Morphobank

Project 5161: D. Conedera, Y. Pochat-Cottilloux, N. Rinder, J. Adrien, J. E. Martin. 2024. An anatomical reappraisal of the dwarf crocodylian Arambourgia gaudryi from the Eocene of Quercy (France) using CT data and its implications for the phylogeny and paleoecology of basally branching alligatoroids. Journal of Vertebrate Paleontology. null (null):1-31.

Specimen: † Crocodylus siamensis (unvouchered)
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Abstract

After about a century, the skull of the holotype and sole specimen of Arambourgia gaudryi from the Eocene of Quercy (France) is here redescribed. Its interest resides in its extreme morphology, such as its very small skull and altirostral condition, that raises questions about its maturity, taxonomic validity, and paleoecology. In order to reveal previously hidden anatomical details, we used computed microtomography (µCT-scan) data to digitally extract the bones of the cranium and mandible, as well as the endocranial structures. Here, we update the osteological description of A. gaudryi, outlining new characters and re-evaluating anatomical details. Comparing it with ontogenetic series of extant crocodylians, we confirm that the specimen is a mature individual of a dwarf species with pedomorphic features. Our phylogenetic analyses indicate that A. gaudryi is an alligatorine. Comparisons with other small alligatoroid taxa show that it is indeed a valid species. The depositional environment of the finding site suggests a continental context with a karstic network. Taking this into account, together with its peculiar anatomy, we propose that A. gaudryi was a semi-terrestrial animal occupying an ecological niche similar to that of extant juvenile or dwarf crocodylians. Finally, we discuss the current knowledge of the paleobiogeography of Alligatoroidea in Europe during the Paleogene. Arambourgia gaudryi was likely part of a group of small early globidontans that arrived in the Old World, probably from North America, around the end of the Paleocene and that dispersed to at least four localities of the ancient European archipelago.

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Article DOI: 10.1080/02724634.2024.2313612

Project DOI: 10.7934/P5161, http://dx.doi.org/10.7934/P5161

This project contains	Matrices
1 Media 2 Matrices 2 Documents 161 Taxa 1 Specimen 517 Characters Total size of project's media files: 908.72k Download Project SDD File	Total scored cells: 36789 Total media associated with cells: 0 Total labels associated with cell media: 0
	Characters
	Total characters: 517 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1185 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:517/0

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MorphoBank Project 5161

Creation Date:
27 March 2024
Publication Date:
27 March 2024
Project views: 870
Matrix downloads: 1

Authors' Institutions

Université de Lyon (University of Lyon)

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MorphoBank Curator
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Taxonomic Overview for Matrix 'M29750' (144 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Bernissartia fagesii Last Modified in 03/27/24	140	190	190	0	0	0	0	1
[2] † Isisfordia duncani Last Modified in 03/27/24	116	214	214	0	0	0	0	1
[3] † Acynodon iberoccitanus Last Modified in 03/27/24	189	142	141	0	0	0	0	1
[4] † Agaresuchus fontisensis Last Modified in 03/27/24	132	198	198	0	0	0	0	1
[5] † Allodaposuchus precedens Last Modified in 03/27/24	144	186	186	0	0	0	0	1
[6] † Glen rose form Last Modified in 03/27/24	156	174	174	0	0	0	0	1
[7] † Hylaeochampsa vectiana Last Modified in 03/27/24	209	121	121	0	0	0	0	1
[8] † Iharkutosuchus makadii Last Modified in 03/27/24	156	174	174	0	0	0	0	1
[9] † Lohuecosuchus megadontos Last Modified in 03/27/24	130	201	200	0	0	0	0	1
[10] † Portugalosuchus azenhae Last Modified in 03/27/24	215	115	115	0	0	0	0	1
[11] † Shamosuchus djadochtaensis Last Modified in 03/27/24	175	155	155	0	0	0	0	1
[12] † Wannchampsus kirpachi Last Modified in 03/27/24	176	155	154	0	0	0	0	1
[13] † Acresuchus pachytemporalis Last Modified in 03/27/24	142	188	188	0	0	0	0	1
[14] † Aktiogavialis caribesi Last Modified in 03/27/24	214	116	116	0	0	0	0	1
[15] † Alligator mcgrewi Last Modified in 03/27/24	63	271	267	0	0	0	0	1
[16] † Alligator mefferdi Last Modified in 03/27/24	98	232	232	0	0	0	0	1
[17] Alligator mississippiensis Last Modified in 03/27/24	12	329	318	0	0	0	0	1
[18] † Alligator olseni Last Modified in 03/27/24	117	213	213	0	0	0	0	1
[19] † Alligator prenasalis Last Modified in 03/27/24	76	260	254	0	0	0	0	1
[20] † Alligator sinensis Last Modified in 03/27/24	21	311	309	0	0	0	0	1
[21] † Hassiacosuchus haupti Last Modified in 03/27/24	182	149	148	0	0	0	0	1
[22] † Navajosuchus mooki Last Modified in 03/27/24	143	189	187	0	0	0	0	1
[23] † Allognathosuchus polyodon Last Modified in 03/27/24	250	80	80	0	0	0	0	1
[24] † Allognathosuchus wartheni Last Modified in 03/27/24	189	141	141	0	0	0	0	1
[25] † Arambourgia gaudryi Last Modified in 03/27/24	115	217	215	0	0	0	0	1
[26] † Argochampsa krebsi Last Modified in 03/27/24	153	177	177	0	0	0	0	1
[27] † Asiatosuchus germanicus Last Modified in 03/27/24	127	203	203	0	0	0	0	1
[28] † Asiatosuchus nanlingensis Last Modified in 03/27/24	298	32	32	0	0	0	0	1
[29] † Australosuchus clarkae Last Modified in 03/27/24	159	171	171	0	0	0	0	1
[30] † Baru darrowi Last Modified in 03/27/24	218	113	112	0	0	0	0	1
[31] † Baru huberi Last Modified in 03/27/24	261	69	69	0	0	0	0	1
[32] † Baru wickeni Last Modified in 03/27/24	151	179	179	0	0	0	0	1
[33] † Borealosuchus acutidentatus Last Modified in 03/27/24	228	102	102	0	0	0	0	1
[34] † Borealosuchus formidabilis Last Modified in 03/27/24	62	268	268	0	0	0	0	1
[35] † Borealosuchus sternbergii Last Modified in 03/27/24	67	269	263	0	0	0	0	1
[36] † Borealosuchus threensis Last Modified in 03/27/24	287	43	43	0	0	0	0	1
[37] † Borealosuchus wilsoni Last Modified in 03/27/24	197	133	133	0	0	0	0	1
[38] † Bottosaurus harlani Last Modified in 03/27/24	221	109	109	0	0	0	0	1
[39] † Boverisuchus magnifrons Last Modified in 03/27/24	223	107	107	0	0	0	0	1
[40] † Boverisuchus vorax Last Modified in 03/27/24	98	232	232	0	0	0	0	1
[41] † Brachychampsa montana Last Modified in 03/27/24	69	261	261	0	0	0	0	1
[42] † Brochuchus pigotti Last Modified in 03/27/24	116	214	214	0	0	0	0	1
[43] † Caiman brevirostris Last Modified in 03/27/24	231	99	99	0	0	0	0	1
[44] Caiman crocodilus Last Modified in 03/27/24	12	334	318	0	0	0	0	1
[45] † Caiman gasparinae Last Modified in 03/27/24	262	68	68	0	0	0	0	1
[46] † Caiman latirostris Last Modified in 03/27/24	23	314	307	0	0	0	0	1
[47] † Caiman lutescens MACN Last Modified in 03/27/24	297	33	33	0	0	0	0	1
[48] † UCMP 39978 Last Modified in 03/27/24	230	100	100	0	0	0	0	1
[49] † Caiman wannlangstoni Last Modified in 03/27/24	207	123	123	0	0	0	0	1
[50] † Caiman yacare Last Modified in 03/27/24	21	319	309	0	0	0	0	1
[51] † Ceratosuchus burdoschi Last Modified in 03/27/24	197	133	133	0	0	0	0	1
[52] † Crocodylus acutus Last Modified in 03/27/24	20	312	310	0	0	0	0	1
[53] † Crocodylus affinis Last Modified in 03/27/24	79	259	251	0	0	0	0	1
[54] † Crocodylus anthropophagus Last Modified in 03/27/24	172	158	158	0	0	0	0	1
[55] † Asiatosuchus depressifrons Last Modified in 03/27/24	71	260	259	0	0	0	0	1
[56] † Crocodylus intermedius Last Modified in 03/27/24	39	295	291	0	0	0	0	1
[57] † Crocodylus johnstoni Last Modified in 03/27/24	17	321	313	0	0	0	0	1
[58] † Crocodylus megarhinus Last Modified in 03/27/24	103	227	227	0	0	0	0	1
[59] † Crocodylus mindorensis Last Modified in 03/27/24	63	267	267	0	0	0	0	1
[60] † Crocodylus moreletii Last Modified in 03/27/24	59	271	271	0	0	0	0	1
[61] Crocodylus niloticus Last Modified in 03/27/24	12	325	318	0	0	0	0	1
[62] † Crocodylus novaeguineae Last Modified in 03/27/24	61	269	269	0	0	0	0	1
[63] † Crocodylus palustris Last Modified in 03/27/24	19	315	311	0	0	0	0	1
[64] † Crocodylus porosus Last Modified in 03/27/24	14	331	316	0	0	0	0	1
[65] † Crocodylus rhombifer Last Modified in 03/27/24	15	317	315	0	0	0	0	1
[66] † Crocodylus siamensis Last Modified in 03/27/24	19	313	311	0	0	0	0	1
[67] † Crocodylus thorbjarnarsoni Last Modified in 03/27/24	135	195	195	0	0	0	0	1
[68] † Crocodylus palaeindicus Last Modified in 03/27/24	129	203	201	0	0	0	0	1
[69] † Dadagavialis gunai Last Modified in 03/27/24	255	75	75	0	0	0	0	1
[70] † Diplocynodon darwini Last Modified in 03/27/24	86	247	244	0	0	0	0	1
[71] † Diplocynodon deponiae Last Modified in 03/27/24	182	150	148	0	0	0	0	1
[72] † Diplocynodon hantoniensis Last Modified in 03/27/24	67	268	263	0	0	0	0	1
[73] † Diplocynodon muelleri Last Modified in 03/27/24	185	145	145	0	0	0	0	1
[74] † Diplocynodon ratelii Last Modified in 03/27/24	70	260	260	0	0	0	0	1
[75] † Diplocynodon remensis Last Modified in 03/27/24	129	202	201	0	0	0	0	1
[76] † Diplocynodon tormis Last Modified in 03/27/24	199	131	131	0	0	0	0	1
[77] † Dollosuchoides densmorei Last Modified in 03/27/24	178	152	152	0	0	0	0	1
[78] † Eocaiman cavernensis Last Modified in 03/27/24	237	93	93	0	0	0	0	1
[79] † Eocaiman palaeocenicus Last Modified in 03/27/24	298	32	32	0	0	0	0	1
[80] † Eogavialis africanum Last Modified in 03/27/24	102	234	228	0	0	0	0	1
[81] † Eosuchus lerichei Last Modified in 03/27/24	132	198	198	0	0	0	0	1
[82] † Eosuchus minor Last Modified in 03/27/24	116	214	214	0	0	0	0	1
[83] † Eothoracosaurus mississippiensi Last Modified in 03/27/24	167	164	163	0	0	0	0	1
[84] † Euthecodon arambourgi Last Modified in 03/27/24	190	140	140	0	0	0	0	1
[85] † Gavialis browni Last Modified in 03/27/24	218	112	112	0	0	0	0	1
[86] Gavialis gangeticus Last Modified in 03/27/24	15	319	315	0	0	0	0	1
[87] † Gavialis lewisi Last Modified in 03/27/24	172	158	158	0	0	0	0	1
[88] † Gavialosuchus eggenburgensis Last Modified in 03/27/24	211	119	119	0	0	0	0	1
[89] † Globidentosuchus brachyrostris Last Modified in 03/27/24	220	110	110	0	0	0	0	1
[90] † Gnatusuchus pebasensis Last Modified in 03/27/24	190	140	140	0	0	0	0	1
[91] † Gryposuchus colombianus Last Modified in 03/27/24	138	192	192	0	0	0	0	1
[92] † Gryposuchus croizati Last Modified in 03/27/24	247	83	83	0	0	0	0	1
[93] † Gryposuchus neogaeus Last Modified in 03/27/24	225	105	105	0	0	0	0	1
[94] † Gryposuchus pachakamue Last Modified in 03/27/24	153	178	177	0	0	0	0	1
[95] † Ikanogavialis gameroi Last Modified in 03/27/24	202	128	128	0	0	0	0	1
[96] † Jiangxisuchus nankangensis Last Modified in 03/27/24	160	170	170	0	0	0	0	1
[97] † Kambara implexidens Last Modified in 03/27/24	129	201	201	0	0	0	0	1
[98] † Kambara murgonensis Last Modified in 03/27/24	255	75	75	0	0	0	0	1
[99] † Kambara taraina Last Modified in 03/27/24	178	153	152	0	0	0	0	1
[100] † Kentisuchus spenceri Last Modified in 03/27/24	183	150	147	0	0	0	0	1
[101] † Kuttanacaiman iquitosensis Last Modified in 03/27/24	209	121	121	0	0	0	0	1
[102] † Leidyosuchus canadensis Last Modified in 03/27/24	90	243	240	0	0	0	0	1
[103] † Mecistops cataphractus Last Modified in 03/27/24	15	317	315	0	0	0	0	1
[104] † Mekosuchus inexpectatus Last Modified in 03/27/24	206	124	124	0	0	0	0	1
[105] † Mekosuchus sanderi Last Modified in 03/27/24	261	69	69	0	0	0	0	1
[106] † Mekosuchus whitehunterensis Last Modified in 03/27/24	273	57	57	0	0	0	0	1
[107] † Melanosuchus niger Last Modified in 03/27/24	20	312	310	0	0	0	0	1
[108] † Moroccosuchus zennaroi Last Modified in 03/27/24	142	189	188	0	0	0	0	1
[109] † Mourasuchus amazonensis Last Modified in 03/27/24	180	150	150	0	0	0	0	1
[110] † Mourasuchus arendsi Last Modified in 03/27/24	158	174	172	0	0	0	0	1
[111] † Mourasuchus atopus Last Modified in 03/27/24	200	130	130	0	0	0	0	1
[112] † Necrosuchus ionensis Last Modified in 03/27/24	292	38	38	0	0	0	0	1
[113] † Osteolaemus tetraspis Last Modified in 03/27/24	13	322	317	0	0	0	0	1
[114] † Paleosuchus palpebrosus Last Modified in 03/27/24	24	308	306	0	0	0	0	1
[115] † Paleosuchus trigonatus Last Modified in 03/27/24	24	310	306	0	0	0	0	1
[116] † Paratomistoma courti Last Modified in 03/27/24	243	87	87	0	0	0	0	1
[117] † Penghusuchus pani Last Modified in 03/27/24	132	198	198	0	0	0	0	1
[118] † Piscogavialis jugaliperforatus Last Modified in 03/27/24	123	207	207	0	0	0	0	1
[119] † Planocrania datangensis Last Modified in 03/27/24	228	102	102	0	0	0	0	1
[120] † Planocrania hengdongensis Last Modified in 03/27/24	237	93	93	0	0	0	0	1
[121] † Procaimanoidea utahensis Last Modified in 03/27/24	139	191	191	0	0	0	0	1
[122] † Protocaiman peligrensis Last Modified in 03/27/24	283	47	47	0	0	0	0	1
[123] † Purussaurus barasiliensis Last Modified in 03/27/24	133	197	197	0	0	0	0	1
[124] † Purussaurus mirandai Last Modified in 03/27/24	216	114	114	0	0	0	0	1
[125] † Purussaurus neivensis Last Modified in 03/27/24	92	238	238	0	0	0	0	1
[126] † Quinkana Last Modified in 03/27/24	243	99	87	0	0	0	0	1
[127] † Siquisiquesuchus venezuelensis Last Modified in 03/27/24	248	82	82	0	0	0	0	1
[128] † Stangerochampsa mccabei Last Modified in 03/27/24	100	230	230	0	0	0	0	1
[129] † Thecachampsa antiquus Last Modified in 03/27/24	195	135	135	0	0	0	0	1
[130] † Thecachampsa sericodon Last Modified in 03/27/24	134	196	196	0	0	0	0	1
[131] † Thoracosaurus isorhynchus Last Modified in 03/27/24	170	160	160	0	0	0	0	1
[132] † Thoracosaurus neocesariensis Last Modified in 03/27/24	255	76	75	0	0	0	0	1
[133] † Tomistoma cairense Last Modified in 03/27/24	173	157	157	0	0	0	0	1
[134] † Tomistoma dowsoni Last Modified in 03/27/24	214	116	116	0	0	0	0	1
[135] † Tomistoma lusitanica Last Modified in 03/27/24	171	159	159	0	0	0	0	1
[136] † Maomingosuchus petrolica Last Modified in 03/27/24	105	228	225	0	0	0	0	1
[137] † Tomistoma schlegelii Last Modified in 03/27/24	13	322	317	0	0	0	0	1
[138] † Toyotamaphimeia machikanensis Last Modified in 03/27/24	93	237	237	0	0	0	0	1
[139] † Trilophosuchus rackhami Last Modified in 03/27/24	201	129	129	0	0	0	0	1
[140] † Tsoabichi greenriverensis Last Modified in 03/27/24	230	102	100	0	0	0	0	1
[141] † Ultrastenos willisi Last Modified in 03/27/24	228	102	102	0	0	0	0	1
[142] † Voay robustus Last Modified in 03/27/24	64	267	266	0	0	0	0	1
[143] † Wannaganosuchus brachymanus Last Modified in 03/27/24	178	152	152	0	0	0	0	1
[144] † Theriosuchus pusillus Last Modified in 03/27/24	141	189	189	0	0	0	0	1

Taxonomic Overview for Matrix 'M29751' (94 Taxa)

taxon	unscored cells	scored cells	no cell support	NPA cells	"-" cells	cell images	labels on cell images	member access
[1] † Bernissartia fagesii Last Modified in 03/27/24	80	107	107	0	0	0	0	1
[2] † Allodaposuchus precedens Last Modified in 03/27/24	101	86	86	0	0	0	0	1
[3] † Acynodon iberoccitanus Last Modified in 03/27/24	100	87	87	0	0	0	0	1
[4] † Acynodon adriaticus Last Modified in 03/27/24	106	81	81	0	0	0	0	1
[5] † Iharkutosuchus makadii Last Modified in 03/27/24	98	89	89	0	0	0	0	1
[6] † Hylaeochampsa vectiana Last Modified in 03/27/24	110	77	77	0	0	0	0	1
[7] † Eothoracosaurus mississippiensis Last Modified in 03/27/24	86	101	101	0	0	0	0	1
[8] † Eosuchus minor Last Modified in 03/27/24	53	134	134	0	0	0	0	1
[9] † Eogavialis africanum Last Modified in 03/27/24	63	124	124	0	0	0	0	1
[10] † Gryposuchus colombianus Last Modified in 03/27/24	53	134	134	0	0	0	0	1
[11] Gavialis gangeticus Last Modified in 03/27/24	2	185	185	0	0	0	0	1
[12] † Borealosuchus sternbergii Last Modified in 03/27/24	17	170	170	0	0	0	0	1
[13] † Borealosuchus formidabilis Last Modified in 03/27/24	23	164	164	0	0	0	0	1
[14] † Borealosuchus acutidentatus Last Modified in 03/27/24	117	70	70	0	0	0	0	1
[15] † Borealosuchus wilsoni Last Modified in 03/27/24	55	132	132	0	0	0	0	1
[16] † Planocrania hengdongensis Last Modified in 03/27/24	123	64	64	0	0	0	0	1
[17] † Planocrania datangensis Last Modified in 03/27/24	131	56	56	0	0	0	0	1
[18] † Boverisuchus magnifrons Last Modified in 03/27/24	82	105	105	0	0	0	0	1
[19] † Boverisuchus vorax Last Modified in 03/27/24	50	137	137	0	0	0	0	1
[20] † Asiatosuchus germanicus Last Modified in 03/27/24	55	132	132	0	0	0	0	1
[21] † Crocodylus affinis Last Modified in 03/27/24	24	163	163	0	0	0	0	1
[22] † Crocodylus acer Last Modified in 03/27/24	95	92	92	0	0	0	0	1
[23] † Kentisuchus spenceri Last Modified in 03/27/24	98	89	89	0	0	0	0	1
[24] † Thecachampsa americana Last Modified in 03/27/24	66	121	121	0	0	0	0	1
[25] † Tomistoma schlegelii Last Modified in 03/27/24	1	186	186	0	0	0	0	1
[26] † Kambara implexidens Last Modified in 03/27/24	49	138	138	0	0	0	0	1
[27] † Australosuchus clarkae Last Modified in 03/27/24	66	121	121	0	0	0	0	1
[28] † Trilophosuchus rackhami Last Modified in 03/27/24	115	72	72	0	0	0	0	1
[29] † Crocodylus megarhinus Last Modified in 03/27/24	71	116	116	0	0	0	0	1
[30] † Rimasuchus lloydi Last Modified in 03/27/24	99	88	88	0	0	0	0	1
[31] † Voay robustus Last Modified in 03/27/24	42	145	145	0	0	0	0	1
[32] † Osteolaemus tetraspis Last Modified in 03/27/24	3	184	184	0	0	0	0	1
[33] † Osteolaemus osborni Last Modified in 03/27/24	12	175	175	0	0	0	0	1
[34] † Mecistops cataphractus Last Modified in 03/27/24	2	185	185	0	0	0	0	1
[35] † Crocodylus porosus Last Modified in 03/27/24	0	187	187	0	0	0	0	1
[36] † Crocodylus acutus Last Modified in 03/27/24	0	187	187	0	0	0	0	1
[37] Crocodylus niloticus Last Modified in 03/27/24	0	187	187	0	0	0	0	1
[38] † Prodiplocynodon langi Last Modified in 03/27/24	104	83	83	0	0	0	0	1
[39] † Leidyosuchus canadensis Last Modified in 03/27/24	37	150	150	0	0	0	0	1
[40] † Diplocynodon deponiae Last Modified in 03/27/24	88	99	99	0	0	0	0	1
[41] † Diplocynodon darwini Last Modified in 03/27/24	34	153	153	0	0	0	0	1
[42] † Diplocynodon ratelii Last Modified in 03/27/24	41	146	146	0	0	0	0	1
[43] † Diplocynodon hantoniensis Last Modified in 03/27/24	36	151	151	0	0	0	0	1
[44] † Diplocynodon muelleri Last Modified in 03/27/24	69	118	118	0	0	0	0	1
[45] † Diplocynodon tormis Last Modified in 03/27/24	80	107	107	0	0	0	0	1
[46] † Ceratosuchus burdoshi Last Modified in 03/27/24	113	74	74	0	0	0	0	1
[47] † Navajosuchus mooki Last Modified in 03/27/24	73	114	114	0	0	0	0	1
[48] † Hassiacosuchus haupti Last Modified in 03/27/24	85	102	102	0	0	0	0	1
[49] † Allognathosuchus polyodon Last Modified in 03/27/24	112	75	75	0	0	0	0	1
[50] † Allognathosuchus wartheni Last Modified in 03/27/24	51	136	136	0	0	0	0	1
[51] † Procaimanoidea kayi Last Modified in 03/27/24	61	126	126	0	0	0	0	1
[52] † Procaimanoidea utahensis Last Modified in 03/27/24	77	110	110	0	0	0	0	1
[53] † Arambourgia gaudryi Last Modified in 03/27/24	68	119	119	0	0	0	0	1
[54] † Wannaganosuchus brachymanus Last Modified in 03/27/24	78	109	109	0	0	0	0	1
[55] † Alligator sinensis Last Modified in 03/27/24	3	184	184	0	0	0	0	1
[56] Alligator mississippiensis Last Modified in 03/27/24	0	187	187	0	0	0	0	1
[57] † Alligator thomsoni Last Modified in 03/27/24	75	112	112	0	0	0	0	1
[58] † Alligator prenasalis Last Modified in 03/27/24	28	159	159	0	0	0	0	1
[59] † Alligator mefferdi Last Modified in 03/27/24	59	128	128	0	0	0	0	1
[60] † Alligator olseni Last Modified in 03/27/24	57	130	130	0	0	0	0	1
[61] † Alligator mcgrewi Last Modified in 03/27/24	26	161	161	0	0	0	0	1
[62] † Stangerochampsa mccabei Last Modified in 03/27/24	41	146	146	0	0	0	0	1
[63] † Albertochampsa langstoni Last Modified in 03/27/24	113	74	74	0	0	0	0	1
[64] † Brachychampsa montana Last Modified in 03/27/24	26	161	161	0	0	0	0	1
[65] † Brachychampsa sealeyi Last Modified in 03/27/24	125	62	62	0	0	0	0	1
[66] † Culebrasuchus mesoamericanus Last Modified in 03/27/24	156	31	31	0	0	0	0	1
[67] † Gnatusuchus pebasensis Last Modified in 03/27/24	88	99	99	0	0	0	0	1
[68] † Globidentosuchus brachyrostris Last Modified in 03/27/24	118	69	69	0	0	0	0	1
[69] † Purussaurus mirandai Last Modified in 03/27/24	74	113	113	0	0	0	0	1
[70] † Purussaurus neivensis Last Modified in 03/27/24	45	142	142	0	0	0	0	1
[71] † Mourasuchus amazonensis Last Modified in 03/27/24	148	39	39	0	0	0	0	1
[72] † Mourasuchus arendsi Last Modified in 03/27/24	119	68	68	0	0	0	0	1
[73] † Mourasuchus atopus Last Modified in 03/27/24	119	68	68	0	0	0	0	1
[74] † Mourasuchus pattersoni Last Modified in 03/27/24	139	48	48	0	0	0	0	1
[75] † Caiman yacare Last Modified in 03/27/24	0	187	187	0	0	0	0	1
[76] Caiman crocodilus Last Modified in 03/27/24	0	187	187	0	0	0	0	1
[77] † Caiman latirostris Last Modified in 03/27/24	4	183	183	0	0	0	0	1
[78] † Melanosuchus niger Last Modified in 03/27/24	3	184	184	0	0	0	0	1
[79] † Paleosuchus trigonatus Last Modified in 03/27/24	1	186	186	0	0	0	0	1
[80] † Paleosuchus palpebrosus Last Modified in 03/27/24	2	185	185	0	0	0	0	1
[81] † Caiman brevirostris Last Modified in 03/27/24	130	57	57	0	0	0	0	1
[82] † Tsoabichi greenriverensis Last Modified in 03/27/24	127	60	60	0	0	0	0	1
[83] † Centenariosuchus gilmorei Last Modified in 03/27/24	148	39	39	0	0	0	0	1
[84] † Eocaiman cavernensis Last Modified in 03/27/24	150	37	37	0	0	0	0	1
[85] † Eocaiman palaeocenicus Last Modified in 03/27/24	172	15	15	0	0	0	0	1
[86] † Eocaiman itaboraiensis Last Modified in 03/27/24	181	6	6	0	0	0	0	1
[87] † Kuttanacaiman iquitosensis Last Modified in 03/27/24	78	109	109	0	0	0	0	1
[88] † Caiman wannlangstoni Last Modified in 03/27/24	99	88	88	0	0	0	0	1
[89] † Acresuchus pachytemporalis Last Modified in 03/27/24	110	77	77	0	0	0	0	1
[90] † Purussaurus brasiliensis Last Modified in 03/27/24	106	81	81	0	0	0	0	1
[91] † Necrosuchus ionensis Last Modified in 03/27/24	157	30	30	0	0	0	0	1
[92] † Caiman gasparinae Last Modified in 03/27/24	157	30	30	0	0	0	0	1
[93] † Bottosaurus harlani Last Modified in 03/27/24	128	59	59	0	0	0	0	1
[94] † Protocaiman peligrensis Last Modified in 03/27/24	160	27	27	0	0	0	0	1

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