Project 540: J. C. Lamsdell, P. A. Selden. 2015. Phylogenetic support for the monophyly of proetide trilobites. Lethaia. 48 (3):375–386.
Specimen: † Cyphaspis christyi (FMNH:PE6796)

Abstract

The monophyly of the order Proetida, the only trilobite group to survive the end- Devonian mass extinction, has been regularly questioned since its erection almost three decades ago. Through analysis of a novel phylogenetic dataset comprising 114 characters coded for 55 taxa, including both traditional members of the Proetida along with a number of other trilobite groups, the monophyly of proetide trilobites is rigorously tested for the first time. Proetida is shown to be monophyletic, united by the initial compound eye formation in early protaspids occurring at the lateral margin rather than the anterior margin, and the form of the protaspid glabella being tapering with a preglabellar field. A number of adult characters, including the possession of a quadrate or shield-shaped hypostome with angular posterior margins, the hypostome median body being divided by a deep groove that entirely traverses the median body, the presence of an enlarged thoracic spine on the sixth tergite, and a tergite count of between 7 and 10, also define the basal node. Hystricurid and dimeropygid trilobites are shown to resolve at the base of the group, while the remaining proetide taxa are divided between large proetoid and aulacopleuroid clades. Some taxa previously allied with Aulacopleuroidea, such as rorringtoniids and scharyiids, are retrieved as basal members of the Proetoidea.


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Article DOI: 10.1111/let.12113

Project DOI: 10.7934/P540, http://dx.doi.org/10.7934/P540
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 56 Taxa
  • 1 Specimen
  • 114 Characters
Total size of project's media files: 18.06M

Download Project SDD File
Total scored cells: 5249
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 114
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 258
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:114/0
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MorphoBank Project 540

    Authors' Institutions

    • Natural History Museum, London

    • University of Kansas



    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
    cell
    medialabels
    rules
    James Lamsdell
    Project Administrator
    5611114005249
    (4480, 0, 769)
    000


    Taxonomic Overview for Matrix 'M2775' (55 Taxa)

    taxon unscored
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    scored
    cells
    no cell
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    NPA
    cells
    "-" cellscell images labels on
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    [1] † Eoredlichia intermedia
    Taxon name last Modified on 07/21/15
    118686017001
    [2] † Altiocculus harrisi
    Taxon name last Modified on 07/21/15
    277474013001
    [3] † Elrathia kingii
    Taxon name last Modified on 07/21/15
    99292013001
    [4] † Ehmaniella apolabella
    Taxon name last Modified on 07/21/15
    247676014001
    [5] † Lermontovia dzevanovskii
    Taxon name last Modified on 07/21/15
    277474013001
    [6] † Modocia kohli
    Taxon name last Modified on 07/21/15
    277777010001
    [7] † Marjumia typa
    Taxon name last Modified on 07/21/15
    277373014001
    [8] † Olenus wahlenbergi
    Taxon name last Modified on 07/21/15
    0102102012001
    [9] † Parabolina spinulosa
    Taxon name last Modified on 07/21/15
    0100100014001
    [10] † Leptoplastus crassicorne
    Taxon name last Modified on 07/21/15
    0100100014001
    [11] † Aphelaspis brachyphasis
    Taxon name last Modified on 07/21/15
    69393015001
    [12] † Crepicephalus buttsi montanaensis
    Taxon name last Modified on 07/21/15
    148787013001
    [13] † Coosella prolifica
    Taxon name last Modified on 07/21/15
    227777015001
    [14] † Tricrepicephalus taxanus
    Taxon name last Modified on 07/21/15
    277676011001
    [15] † Solenopleura canaliculata
    Taxon name last Modified on 07/21/15
    514949014001
    [16] † Paraselenopleura gregaria
    Taxon name last Modified on 07/21/15
    276969018001
    [17] † Kawina arnoldi
    Taxon name last Modified on 07/21/15
    187979017001
    [18] † Anacheirurus frederici
    Taxon name last Modified on 07/21/15
    218282011001
    [19] † Ceraurinella typa
    Taxon name last Modified on 07/21/15
    18878709001
    [20] † Ceraurinella nahanniensis
    Taxon name last Modified on 07/21/15
    3100100011001
    [21] † Parapilekia olesnaensis
    Taxon name last Modified on 07/21/15
    188484012001
    [22] † Calyptaulax annulata
    Taxon name last Modified on 07/21/15
    19494019001
    [23] † Flexicalymene senaria
    Taxon name last Modified on 07/21/15
    09595019001
    [24] † Isotelus parvirugosus
    Taxon name last Modified on 07/21/15
    08181033001
    [25] † Lopnorites rectispicatus
    Taxon name last Modified on 07/21/15
    277575012001
    [26] † Proceratopyge promisca
    Taxon name last Modified on 07/21/15
    277777010001
    [27] † Remopleurides caelatus
    Taxon name last Modified on 07/21/15
    88686020001
    [28] † Remopleurides eximius
    Taxon name last Modified on 07/21/15
    187878018001
    [29] † Asaphus expansus
    Taxon name last Modified on 07/21/15
    187272024001
    [30] † Harpides atlanticus
    Taxon name last Modified on 07/21/15
    63424209001
    [31] † Eskoharpes palanasus
    Taxon name last Modified on 07/21/15
    297171014001
    [32] † Harpes macrocephalus
    Taxon name last Modified on 07/21/15
    277272015001
    [33] † Eoharpes benignensis
    Taxon name last Modified on 07/21/15
    187979017001
    [34] † Entomaspis radiatus
    Taxon name last Modified on 07/21/15
    356565014001
    [35] † Holotrachelus punctillosus
    Taxon name last Modified on 07/21/15
    188080016001
    [36] † Aulacopleura wulongensis
    Taxon name last Modified on 07/21/15
    99292013001
    [37] † Aulacopleura konincki
    Taxon name last Modified on 07/21/15
    119090013001
    [38] † Aulacopleura longecornuta
    Taxon name last Modified on 07/21/15
    277676011001
    [39] † Cyphaspis dabrowni
    Taxon name last Modified on 07/21/15
    109494010001
    [40] † Otarion huddyi
    Taxon name last Modified on 07/21/15
    119292011001
    [41] † Maurotarion periergum
    Taxon name last Modified on 07/21/15
    18878709001
    [42] † Maurotarion struszi
    Taxon name last Modified on 07/21/15
    45646405001
    [43] † Brachymetopus germanicus
    Taxon name last Modified on 07/21/15
    277171016001
    [44] † Rorringtonia kennedyi
    Taxon name last Modified on 07/21/15
    277575012001
    [45] † Scharyia micropyga
    Taxon name last Modified on 07/21/15
    99191014001
    [46] † Bathyurellus nitidus
    Taxon name last Modified on 07/21/15
    207878016001
    [47] † Bathyurus ulu
    Taxon name last Modified on 07/21/15
    118686017001
    [48] † Proetus talenti
    Taxon name last Modified on 07/21/15
    79292015001
    [49] † Phillipsia belgica
    Taxon name last Modified on 07/21/15
    188484012001
    [50] † Gerastos tuberculatus macrocensis
    Taxon name last Modified on 07/21/15
    277575012001
    [51] † Tropidocoryphe bassei
    Taxon name last Modified on 07/21/15
    277171016001
    [52] † Hintzecurus paragenalatus
    Taxon name last Modified on 07/21/15
    148989011001
    [53] † Politicurus politus
    Taxon name last Modified on 07/21/15
    277676011001
    [54] † Toernquistia sanchezae
    Taxon name last Modified on 07/21/15
    510010009001
    [55] † Dimeropyge speyeri
    Taxon name last Modified on 07/21/15
    59393016001


    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project353
    Media downloads1M333749 (1 download);
    Matrix downloads19Proetide monophyly (19 downloads);
    Project downloads333