Project 540: J. C. Lamsdell,P. A. Selden. 2015. Phylogenetic support for the monophyly of proetide trilobites. Lethaia. 48 (3):375–386.
Specimen: † Cyphaspis christyi (FMNH:PE6796)

Abstract

The monophyly of the order Proetida, the only trilobite group to survive the end- Devonian mass extinction, has been regularly questioned since its erection almost three decades ago. Through analysis of a novel phylogenetic dataset comprising 114 characters coded for 55 taxa, including both traditional members of the Proetida along with a number of other trilobite groups, the monophyly of proetide trilobites is rigorously tested for the first time. Proetida is shown to be monophyletic, united by the initial compound eye formation in early protaspids occurring at the lateral margin rather than the anterior margin, and the form of the protaspid glabella being tapering with a preglabellar field. A number of adult characters, including the possession of a quadrate or shield-shaped hypostome with angular posterior margins, the hypostome median body being divided by a deep groove that entirely traverses the median body, the presence of an enlarged thoracic spine on the sixth tergite, and a tergite count of between 7 and 10, also define the basal node. Hystricurid and dimeropygid trilobites are shown to resolve at the base of the group, while the remaining proetide taxa are divided between large proetoid and aulacopleuroid clades. Some taxa previously allied with Aulacopleuroidea, such as rorringtoniids and scharyiids, are retrieved as basal members of the Proetoidea.


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Article DOI: 10.1111/let.12113

Project DOI: 10.7934/P540, http://dx.doi.org/10.7934/P540
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 56 Taxa
  • 1 Specimen
  • 114 Characters
Total size of project's media files: 18 MB

Download Project SDD File
Total scored cells: 5249
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 114
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 258
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:114/0
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MorphoBank Project 540


    Members

    member name taxa specimens media chars character
    media
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    cell scorings
    (scored, NPA, "-")
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    James Lamsdell
    Project Administrator
    Last logged in 07/21/15
    5611114005249
    (4480, 0, 769)
    000


    Taxonomic Overview for Matrix 'M2775' (55 Taxa)

    taxon unscored
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    [1] † Eoredlichia intermedia
    Last Modified in 07/21/15
    118686017001
    [2] † Altiocculus harrisi
    Last Modified in 07/21/15
    277474013001
    [3] † Elrathia kingii
    Last Modified in 07/21/15
    99292013001
    [4] † Ehmaniella apolabella
    Last Modified in 07/21/15
    247676014001
    [5] † Lermontovia dzevanovskii
    Last Modified in 07/21/15
    277474013001
    [6] † Modocia kohli
    Last Modified in 07/21/15
    277777010001
    [7] † Marjumia typa
    Last Modified in 07/21/15
    277373014001
    [8] † Olenus wahlenbergi
    Last Modified in 07/21/15
    0102102012001
    [9] † Parabolina spinulosa
    Last Modified in 07/21/15
    0100100014001
    [10] † Leptoplastus crassicorne
    Last Modified in 07/21/15
    0100100014001
    [11] † Aphelaspis brachyphasis
    Last Modified in 07/21/15
    69393015001
    [12] † Crepicephalus buttsi montanaensis
    Last Modified in 07/21/15
    148787013001
    [13] † Coosella prolifica
    Last Modified in 07/21/15
    227777015001
    [14] † Tricrepicephalus taxanus
    Last Modified in 07/21/15
    277676011001
    [15] † Solenopleura canaliculata
    Last Modified in 07/21/15
    514949014001
    [16] † Paraselenopleura gregaria
    Last Modified in 07/21/15
    276969018001
    [17] † Kawina arnoldi
    Last Modified in 07/21/15
    187979017001
    [18] † Anacheirurus frederici
    Last Modified in 07/21/15
    218282011001
    [19] † Ceraurinella typa
    Last Modified in 07/21/15
    18878709001
    [20] † Ceraurinella nahanniensis
    Last Modified in 07/21/15
    3100100011001
    [21] † Parapilekia olesnaensis
    Last Modified in 07/21/15
    188484012001
    [22] † Calyptaulax annulata
    Last Modified in 07/21/15
    19494019001
    [23] † Flexicalymene senaria
    Last Modified in 07/21/15
    09595019001
    [24] † Isotelus parvirugosus
    Last Modified in 07/21/15
    08181033001
    [25] † Lopnorites rectispicatus
    Last Modified in 07/21/15
    277575012001
    [26] † Proceratopyge promisca
    Last Modified in 07/21/15
    277777010001
    [27] † Remopleurides caelatus
    Last Modified in 07/21/15
    88686020001
    [28] † Remopleurides eximius
    Last Modified in 07/21/15
    187878018001
    [29] † Asaphus expansus
    Last Modified in 07/21/15
    187272024001
    [30] † Harpides atlanticus
    Last Modified in 07/21/15
    63424209001
    [31] † Eskoharpes palanasus
    Last Modified in 07/21/15
    297171014001
    [32] † Harpes macrocephalus
    Last Modified in 07/21/15
    277272015001
    [33] † Eoharpes benignensis
    Last Modified in 07/21/15
    187979017001
    [34] † Entomaspis radiatus
    Last Modified in 07/21/15
    356565014001
    [35] † Holotrachelus punctillosus
    Last Modified in 07/21/15
    188080016001
    [36] † Aulacopleura wulongensis
    Last Modified in 07/21/15
    99292013001
    [37] † Aulacopleura konincki
    Last Modified in 07/21/15
    119090013001
    [38] † Aulacopleura longecornuta
    Last Modified in 07/21/15
    277676011001
    [39] † Cyphaspis dabrowni
    Last Modified in 07/21/15
    109494010001
    [40] † Otarion huddyi
    Last Modified in 07/21/15
    119292011001
    [41] † Maurotarion periergum
    Last Modified in 07/21/15
    18878709001
    [42] † Maurotarion struszi
    Last Modified in 07/21/15
    45646405001
    [43] † Brachymetopus germanicus
    Last Modified in 07/21/15
    277171016001
    [44] † Rorringtonia kennedyi
    Last Modified in 07/21/15
    277575012001
    [45] † Scharyia micropyga
    Last Modified in 07/21/15
    99191014001
    [46] † Bathyurellus nitidus
    Last Modified in 07/21/15
    207878016001
    [47] † Bathyurus ulu
    Last Modified in 07/21/15
    118686017001
    [48] † Proetus talenti
    Last Modified in 07/21/15
    79292015001
    [49] † Phillipsia belgica
    Last Modified in 07/21/15
    188484012001
    [50] † Gerastos tuberculatus macrocensis
    Last Modified in 07/21/15
    277575012001
    [51] † Tropidocoryphe bassei
    Last Modified in 07/21/15
    277171016001
    [52] † Hintzecurus paragenalatus
    Last Modified in 07/21/15
    148989011001
    [53] † Politicurus politus
    Last Modified in 07/21/15
    277676011001
    [54] † Toernquistia sanchezae
    Last Modified in 07/21/15
    510010009001
    [55] † Dimeropyge speyeri
    Last Modified in 07/21/15
    59393016001


    Project views

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    Total project views14820
    Project overview1958
    Media views2943Media search (2553 views); M333749 (390 views);
    Matrix views1358Matrix landing page (1230 views); Proetide monophyly (128 views);
    Taxon list6768
    Specimen list1279
    Bibliography514




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    Total downloads from project157
    Media downloads1M333749 (1 download);
    Matrix downloads6Proetide monophyly (6 downloads);
    Project downloads150