Project 5144: F. M. Rotatori, L. Ferrari, C. Sequero, B. Camilo, O. Mateus, M. Moreno-Azanza. 2024. An unexpected early-diverging iguanodontian dinosaur (Ornithischia, Ornithopoda) from the Upper Jurassic of Portugal. Journal of Vertebrate Paleontology. 43 (3):null.
Abstract
Iguanodontia is a diverse clade of herbivorous ornithischian dinosaurs that were speciose and abundant during the Jurassic and Cretaceous. Although the monophyly of Iguanodontia is well supported, their internal relationships have sparked heated debate due to several phylogenetic paradigm shifts. Late Jurassic basally branching iguanodontians in particular are not well understood in terms of their systematic affinities and evolutionary relevance. Their fossil record in Europe is meager compared with North America, with only a few species currently recognized. Two taxa are currently known from the Upper Jurassic of England, the basally branching styracosternan Cumnoria prestwichii and the putative dryosaurid Callovosaurus leedsi. In the Upper Jurassic of Portugal, the styracosternan Draconyx loureiroi and the dryosaurid Eousdryosaurus nanohallucis are presently the only described basally branching iguanodontians. Here we report a new species of early diverging iguanodontian from the Upper Jurassic Lourinhã Formation of western-central Portugal. The new species is clearly distinguished from all other coeval taxa by an exclusive combination of characters that include a tibia with a cnemial crest that is directed craniolaterally and a fibular condyle that is angled at 90° with respect to the proximal epiphysis, a fibula with symmetrical proximal margins, and a reduced metatarsal I. The phylogenetic relationships of the Lourinhã iguanodontian were explored using maximum parsimony and Bayesian inference. The two analyses recover the Lourinhã iguanodontian as an indeterminate dryomorphan, with more precise affinities precluded due to the current available material. Body size is estimated between 3 and 4 meters for the holotype specimen, adding to the diversity of small ornithopods already recognized in the paleoichnological record of the Lourinhã Formation.Read the article »
Article DOI: 10.1080/02724634.2024.2310066
Project DOI: 10.7934/P5144, http://dx.doi.org/10.7934/P5144
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 11607 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 343 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 765 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:343/0 |
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MorphoBank Project 5144
MorphoBank Project 5144
- Creation Date:
25 March 2024 - Publication Date:
25 March 2024 - Project views: 832
- Matrix downloads: 1
Authors' Institutions
- Opole University
- Museu da Lourinha
- Staatliches Museum fur Naturkunde Stuttgart
- Rheinische Friedrich-Wilhelms-Universität Bonn (University of Bonn)
- Universidade Nova de Lisboa
- Universidad de Zaragoza
- Sociedade de História Natural
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 73 | 1 | 1 | 343 | 0 | 0 | 11607 (11607, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29731' (73 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Herrerasaurus ischigualastensis Last Modified in 03/25/24 | 73 | 270 | 270 | 0 | 0 | 0 | 0 | 1 |
[2] † Abrictosaurus consors Last Modified in 03/25/24 | 224 | 119 | 119 | 0 | 0 | 0 | 0 | 1 |
[3] † Agilisaurus louderbacki Last Modified in 03/25/24 | 92 | 251 | 251 | 0 | 0 | 0 | 0 | 1 |
[4] † Anabisetia saldiviai Last Modified in 03/25/24 | 202 | 142 | 141 | 0 | 0 | 0 | 0 | 1 |
[5] † Ankylosauria Last Modified in 03/25/24 | 172 | 172 | 171 | 0 | 0 | 0 | 0 | 1 |
[6] † Archaeoceratops oshimai Last Modified in 03/25/24 | 141 | 202 | 202 | 0 | 0 | 0 | 0 | 1 |
[7] † Camptosaurus aphanoecetes Last Modified in 03/25/24 | 184 | 160 | 159 | 0 | 0 | 0 | 0 | 1 |
[8] † Camptosaurus dispar Last Modified in 03/25/24 | 30 | 315 | 313 | 0 | 0 | 0 | 0 | 1 |
[9] † Changchunsaurus parvus Last Modified in 03/25/24 | 146 | 197 | 197 | 0 | 0 | 0 | 0 | 1 |
[10] † Chaoyangsaurus youngi Last Modified in 03/25/24 | 246 | 97 | 97 | 0 | 0 | 0 | 0 | 1 |
[11] † Chilesaurus diegosuarezi Last Modified in 03/25/24 | 194 | 149 | 149 | 0 | 0 | 0 | 0 | 1 |
[12] † Convolosaurus marri Last Modified in 03/25/24 | 63 | 281 | 280 | 0 | 0 | 0 | 0 | 1 |
[13] † Dryosaurus Last Modified in 03/25/24 | 28 | 315 | 315 | 0 | 0 | 0 | 0 | 1 |
[14] † Dysalotosaurus lettowvorbecki Last Modified in 03/25/24 | 48 | 295 | 295 | 0 | 0 | 0 | 0 | 1 |
[15] † Echinodon becklesii Last Modified in 03/25/24 | 299 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[16] † Emausaurus ernstii Last Modified in 03/25/24 | 233 | 110 | 110 | 0 | 0 | 0 | 0 | 1 |
[17] † Eocursor parvus Last Modified in 03/25/24 | 224 | 119 | 119 | 0 | 0 | 0 | 0 | 1 |
[18] † Eousdryosaurus nanohallucis Last Modified in 03/25/24 | 274 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[19] † Fostoria dhimbangunmal Last Modified in 03/25/24 | 300 | 43 | 43 | 0 | 0 | 0 | 0 | 1 |
[20] † Fruitadens haagarorum Last Modified in 03/25/24 | 267 | 76 | 76 | 0 | 0 | 0 | 0 | 1 |
[21] † Gasparinisaura cincosaltensis Last Modified in 03/25/24 | 119 | 225 | 224 | 0 | 0 | 0 | 0 | 1 |
[22] † Goyocephale lattimorei Last Modified in 03/25/24 | 202 | 142 | 141 | 0 | 0 | 0 | 0 | 1 |
[23] † Haya griva Last Modified in 03/25/24 | 84 | 259 | 259 | 0 | 0 | 0 | 0 | 1 |
[24] † Heterodontosaurus tucki Last Modified in 03/25/24 | 31 | 312 | 312 | 0 | 0 | 0 | 0 | 1 |
[25] † Hexinlusaurus multidens Last Modified in 03/25/24 | 114 | 229 | 229 | 0 | 0 | 0 | 0 | 1 |
[26] † Homalocephale calathocercos Last Modified in 03/25/24 | 194 | 149 | 149 | 0 | 0 | 0 | 0 | 1 |
[27] † Hypsilophodon foxii Last Modified in 03/25/24 | 13 | 330 | 330 | 0 | 0 | 0 | 0 | 1 |
[28] † Iguanodon bernissartensis Last Modified in 03/25/24 | 26 | 317 | 317 | 0 | 0 | 0 | 0 | 1 |
[29] † Isaberrysaura mollensis Last Modified in 03/25/24 | 258 | 85 | 85 | 0 | 0 | 0 | 0 | 1 |
[30] † Jeholosaurus shangyuanensis Last Modified in 03/25/24 | 87 | 256 | 256 | 0 | 0 | 0 | 0 | 1 |
[31] † Kangnasaurus coetzeei Last Modified in 03/25/24 | 308 | 35 | 35 | 0 | 0 | 0 | 0 | 1 |
[32] † Koreanosaurus boseongensis Last Modified in 03/25/24 | 281 | 62 | 62 | 0 | 0 | 0 | 0 | 1 |
[33] † Kulindadromeus zabaikalicus Last Modified in 03/25/24 | 180 | 163 | 163 | 0 | 0 | 0 | 0 | 1 |
[34] † Lesothosaurus diagnosticus Last Modified in 03/25/24 | 47 | 296 | 296 | 0 | 0 | 0 | 0 | 1 |
[35] † Laquintasaura venezuelae Last Modified in 03/25/24 | 270 | 73 | 73 | 0 | 0 | 0 | 0 | 1 |
[36] † Liaoceratops yanzigouensis Last Modified in 03/25/24 | 187 | 156 | 156 | 0 | 0 | 0 | 0 | 1 |
[37] † Lycorhinus angustidens Last Modified in 03/25/24 | 314 | 29 | 29 | 0 | 0 | 0 | 0 | 1 |
[38] † Macrogryphosaurus gondwanicus Last Modified in 03/25/24 | 291 | 52 | 52 | 0 | 0 | 0 | 0 | 1 |
[39] † Mahuidacursor lipanglef Last Modified in 03/25/24 | 302 | 41 | 41 | 0 | 0 | 0 | 0 | 1 |
[40] † Mochlodon suessi Last Modified in 03/25/24 | 304 | 39 | 39 | 0 | 0 | 0 | 0 | 1 |
[41] † Mochlodon vorosi Last Modified in 03/25/24 | 270 | 73 | 73 | 0 | 0 | 0 | 0 | 1 |
[42] † Morrosaurus antarcticus Last Modified in 03/25/24 | 322 | 21 | 21 | 0 | 0 | 0 | 0 | 1 |
[43] † Muttaburrasaurus langdoni Last Modified in 03/25/24 | 196 | 147 | 147 | 0 | 0 | 0 | 0 | 1 |
[44] † Nanosaurus agilis Last Modified in 03/25/24 | 189 | 154 | 154 | 0 | 0 | 0 | 0 | 1 |
[45] † Orodromeus makelai Last Modified in 03/25/24 | 56 | 287 | 287 | 0 | 0 | 0 | 0 | 1 |
[46] † Pachycephalosaurus wyomingensis Last Modified in 03/25/24 | 251 | 92 | 92 | 0 | 0 | 0 | 0 | 1 |
[47] † Parksosaurus warreni Last Modified in 03/25/24 | 164 | 179 | 179 | 0 | 0 | 0 | 0 | 1 |
[48] † Prenocephale prenes Last Modified in 03/25/24 | 213 | 130 | 130 | 0 | 0 | 0 | 0 | 1 |
[49] † Psittacosaurus mongoliensis Last Modified in 03/25/24 | 100 | 243 | 243 | 0 | 0 | 0 | 0 | 1 |
[50] † Psittacosaurus major Last Modified in 03/25/24 | 178 | 165 | 165 | 0 | 0 | 0 | 0 | 1 |
[51] † Rhabdodon priscus Last Modified in 03/25/24 | 301 | 42 | 42 | 0 | 0 | 0 | 0 | 1 |
[52] † Rhabdodon sp1 Last Modified in 03/25/24 | 261 | 83 | 82 | 0 | 0 | 0 | 0 | 1 |
[53] † Scelidosaurus harrisonii Last Modified in 03/25/24 | 154 | 189 | 189 | 0 | 0 | 0 | 0 | 1 |
[54] † Scutellosaurus lawleri Last Modified in 03/25/24 | 198 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
[55] † Stegoceras validum Last Modified in 03/25/24 | 111 | 233 | 232 | 0 | 0 | 0 | 0 | 1 |
[56] † Stegosauria Last Modified in 03/25/24 | 97 | 255 | 246 | 0 | 0 | 0 | 0 | 1 |
[57] † Stenopelix valdensis Last Modified in 03/25/24 | 287 | 56 | 56 | 0 | 0 | 0 | 0 | 1 |
[58] † Talenkauen santacrucensis Last Modified in 03/25/24 | 192 | 151 | 151 | 0 | 0 | 0 | 0 | 1 |
[59] † Tenontosaurus dossi Last Modified in 03/25/24 | 78 | 265 | 265 | 0 | 0 | 0 | 0 | 1 |
[60] † Tenontosaurus tilletti Last Modified in 03/25/24 | 18 | 325 | 325 | 0 | 0 | 0 | 0 | 1 |
[61] † Thescelosaurus neglectus Last Modified in 03/25/24 | 32 | 312 | 311 | 0 | 0 | 0 | 0 | 1 |
[62] † Thescelosaurus assiniboiensis Last Modified in 03/25/24 | 270 | 73 | 73 | 0 | 0 | 0 | 0 | 1 |
[63] † Tianyulong confuciusi Last Modified in 03/25/24 | 231 | 112 | 112 | 0 | 0 | 0 | 0 | 1 |
[64] † Valdosaurus canaliculatus Last Modified in 03/25/24 | 261 | 82 | 82 | 0 | 0 | 0 | 0 | 1 |
[65] † Vegagete ornithopod Last Modified in 03/25/24 | 249 | 94 | 94 | 0 | 0 | 0 | 0 | 1 |
[66] † Wannanosaurus yansiensis Last Modified in 03/25/24 | 250 | 93 | 93 | 0 | 0 | 0 | 0 | 1 |
[67] † Yandusaurus hongheensis Last Modified in 03/25/24 | 268 | 75 | 75 | 0 | 0 | 0 | 0 | 1 |
[68] † Yueosaurus tiantaiensis Last Modified in 03/25/24 | 299 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[69] † Yinlong downsi Last Modified in 03/25/24 | 84 | 262 | 259 | 0 | 0 | 0 | 0 | 1 |
[70] † Zalmoxes robustus Last Modified in 03/25/24 | 106 | 237 | 237 | 0 | 0 | 0 | 0 | 1 |
[71] † Zalmoxes shqiperorum Last Modified in 03/25/24 | 172 | 171 | 171 | 0 | 0 | 0 | 0 | 1 |
[72] † Zephyrosaurus schaffi Last Modified in 03/25/24 | 217 | 126 | 126 | 0 | 0 | 0 | 0 | 1 |
[73] † ML 2700 Last Modified in 03/25/24 | 328 | 15 | 15 | 0 | 0 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 832 | |
Project overview | 49 | |
Taxon list | 472 | |
Matrix views | 40 | Matrix landing page (35 views); Matrix (5 views); |
Specimen list | 67 | |
Views for media list | 38 | |
Media views | 133 | Media search (116 views); M902691 (17 views); |
Bibliography | 33 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 1 | |
Matrix downloads | 1 | Matrix (1 download); |