Project 5228: D. E. Perez, L. M. Giachetti, M. Hautmann. 2020. Is Cyclocardia (Conrad) a wastebasket taxon? Exploring the phylogeny of the most diverse genus of the Carditidae (Archiheterodonta, Bivalvia). Palaeontology. 63 (3):477-495.
Abstract
The carditid genus Cyclocardia is currently the most diverse genus of the family, including nearly 180 nominal species encompassing wide stratigraphical (Cretaceous–Recent) and geographical (Antarctica, South and North America, Europe, Africa, Alaska, Russia, Japan and New Zealand) ranges. Due to the lack of autapomorphies in the diagnosis of the genus and its large account of species, we re‐evaluate the systematic and phylogenetic status of Cyclocardia. We applied three approaches: bibliographic revision, phylogenetic analysis and an exploration of morphological disparity. We used a shell–character matrix comprising 65 taxa (2 outgroups, 29 non‐Cyclocardia carditids and 28 species of Cyclocardia) for phylogenetic and disparity analyses. Maximum Observable Rescaled Distances was used to construct a distance matrix to compare Cyclocardia species and other carditid groups. According to our results, Cyclocardia represents a non‐monophyletic taxon and is thus a ‘wastebasket taxon’, chiefly because its diagnosis was based mainly on plesiomorphic characters. The European species C. kickxi and C. chameformis are placed within Scalaricardita, and the previously proposed genus Crassicardia is monophyletic (including C. crassidens, C. crebricostata, C. isaotakii and C. rjabininae). Three new genera are proposed for new groups identified by the phylogenetic analysis: South American Oesterheldia gen. nov. (including O. cannada and O. dalek), western North American Coanicardita gen. nov. (including C. ventricosa and C. occidentalis), and North Pacific Hippocampocardia gen. nov. (including H. barbarensis, H. hamiltonensis and H. yakatagensis). The newly defined monophyletic Cyclocardia is restricted to the Atlantic Ocean species C. borealis, C. novangliae and C. compressa.Read the article »
Article DOI: 10.1111/pala.12467
Project DOI: 10.7934/P5228, http://dx.doi.org/10.7934/P5228
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 8581 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 146 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 446 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:146/0 |
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MorphoBank Project 5228
MorphoBank Project 5228
- Creation Date:
29 April 2024 - Publication Date:
02 July 2024 - Media downloads: 1
- Matrix downloads: 2
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| Shreya Jariwala Project Administrator | 59 | 8 | 8 | 146 | 0 | 0 | 8581 (8581, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M29806' (59 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] † Crassatella ponderosa Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [2] † Astarte sulcata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [3] † Palaeocardita stoecklini Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [4] † Claibornicardia alticostata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [5] † Glyptoactis hadra Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [6] † Arcturellina asperula Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [7] † Purpurocardia purpurata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [8] † Darwinicardia patagonica Taxon name last Modified on 04/29/24 | 0 | 149 | 146 | 0 | 0 | 0 | 0 | 1 |
| [9] † Leuroactis pilsbryi Taxon name last Modified on 04/29/24 | 1 | 146 | 145 | 0 | 0 | 0 | 0 | 1 |
| [10] † Pacificor mulleri Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [11] † Beguina semiorbiculata Taxon name last Modified on 04/29/24 | 9 | 137 | 137 | 0 | 0 | 0 | 0 | 1 |
| [12] † Cardita variegata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [13] † Thecalia concamerata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [14] † Venericardia imbricata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [15] † Cardites antiquatus Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [16] † Bathycardita raouli Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [17] † Megacardita jouanetti Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [18] † Venericor planicosta Taxon name last Modified on 04/29/24 | 1 | 146 | 145 | 0 | 0 | 0 | 0 | 1 |
| [19] † Coripia unidentata Taxon name last Modified on 04/29/24 | 0 | 147 | 146 | 0 | 0 | 0 | 0 | 1 |
| [20] † Kolmeris tehuelchana Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [21] † Miodomeris cossmanni Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [22] † Scalaricardita scalaris Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [23] † Scalaricardita laciarina Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [24] † Cyclocardia kickxi Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [25] † Cyclocardia chamaeformis Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [26] † Vimentum dilectum Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [27] † Pleuromeris tridentata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [28] † Pleuromeris decemcostata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [29] † Pteromeris perplana Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [30] † Glans trapezia Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [31] † Centrocardita aculeata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [32] † Carditamera floridana Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [33] † Byssomera affinis Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [34] † Cyclocardia borealis Taxon name last Modified on 04/29/24 | 1 | 146 | 145 | 0 | 0 | 0 | 0 | 1 |
| [35] † Cyclocardia compressa Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [36] † Cyclocardia mesembria Taxon name last Modified on 04/29/24 | 6 | 140 | 140 | 0 | 0 | 0 | 0 | 1 |
| [37] † Cyclocardia novangliae Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [38] † Cyclocardia cannada Taxon name last Modified on 04/29/24 | 0 | 147 | 146 | 0 | 0 | 0 | 0 | 1 |
| [39] † Cyclocardia dalek Taxon name last Modified on 04/29/24 | 0 | 147 | 146 | 0 | 0 | 0 | 0 | 1 |
| [40] † Cyclocardia castrana Taxon name last Modified on 04/29/24 | 1 | 146 | 145 | 0 | 0 | 0 | 0 | 1 |
| [41] † Cyclocardia granulata Taxon name last Modified on 04/29/24 | 0 | 148 | 146 | 0 | 0 | 0 | 0 | 1 |
| [42] † Cyclocardia moniliata Taxon name last Modified on 04/29/24 | 0 | 150 | 146 | 0 | 0 | 0 | 0 | 1 |
| [43] † Cyclocardia velutinus Taxon name last Modified on 04/29/24 | 0 | 147 | 146 | 0 | 0 | 0 | 0 | 1 |
| [44] † Cyclocardia awamoensis Taxon name last Modified on 04/29/24 | 0 | 147 | 146 | 0 | 0 | 0 | 0 | 1 |
| [45] † Cyclocardia nortensis Taxon name last Modified on 04/29/24 | 0 | 147 | 146 | 0 | 0 | 0 | 0 | 1 |
| [46] † Cyclocardia christiei Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [47] † Cyclocardia marama Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [48] † Crassicardia crassidens Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [49] † Cyclocardia ventricosa Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [50] † Crassicardia crebicostata Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
| [51] † Crassicardia isaotakii Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [52] † Crassicardia rjabininae Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [53] † Cyclocardia barbarensis Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [54] † Cyclocardia californica Taxon name last Modified on 04/29/24 | 0 | 147 | 146 | 0 | 0 | 0 | 0 | 1 |
| [55] † Cyclocardia hamiltonensis Taxon name last Modified on 04/29/24 | 1 | 147 | 145 | 0 | 0 | 0 | 0 | 1 |
| [56] † Cyclocardia yakatagensis Taxon name last Modified on 04/29/24 | 1 | 145 | 145 | 0 | 0 | 0 | 0 | 1 |
| [57] † Cyclocardia calvertensis Taxon name last Modified on 04/29/24 | 11 | 136 | 135 | 0 | 0 | 0 | 0 | 1 |
| [58] † Cyclocardia vautrotorum Taxon name last Modified on 04/29/24 | 11 | 135 | 135 | 0 | 0 | 0 | 0 | 1 |
| [59] † Cyclocardia occidentalis Taxon name last Modified on 04/29/24 | 0 | 146 | 146 | 0 | 0 | 0 | 0 | 1 |
Project downloads
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 3 | |
| Matrix downloads | 2 | Perez and Giachetti 2020 (2 downloads); |
| Media downloads | 1 | M903510 (1 download); |