
    32625405001
    [63] † Daspletosaurus
    Last Modified in 03/15/24
    01178803001
    [64] † Tyrannosaurus
    Last Modified in 03/15/24
    11088703001
    [65] † Compsognathus
    Last Modified in 03/15/24
    65649036001
    [66] † Juravenator
    Last Modified in 03/15/24
    105851030001
    [67] † Scipionyx
    Last Modified in 03/15/24
    154540036001
    [68] † Ornitholestes
    Last Modified in 03/15/24
    67461024001
    [69] † Nqwebasaurus
    Last Modified in 03/15/24
    351818038001
    [70] † Pelecanimimus
    Last Modified in 03/15/24
    83531052001
    [71] † Shenzhousaurus
    Last Modified in 03/15/24
    73030054001
    [72] † Aorun
    Last Modified in 03/15/24
    154439037001
    [73] † Haplocheirus
    Last Modified in 03/15/24
    136458020001
    [74] † Shuvuuia
    Last Modified in 03/15/24
    63333052001
    [75] † Mononykus
    Last Modified in 03/15/24
    332020038001
    [76] † Eshanosaurus
    Last Modified in 03/15/24
    32595108001
    [77] † Falcarius
    Last Modified in 03/15/24
    08771020001
    [78] † Jianchangosaurus
    Last Modified in 03/15/24
    3828008001
    [79] † Segnosaurus
    Last Modified in 03/15/24
    34685007001
    [80] † Erlikosaurus
    Last Modified in 03/15/24
    57371015001
    [81] † Incisivosaurus
    Last Modified in 03/15/24
    265449016001
    [82] † Caudipteryx
    Last Modified in 03/15/24
    22218071001
    [83] † Halszkaraptor
    Last Modified in 03/15/24
    162827048001
    [84] † Buitreraptor
    Last Modified in 03/15/24
    312621039001
    [85] † Microraptor
    Last Modified in 03/15/24
    76652032001
    [86] † Sinornithosaurus
    Last Modified in 03/15/24
    61007906001
    [87] † Graciliraptor
    Last Modified in 03/15/24
    40514506001
    [88] † Dromaeosaurus
    Last Modified in 03/15/24
    6937906001
    [89] † Atrociraptor
    Last Modified in 03/15/24
    21038306001
    [90] † Bambiraptor
    Last Modified in 03/15/24
    36544807001
    [91] † Tsaagan
    Last Modified in 03/15/24
    95753029001
    [92] † Velociraptor
    Last Modified in 03/15/24
    97266016001
    [93] † Deinonychus
    Last Modified in 03/15/24
    1998406001
    [94] † Saurornitholestes
    Last Modified in 03/15/24
    01028506001
    [95] † MPC d100 1128
    Last Modified in 03/15/24
    34036052001
    [96] † Almas
    Last Modified in 03/15/24
    63633052001
    [97] † Sinusonasus
    Last Modified in 03/15/24
    135343035001
    [98] † Byronosaurus
    Last Modified in 03/15/24
    74433051001
    [99] † Zanabazar
    Last Modified in 03/15/24
    18746508001
    [100] † Troodon
    Last Modified in 03/15/24
    2998405001
    [101] † Epidexipteryx
    Last Modified in 03/15/24
    73330054001
    [102] † Archaeopteryx
    Last Modified in 03/15/24
    63533052001
    [103] † Kem-Morpho1a
    Last Modified in 03/15/24
    64292601001
    [104] † Kem-Morpho1b
    Last Modified in 03/15/24
    37584707001
    [105] † Kem-Morpho1
    Last Modified in 03/15/24
    12867207001
    [106] † Kem-Morpho2
    Last Modified in 03/15/24
    374025029001
    [107] † Kem-Morpho3a
    Last Modified in 03/15/24
    63372800001
    [108] † Kem-Morpho3b
    Last Modified in 03/15/24
    40644803001
    [109] † Kem-Morpho3
    Last Modified in 03/15/24
    131007503001
    [110] † Kem-Morpho4
    Last Modified in 03/15/24
    44424007001


    Taxonomic Overview for Matrix 'M29711' (115 Taxa)

    taxon unscored
    cells
    scored
    cells
    no cell
    support
    NPA
    cells
    "-" cellscell images labels on
    cell images
    member
    access
    [1] † Herrerasaurus
    Last Modified in 03/15/24
    30108101015001
    [2] † Daemonosaurus
    Last Modified in 03/15/24
    3810210008001
    [3] † Eoraptor
    Last Modified in 03/15/24
    3711310108001
    [4] † Eodromaeus
    Last Modified in 03/15/24
    70767105001
    [5] † Coelophysis
    Last Modified in 03/15/24
    12124113021001
    [6] † Liliensternus
    Last Modified in 03/15/24
    47989009001
    [7] † Dracovenator
    Last Modified in 03/15/24
    50909006001
    [8] † Dilophosaurus
    Last Modified in 03/15/24
    2412011507001
    [9] † Ceratosaurus
    Last Modified in 03/15/24
    117114104001
    [10] † Genyodectes
    Last Modified in 03/15/24
    2912211007001
    [11] † Berberosaurus
    Last Modified in 03/15/24
    96474406001
    [12] † Limusaurus juvenile
    Last Modified in 03/15/24
    335750063001
    [13] † Limusaurus adult
    Last Modified in 03/15/24
    0440142001
    [14] † Noasaurus
    Last Modified in 03/15/24
    85555407001
    [15] † Masiakasaurus
    Last Modified in 03/15/24
    3412210804001
    [16] † Kryptops
    Last Modified in 03/15/24
    89525007001
    [17] † Rugops
    Last Modified in 03/15/24
    64787507001
    [18] † Abelisaurus
    Last Modified in 03/15/24
    52989004001
    [19] † Arcovenator
    Last Modified in 03/15/24
    89565007001
    [20] † Chenanisaurus
    Last Modified in 03/15/24
    61888005001
    [21] † Indosuchus
    Last Modified in 03/15/24
    481019008001
    [22] † Majungasaurus
    Last Modified in 03/15/24
    415613903001
    [23] † Aucasaurus
    Last Modified in 03/15/24
    68787206001
    [24] † Skorpiovenator
    Last Modified in 03/15/24
    52928905001
    [25] † Chilesaurus
    Last Modified in 03/15/24
    606058028001
    [26] † Piatnitzkysaurus
    Last Modified in 03/15/24
    71867302001
    [27] † Marshosaurus
    Last Modified in 03/15/24
    2312611706001
    [28] † Monolophosaurus
    Last Modified in 03/15/24
    3611910703001
    [29] † Sciuruminus
    Last Modified in 03/15/24
    428276028001
    [30] † Eustreptospondylus
    Last Modified in 03/15/24
    48939107001
    [31] † Afrovenator
    Last Modified in 03/15/24
    79666304001
    [32] † Dubreuillosaurus
    Last Modified in 03/15/24
    2012511808001
    [33] † Duriavenator
    Last Modified in 03/15/24
    421059707001
    [34] † Megalosaurus
    Last Modified in 03/15/24
    70867303001
    [35] † Torvosaurus
    Last Modified in 03/15/24
    3012411204001
    [36] † Baryonyx
    Last Modified in 03/15/24
    2213012103001
    [37] † Suchomimus
    Last Modified in 03/15/24
    3012711303001
    [38] † Irritator angaturama
    Last Modified in 03/15/24
    685452026001
    [39] † Spinosaurus
    Last Modified in 03/15/24
    229687037001
    [40] † Erectopus
    Last Modified in 03/15/24
    87565306001
    [41] † Yangchuanosaurus
    Last Modified in 03/15/24
    85655902001
    [42] † Sinraptor dongi
    Last Modified in 03/15/24
    2014812204001
    [43] † Sinraptor hepingensis
    Last Modified in 03/15/24
    57928504001
    [44] † Allosaurus
    Last Modified in 03/15/24
    018514303001
    [45] † Neovenator
    Last Modified in 03/15/24
    49999403001
    [46] † Fukuiraptor
    Last Modified in 03/15/24
    63938003001
    [47] † Australovenator
    Last Modified in 03/15/24
    75766704001
    [48] † Megaraptor
    Last Modified in 03/15/24
    468479021001
    [49] † Orkoraptor
    Last Modified in 03/15/24
    953733018001
    [50] † Acrocanthosaurus
    Last Modified in 03/15/24
    2513611803001
    [51] † Eocarcharia
    Last Modified in 03/15/24
    83635706001
    [52] † Carcharodontosaurus
    Last Modified in 03/15/24
    68867701001
    [53] † Giganotosaurus
    Last Modified in 03/15/24
    2713011702001
    [54] † Mapusaurus
    Last Modified in 03/15/24
    461139703001
    [55] † Bicentenaria
    Last Modified in 03/15/24
    66757406001
    [56] † Zuolong
    Last Modified in 03/15/24
    53948805001
    [57] † Proceratosaurus
    Last Modified in 03/15/24
    2712411207001
    [58] † Guanlong
    Last Modified in 03/15/24
    401149907001
    [59] † Dilong
    Last Modified in 03/15/24
    2214511806001
    [60] † Eotyrannus
    Last Modified in 03/15/24
    3111210708001
    [61] † Raptorex
    Last Modified in 03/15/24
    2412611606001
    [62] † Gorgosaurus
    Last Modified in 03/15/24
    116513906001
    [63] † Alioramus
    Last Modified in 03/15/24
    67827405001
    [64] † Daspletosaurus
    Last Modified in 03/15/24
    017214006001
    [65] † Tyrannosaurus
    Last Modified in 03/15/24
    115913807001
    [66] † Compsognathus
    Last Modified in 03/15/24
    338477036001
    [67] † Juravenator
    Last Modified in 03/15/24
    457871030001
    [68] † Scipionyx
    Last Modified in 03/15/24
    367873037001
    [69] † Ornitholestes
    Last Modified in 03/15/24
    2511197024001
    [70] † Nqwebasaurus
    Last Modified in 03/15/24
    852323038001
    [71] † Pelecanimimus
    Last Modified in 03/15/24
    405854052001
    [72] † Shenzhousaurus
    Last Modified in 03/15/24
    264040080001
    [73] † Garudimimus
    Last Modified in 03/15/24
    0440142001
    [74] † Struthiomimus
    Last Modified in 03/15/24
    0440142001
    [75] † Aorun
    Last Modified in 03/15/24
    397570037001
    [76] † Haplocheirus
    Last Modified in 03/15/24
    369790020001
    [77] † Shuvuuia
    Last Modified in 03/15/24
    425252052001
    [78] † Mononykus
    Last Modified in 03/15/24
    882020038001
    [79] † Eshanosaurus
    Last Modified in 03/15/24
    80665808001
    [80] † Falcarius
    Last Modified in 03/15/24
    2311698025001
    [81] † Jianchangosaurus
    Last Modified in 03/15/24
    259896025001
    [82] † Segnosaurus
    Last Modified in 03/15/24
    83745607001
    [83] † Erlikosaurus
    Last Modified in 03/15/24
    288987031001
    [84] † Incisivosaurus
    Last Modified in 03/15/24
    459185016001
    [85] † Caudipteryx
    Last Modified in 03/15/24
    293127090001
    [86] † Chirostenotes
    Last Modified in 03/15/24
    0440142001
    [87] † Citipati
    Last Modified in 03/15/24
    0440142001
    [88] † Halszkaraptor
    Last Modified in 03/15/24
    415857048001
    [89] † Buitreraptor
    Last Modified in 03/15/24
    753732039001
    [90] † Microraptor
    Last Modified in 03/15/24
    3010084032001
    [91] † Sinornithosaurus
    Last Modified in 03/15/24
    2813311206001
    [92] † Graciliraptor
    Last Modified in 03/15/24
    91554906001
    [93] † Dromaeosaurus
    Last Modified in 03/15/24
    13136122011001
    [94] † Atrociraptor
    Last Modified in 03/15/24
    2313711706001
    [95] † Bambiraptor
    Last Modified in 03/15/24
    508579017001
    [96] † Saurornitholestes
    Last Modified in 03/15/24
    17135117012001
    [97] † Tsaagan
    Last Modified in 03/15/24
    309187029001
    [98] † Velociraptor
    Last Modified in 03/15/24
    13119111022001
    [99] † Deinonychus
    Last Modified in 03/15/24
    3147132011001
    [100] † MPC d100 1128
    Last Modified in 03/15/24
    405854052001
    [101] † Almas
    Last Modified in 03/15/24
    137471062001
    [102] † Sinusonasus
    Last Modified in 03/15/24
    457766035001
    [103] † Byronosaurus
    Last Modified in 03/15/24
    118574061001
    [104] † Zanabazar
    Last Modified in 03/15/24
    3611210208001
    [105] † Troodon
    Last Modified in 03/15/24
    27123108011001
    [106] † Epidexipteryx
    Last Modified in 03/15/24
    326158056001
    [107] † Archaeopteryx
    Last Modified in 03/15/24
    277167052001
    [108] † Kem-Morpho1a
    Last Modified in 03/15/24
    119292601001
    [109] † Kem-Morpho1b
    Last Modified in 03/15/24
    92584707001
    [110] † Kem-Morpho1
    Last Modified in 03/15/24
    67867207001
    [111] † Kem-Morpho2
    Last Modified in 03/15/24
    924025029001
    [112] † Kem-Morpho3a
    Last Modified in 03/15/24
    118372800001
    [113] † Kem-Morpho3b
    Last Modified in 03/15/24
    95644803001
    [114] † Kem-Morpho3
    Last Modified in 03/15/24
    681007503001
    [115] † Kem-Morpho4
    Last Modified in 03/15/24
    99424007001


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views921
    Project overview71
    Matrix views54Matrix landing page (42 views); KemKem Dentition data matrix (6 views); KemKem Crown Data Matrix (6 views);
    Media views168Media search (145 views); M902662 (23 views);
    Specimen list72
    Views for media list51
    Bibliography37
    Taxon list468