Morphobank

Project 5137: E. Martin-Silverstone, D. M. Unwin, A. R. Cuff, E. E. Brown, L. Allington-Jones, P. M. Barrett. 2024. A new pterosaur from the Middle Jurassic of Skye, Scotland and the early diversification of flying reptiles. Journal of Vertebrate Paleontology. null:null.

Specimen: † Rhamphorhynchus (unvouchered)
View: skeleton

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Abstract

The Middle Jurassic was a critical time in pterosaur evolution, witnessing the appearance of major morphological innovations that underpinned successive radiations by rhamphorhynchids, basally branching monofenestratans, and pterodactyloids. Frustratingly, this interval is particularly sparsely sampled, with a record consisting almost exclusively of isolated fragmentary remains. Here, we describe new material from the Bathonian-aged Kilmaluag Formation of Skye, Scotland, which helps close this gap. Ceoptera evansae (gen. et sp. nov.) is based on a three-dimensionally preserved partial skeleton, which represents one of the only associated Middle Jurassic pterosaurs. Ceoptera is among the first pterosaurs to be fully digitally prepared, and µCT scanning reveals multiple elements of the skeleton that remain fully embedded within the matrix and otherwise inaccessible. It is diagnosed by unique features of the pectoral and pelvic girdle. The inclusion of this new Middle Jurassic pterosaur in a novel phylogenetic analysis of pterosaur interrelationships provides additional support for the existence of the controversial clade Darwinoptera, adding to our knowledge of pterosaur diversity and evolution.

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Article DOI: 10.1080/02724634.2023.2298741

Project DOI: 10.7934/P5137, http://dx.doi.org/10.7934/P5137

This project contains	Matrices
1 Media 1 Matrix 69 Taxa 1 Specimen 136 Characters Total size of project's media files: 11.82M Download Project SDD File	Total scored cells: 6415 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 136 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 338 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:111/25

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MorphoBank Project 5137

Creation Date:
18 March 2024
Publication Date:
18 March 2024
Project views: 712

Authors' Institutions

Natural History Museum, London

University of Birmingham

University of Leicester

University of Liverpool

University of Bristol

Members

MorphoBank Curator
Project Administrator

136

6415
(6415, 0, 0)

Taxonomic Overview for Matrix 'M29717' (69 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Euparkeria Last Modified in 03/18/24	10	126	126	0	0	0	0	1
[2] † Herrerasaurus Last Modified in 03/18/24	10	126	126	0	0	0	0	1
[3] † Preondactylus Last Modified in 03/18/24	31	105	105	0	0	0	0	1
[4] † Dimorphodon Last Modified in 03/18/24	13	124	123	0	0	0	0	1
[5] † Peteinosaurus Last Modified in 03/18/24	57	79	79	0	0	0	0	1
[6] † Anurognathus Last Modified in 03/18/24	23	113	113	0	0	0	0	1
[7] † Batrachognathus Last Modified in 03/18/24	41	95	95	0	0	0	0	1
[8] † Dendrorhynchoides Last Modified in 03/18/24	52	85	84	0	0	0	0	1
[9] † Jeholopterus Last Modified in 03/18/24	45	91	91	0	0	0	0	1
[10] † Austriadactylus Last Modified in 03/18/24	58	78	78	0	0	0	0	1
[11] † Campylognathoides Last Modified in 03/18/24	4	133	132	0	0	0	0	1
[12] † Eudimorphodon Last Modified in 03/18/24	11	131	125	0	0	0	0	1
[13] † Raeticodactylus Last Modified in 03/18/24	42	94	94	0	0	0	0	1
[14] † Scaphognathus Last Modified in 03/18/24	8	128	128	0	0	0	0	1
[15] † Sordes Last Modified in 03/18/24	14	123	122	0	0	0	0	1
[16] † Pterorhynchus Last Modified in 03/18/24	50	86	86	0	0	0	0	1
[17] † Cacibupteryx Last Modified in 03/18/24	90	46	46	0	0	0	0	1
[18] † Dorygnathus Last Modified in 03/18/24	5	131	131	0	0	0	0	1
[19] † Nesodactylus Last Modified in 03/18/24	94	42	42	0	0	0	0	1
[20] † Dearc Last Modified in 03/18/24	60	76	76	0	0	0	0	1
[21] † Rhamphorhynchus Last Modified in 03/18/24	1	136	135	0	0	0	0	1
[22] † Ceoptera Last Modified in 03/18/24	113	23	23	0	0	0	0	1
[23] † Allkaruen Last Modified in 03/18/24	81	55	55	0	0	0	0	1
[24] † Cuspicephalus Last Modified in 03/18/24	93	43	43	0	0	0	0	1
[25] † Kryptodrakon Last Modified in 03/18/24	123	13	13	0	0	0	0	1
[26] † Darwinopterus Last Modified in 03/18/24	6	136	130	0	0	0	0	1
[27] † Wukongopterus Last Modified in 03/18/24	57	79	79	0	0	0	0	1
[28] † Kunpengopterus Last Modified in 03/18/24	9	128	127	0	0	0	0	1
[29] † Changchengopterus Last Modified in 03/18/24	81	55	55	0	0	0	0	1
[30] † Douzhanopterus Last Modified in 03/18/24	94	42	42	0	0	0	0	1
[31] † Rhamphodactylus Last Modified in 03/18/24	7	130	129	0	0	0	0	1
[32] † Liaodactylus Last Modified in 03/18/24	73	63	63	0	0	0	0	1
[33] † Pterodactylus Last Modified in 03/18/24	5	134	131	0	0	0	0	1
[34] † Cycnorhamphus Last Modified in 03/18/24	15	121	121	0	0	0	0	1
[35] † Eosipterus Last Modified in 03/18/24	87	49	49	0	0	0	0	1
[36] † Aurorazhdarcho Last Modified in 03/18/24	10	127	126	0	0	0	0	1
[37] † Gnathosaurus Last Modified in 03/18/24	66	70	70	0	0	0	0	1
[38] † Gegepterus Last Modified in 03/18/24	50	86	86	0	0	0	0	1
[39] † Gladocephaloideus Last Modified in 03/18/24	25	111	111	0	0	0	0	1
[40] † Ctenochasma Last Modified in 03/18/24	18	120	118	0	0	0	0	1
[41] † Pterodaustro Last Modified in 03/18/24	6	130	130	0	0	0	0	1
[42] † Istiodactylus Last Modified in 03/18/24	27	110	109	0	0	0	0	1
[43] † Nurhachius Last Modified in 03/18/24	34	102	102	0	0	0	0	1
[44] † Haopterus Last Modified in 03/18/24	53	83	83	0	0	0	0	1
[45] † Ornithocheirus Last Modified in 03/18/24	73	63	63	0	0	0	0	1
[46] † Anhanguera Last Modified in 03/18/24	10	128	126	0	0	0	0	1
[47] † Coloborhynchus Last Modified in 03/18/24	12	124	124	0	0	0	0	1
[48] † Ludodactylus Last Modified in 03/18/24	74	62	62	0	0	0	0	1
[49] † Boreopterus Last Modified in 03/18/24	36	101	100	0	0	0	0	1
[50] † Nyctosaurus Last Modified in 03/18/24	24	113	112	0	0	0	0	1
[51] † Muzquizopteryx Last Modified in 03/18/24	68	68	68	0	0	0	0	1
[52] † Pteranodon Last Modified in 03/18/24	21	115	115	0	0	0	0	1
[53] † Germanodactyluscrist Last Modified in 03/18/24	21	116	115	0	0	0	0	1
[54] † Germanodactylusrhamph Last Modified in 03/18/24	17	119	119	0	0	0	0	1
[55] † Dsungaripterus Last Modified in 03/18/24	26	110	110	0	0	0	0	1
[56] † Tatalpterosaur Last Modified in 03/18/24	37	99	99	0	0	0	0	1
[57] † Noripterus Last Modified in 03/18/24	87	49	49	0	0	0	0	1
[58] † Lonchodectes Last Modified in 03/18/24	86	52	50	0	0	0	0	1
[59] † Tapejara Last Modified in 03/18/24	36	100	100	0	0	0	0	1
[60] † Tupandactylus Last Modified in 03/18/24	29	107	107	0	0	0	0	1
[61] † Sinopterus Last Modified in 03/18/24	37	100	99	0	0	0	0	1
[62] † Huaxiapterus Last Modified in 03/18/24	45	92	91	0	0	0	0	1
[63] † Shenzhoupterus Last Modified in 03/18/24	53	83	83	0	0	0	0	1
[64] † Chaoyangopterus Last Modified in 03/18/24	70	66	66	0	0	0	0	1
[65] † Tupuxuara Last Modified in 03/18/24	25	111	111	0	0	0	0	1
[66] † Thalassodromeus Last Modified in 03/18/24	89	47	47	0	0	0	0	1
[67] † Quetzalcoatlus Last Modified in 03/18/24	43	93	93	0	0	0	0	1
[68] † Zhejiangopterus Last Modified in 03/18/24	40	96	96	0	0	0	0	1
[69] † Azhdarcho Last Modified in 03/18/24	93	43	43	0	0	0	0	1

Project views

type	number of views	Individual items viewed (where applicable)
Total project views	712
Project overview	48
Matrix views	36	Matrix landing page (31 views); matrix (5 views);
Media views	133	Media search (114 views); M902669 (19 views);
Views for media list	44
Specimen list	64
Bibliography	26
Taxon list	361