Project 5137: E. Martin-Silverstone, D. M. Unwin, A. R. Cuff, E. E. Brown, L. Allington-Jones, P. M. Barrett. 2024. A new pterosaur from the Middle Jurassic of Skye, Scotland and the early diversification of flying reptiles. Journal of Vertebrate Paleontology. null:null.
Abstract
The Middle Jurassic was a critical time in pterosaur evolution, witnessing the appearance of major morphological innovations that underpinned successive radiations by rhamphorhynchids, basally branching monofenestratans, and pterodactyloids. Frustratingly, this interval is particularly sparsely sampled, with a record consisting almost exclusively of isolated fragmentary remains. Here, we describe new material from the Bathonian-aged Kilmaluag Formation of Skye, Scotland, which helps close this gap. Ceoptera evansae (gen. et sp. nov.) is based on a three-dimensionally preserved partial skeleton, which represents one of the only associated Middle Jurassic pterosaurs. Ceoptera is among the first pterosaurs to be fully digitally prepared, and µCT scanning reveals multiple elements of the skeleton that remain fully embedded within the matrix and otherwise inaccessible. It is diagnosed by unique features of the pectoral and pelvic girdle. The inclusion of this new Middle Jurassic pterosaur in a novel phylogenetic analysis of pterosaur interrelationships provides additional support for the existence of the controversial clade Darwinoptera, adding to our knowledge of pterosaur diversity and evolution.Read the article »
Article DOI: 10.1080/02724634.2023.2298741
Project DOI: 10.7934/P5137, http://dx.doi.org/10.7934/P5137
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 6415 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 136 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 338 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:111/25 |
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MorphoBank Project 5137
MorphoBank Project 5137
- Creation Date:
18 March 2024 - Publication Date:
18 March 2024
Authors' Institutions
- Natural History Museum, London
- University of Birmingham
- University of Leicester
- University of Liverpool
- University of Bristol
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 69 | 1 | 1 | 136 | 0 | 0 | 6415 (6415, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29717' (69 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Euparkeria Taxon name last Modified on 03/18/24 | 10 | 126 | 126 | 0 | 0 | 0 | 0 | 1 |
[2] † Herrerasaurus Taxon name last Modified on 03/18/24 | 10 | 126 | 126 | 0 | 0 | 0 | 0 | 1 |
[3] † Preondactylus Taxon name last Modified on 03/18/24 | 31 | 105 | 105 | 0 | 0 | 0 | 0 | 1 |
[4] † Dimorphodon Taxon name last Modified on 03/18/24 | 13 | 124 | 123 | 0 | 0 | 0 | 0 | 1 |
[5] † Peteinosaurus Taxon name last Modified on 03/18/24 | 57 | 79 | 79 | 0 | 0 | 0 | 0 | 1 |
[6] † Anurognathus Taxon name last Modified on 03/18/24 | 23 | 113 | 113 | 0 | 0 | 0 | 0 | 1 |
[7] † Batrachognathus Taxon name last Modified on 03/18/24 | 41 | 95 | 95 | 0 | 0 | 0 | 0 | 1 |
[8] † Dendrorhynchoides Taxon name last Modified on 03/18/24 | 52 | 85 | 84 | 0 | 0 | 0 | 0 | 1 |
[9] † Jeholopterus Taxon name last Modified on 03/18/24 | 45 | 91 | 91 | 0 | 0 | 0 | 0 | 1 |
[10] † Austriadactylus Taxon name last Modified on 03/18/24 | 58 | 78 | 78 | 0 | 0 | 0 | 0 | 1 |
[11] † Campylognathoides Taxon name last Modified on 03/18/24 | 4 | 133 | 132 | 0 | 0 | 0 | 0 | 1 |
[12] † Eudimorphodon Taxon name last Modified on 03/18/24 | 11 | 131 | 125 | 0 | 0 | 0 | 0 | 1 |
[13] † Raeticodactylus Taxon name last Modified on 03/18/24 | 42 | 94 | 94 | 0 | 0 | 0 | 0 | 1 |
[14] † Scaphognathus Taxon name last Modified on 03/18/24 | 8 | 128 | 128 | 0 | 0 | 0 | 0 | 1 |
[15] † Sordes Taxon name last Modified on 03/18/24 | 14 | 123 | 122 | 0 | 0 | 0 | 0 | 1 |
[16] † Pterorhynchus Taxon name last Modified on 03/18/24 | 50 | 86 | 86 | 0 | 0 | 0 | 0 | 1 |
[17] † Cacibupteryx Taxon name last Modified on 03/18/24 | 90 | 46 | 46 | 0 | 0 | 0 | 0 | 1 |
[18] † Dorygnathus Taxon name last Modified on 03/18/24 | 5 | 131 | 131 | 0 | 0 | 0 | 0 | 1 |
[19] † Nesodactylus Taxon name last Modified on 03/18/24 | 94 | 42 | 42 | 0 | 0 | 0 | 0 | 1 |
[20] † Dearc Taxon name last Modified on 03/18/24 | 60 | 76 | 76 | 0 | 0 | 0 | 0 | 1 |
[21] † Rhamphorhynchus Taxon name last Modified on 03/18/24 | 1 | 136 | 135 | 0 | 0 | 0 | 0 | 1 |
[22] † Ceoptera Taxon name last Modified on 03/18/24 | 113 | 23 | 23 | 0 | 0 | 0 | 0 | 1 |
[23] † Allkaruen Taxon name last Modified on 03/18/24 | 81 | 55 | 55 | 0 | 0 | 0 | 0 | 1 |
[24] † Cuspicephalus Taxon name last Modified on 03/18/24 | 93 | 43 | 43 | 0 | 0 | 0 | 0 | 1 |
[25] † Kryptodrakon Taxon name last Modified on 03/18/24 | 123 | 13 | 13 | 0 | 0 | 0 | 0 | 1 |
[26] † Darwinopterus Taxon name last Modified on 03/18/24 | 6 | 136 | 130 | 0 | 0 | 0 | 0 | 1 |
[27] † Wukongopterus Taxon name last Modified on 03/18/24 | 57 | 79 | 79 | 0 | 0 | 0 | 0 | 1 |
[28] † Kunpengopterus Taxon name last Modified on 03/18/24 | 9 | 128 | 127 | 0 | 0 | 0 | 0 | 1 |
[29] † Changchengopterus Taxon name last Modified on 03/18/24 | 81 | 55 | 55 | 0 | 0 | 0 | 0 | 1 |
[30] † Douzhanopterus Taxon name last Modified on 03/18/24 | 94 | 42 | 42 | 0 | 0 | 0 | 0 | 1 |
[31] † Rhamphodactylus Taxon name last Modified on 03/18/24 | 7 | 130 | 129 | 0 | 0 | 0 | 0 | 1 |
[32] † Liaodactylus Taxon name last Modified on 03/18/24 | 73 | 63 | 63 | 0 | 0 | 0 | 0 | 1 |
[33] † Pterodactylus Taxon name last Modified on 03/18/24 | 5 | 134 | 131 | 0 | 0 | 0 | 0 | 1 |
[34] † Cycnorhamphus Taxon name last Modified on 03/18/24 | 15 | 121 | 121 | 0 | 0 | 0 | 0 | 1 |
[35] † Eosipterus Taxon name last Modified on 03/18/24 | 87 | 49 | 49 | 0 | 0 | 0 | 0 | 1 |
[36] † Aurorazhdarcho Taxon name last Modified on 03/18/24 | 10 | 127 | 126 | 0 | 0 | 0 | 0 | 1 |
[37] † Gnathosaurus Taxon name last Modified on 03/18/24 | 66 | 70 | 70 | 0 | 0 | 0 | 0 | 1 |
[38] † Gegepterus Taxon name last Modified on 03/18/24 | 50 | 86 | 86 | 0 | 0 | 0 | 0 | 1 |
[39] † Gladocephaloideus Taxon name last Modified on 03/18/24 | 25 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[40] † Ctenochasma Taxon name last Modified on 03/18/24 | 18 | 120 | 118 | 0 | 0 | 0 | 0 | 1 |
[41] † Pterodaustro Taxon name last Modified on 03/18/24 | 6 | 130 | 130 | 0 | 0 | 0 | 0 | 1 |
[42] † Istiodactylus Taxon name last Modified on 03/18/24 | 27 | 110 | 109 | 0 | 0 | 0 | 0 | 1 |
[43] † Nurhachius Taxon name last Modified on 03/18/24 | 34 | 102 | 102 | 0 | 0 | 0 | 0 | 1 |
[44] † Haopterus Taxon name last Modified on 03/18/24 | 53 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[45] † Ornithocheirus Taxon name last Modified on 03/18/24 | 73 | 63 | 63 | 0 | 0 | 0 | 0 | 1 |
[46] † Anhanguera Taxon name last Modified on 03/18/24 | 10 | 128 | 126 | 0 | 0 | 0 | 0 | 1 |
[47] † Coloborhynchus Taxon name last Modified on 03/18/24 | 12 | 124 | 124 | 0 | 0 | 0 | 0 | 1 |
[48] † Ludodactylus Taxon name last Modified on 03/18/24 | 74 | 62 | 62 | 0 | 0 | 0 | 0 | 1 |
[49] † Boreopterus Taxon name last Modified on 03/18/24 | 36 | 101 | 100 | 0 | 0 | 0 | 0 | 1 |
[50] † Nyctosaurus Taxon name last Modified on 03/18/24 | 24 | 113 | 112 | 0 | 0 | 0 | 0 | 1 |
[51] † Muzquizopteryx Taxon name last Modified on 03/18/24 | 68 | 68 | 68 | 0 | 0 | 0 | 0 | 1 |
[52] † Pteranodon Taxon name last Modified on 03/18/24 | 21 | 115 | 115 | 0 | 0 | 0 | 0 | 1 |
[53] † Germanodactyluscrist Taxon name last Modified on 03/18/24 | 21 | 116 | 115 | 0 | 0 | 0 | 0 | 1 |
[54] † Germanodactylusrhamph Taxon name last Modified on 03/18/24 | 17 | 119 | 119 | 0 | 0 | 0 | 0 | 1 |
[55] † Dsungaripterus Taxon name last Modified on 03/18/24 | 26 | 110 | 110 | 0 | 0 | 0 | 0 | 1 |
[56] † Tatalpterosaur Taxon name last Modified on 03/18/24 | 37 | 99 | 99 | 0 | 0 | 0 | 0 | 1 |
[57] † Noripterus Taxon name last Modified on 03/18/24 | 87 | 49 | 49 | 0 | 0 | 0 | 0 | 1 |
[58] † Lonchodectes Taxon name last Modified on 03/18/24 | 86 | 52 | 50 | 0 | 0 | 0 | 0 | 1 |
[59] † Tapejara Taxon name last Modified on 03/18/24 | 36 | 100 | 100 | 0 | 0 | 0 | 0 | 1 |
[60] † Tupandactylus Taxon name last Modified on 03/18/24 | 29 | 107 | 107 | 0 | 0 | 0 | 0 | 1 |
[61] † Sinopterus Taxon name last Modified on 03/18/24 | 37 | 100 | 99 | 0 | 0 | 0 | 0 | 1 |
[62] † Huaxiapterus Taxon name last Modified on 03/18/24 | 45 | 92 | 91 | 0 | 0 | 0 | 0 | 1 |
[63] † Shenzhoupterus Taxon name last Modified on 03/18/24 | 53 | 83 | 83 | 0 | 0 | 0 | 0 | 1 |
[64] † Chaoyangopterus Taxon name last Modified on 03/18/24 | 70 | 66 | 66 | 0 | 0 | 0 | 0 | 1 |
[65] † Tupuxuara Taxon name last Modified on 03/18/24 | 25 | 111 | 111 | 0 | 0 | 0 | 0 | 1 |
[66] † Thalassodromeus Taxon name last Modified on 03/18/24 | 89 | 47 | 47 | 0 | 0 | 0 | 0 | 1 |
[67] † Quetzalcoatlus Taxon name last Modified on 03/18/24 | 43 | 93 | 93 | 0 | 0 | 0 | 0 | 1 |
[68] † Zhejiangopterus Taxon name last Modified on 03/18/24 | 40 | 96 | 96 | 0 | 0 | 0 | 0 | 1 |
[69] † Azhdarcho Taxon name last Modified on 03/18/24 | 93 | 43 | 43 | 0 | 0 | 0 | 0 | 1 |