Project 540: J. C. Lamsdell, P. A. Selden. 2015. Phylogenetic support for the monophyly of proetide trilobites. Lethaia. 48 (3):375–386.
Abstract
The monophyly of the order Proetida, the only trilobite group to survive the end- Devonian mass extinction, has been regularly questioned since its erection almost three decades ago. Through analysis of a novel phylogenetic dataset comprising 114 characters coded for 55 taxa, including both traditional members of the Proetida along with a number of other trilobite groups, the monophyly of proetide trilobites is rigorously tested for the first time. Proetida is shown to be monophyletic, united by the initial compound eye formation in early protaspids occurring at the lateral margin rather than the anterior margin, and the form of the protaspid glabella being tapering with a preglabellar field. A number of adult characters, including the possession of a quadrate or shield-shaped hypostome with angular posterior margins, the hypostome median body being divided by a deep groove that entirely traverses the median body, the presence of an enlarged thoracic spine on the sixth tergite, and a tergite count of between 7 and 10, also define the basal node. Hystricurid and dimeropygid trilobites are shown to resolve at the base of the group, while the remaining proetide taxa are divided between large proetoid and aulacopleuroid clades. Some taxa previously allied with Aulacopleuroidea, such as rorringtoniids and scharyiids, are retrieved as basal members of the Proetoidea.Read the article »
Article DOI: 10.1111/let.12113
Project DOI: 10.7934/P540, http://dx.doi.org/10.7934/P540
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 5249 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 114 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 258 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:114/0 |
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MorphoBank Project 540
MorphoBank Project 540
- Creation Date:
27 June 2011 - Publication Date:
21 July 2015 - Project views: 34119
- Media downloads: 1
- Matrix downloads: 14
Authors' Institutions
- Natural History Museum, London
- University of Kansas
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
James Lamsdell Project Administrator | 56 | 1 | 1 | 114 | 0 | 0 | 5249 (4480, 0, 769) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M2775' (55 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Eoredlichia intermedia Last Modified in 07/21/15 | 11 | 86 | 86 | 0 | 17 | 0 | 0 | 1 |
[2] † Altiocculus harrisi Last Modified in 07/21/15 | 27 | 74 | 74 | 0 | 13 | 0 | 0 | 1 |
[3] † Elrathia kingii Last Modified in 07/21/15 | 9 | 92 | 92 | 0 | 13 | 0 | 0 | 1 |
[4] † Ehmaniella apolabella Last Modified in 07/21/15 | 24 | 76 | 76 | 0 | 14 | 0 | 0 | 1 |
[5] † Lermontovia dzevanovskii Last Modified in 07/21/15 | 27 | 74 | 74 | 0 | 13 | 0 | 0 | 1 |
[6] † Modocia kohli Last Modified in 07/21/15 | 27 | 77 | 77 | 0 | 10 | 0 | 0 | 1 |
[7] † Marjumia typa Last Modified in 07/21/15 | 27 | 73 | 73 | 0 | 14 | 0 | 0 | 1 |
[8] † Olenus wahlenbergi Last Modified in 07/21/15 | 0 | 102 | 102 | 0 | 12 | 0 | 0 | 1 |
[9] † Parabolina spinulosa Last Modified in 07/21/15 | 0 | 100 | 100 | 0 | 14 | 0 | 0 | 1 |
[10] † Leptoplastus crassicorne Last Modified in 07/21/15 | 0 | 100 | 100 | 0 | 14 | 0 | 0 | 1 |
[11] † Aphelaspis brachyphasis Last Modified in 07/21/15 | 6 | 93 | 93 | 0 | 15 | 0 | 0 | 1 |
[12] † Crepicephalus buttsi montanaensis Last Modified in 07/21/15 | 14 | 87 | 87 | 0 | 13 | 0 | 0 | 1 |
[13] † Coosella prolifica Last Modified in 07/21/15 | 22 | 77 | 77 | 0 | 15 | 0 | 0 | 1 |
[14] † Tricrepicephalus taxanus Last Modified in 07/21/15 | 27 | 76 | 76 | 0 | 11 | 0 | 0 | 1 |
[15] † Solenopleura canaliculata Last Modified in 07/21/15 | 51 | 49 | 49 | 0 | 14 | 0 | 0 | 1 |
[16] † Paraselenopleura gregaria Last Modified in 07/21/15 | 27 | 69 | 69 | 0 | 18 | 0 | 0 | 1 |
[17] † Kawina arnoldi Last Modified in 07/21/15 | 18 | 79 | 79 | 0 | 17 | 0 | 0 | 1 |
[18] † Anacheirurus frederici Last Modified in 07/21/15 | 21 | 82 | 82 | 0 | 11 | 0 | 0 | 1 |
[19] † Ceraurinella typa Last Modified in 07/21/15 | 18 | 87 | 87 | 0 | 9 | 0 | 0 | 1 |
[20] † Ceraurinella nahanniensis Last Modified in 07/21/15 | 3 | 100 | 100 | 0 | 11 | 0 | 0 | 1 |
[21] † Parapilekia olesnaensis Last Modified in 07/21/15 | 18 | 84 | 84 | 0 | 12 | 0 | 0 | 1 |
[22] † Calyptaulax annulata Last Modified in 07/21/15 | 1 | 94 | 94 | 0 | 19 | 0 | 0 | 1 |
[23] † Flexicalymene senaria Last Modified in 07/21/15 | 0 | 95 | 95 | 0 | 19 | 0 | 0 | 1 |
[24] † Isotelus parvirugosus Last Modified in 07/21/15 | 0 | 81 | 81 | 0 | 33 | 0 | 0 | 1 |
[25] † Lopnorites rectispicatus Last Modified in 07/21/15 | 27 | 75 | 75 | 0 | 12 | 0 | 0 | 1 |
[26] † Proceratopyge promisca Last Modified in 07/21/15 | 27 | 77 | 77 | 0 | 10 | 0 | 0 | 1 |
[27] † Remopleurides caelatus Last Modified in 07/21/15 | 8 | 86 | 86 | 0 | 20 | 0 | 0 | 1 |
[28] † Remopleurides eximius Last Modified in 07/21/15 | 18 | 78 | 78 | 0 | 18 | 0 | 0 | 1 |
[29] † Asaphus expansus Last Modified in 07/21/15 | 18 | 72 | 72 | 0 | 24 | 0 | 0 | 1 |
[30] † Harpides atlanticus Last Modified in 07/21/15 | 63 | 42 | 42 | 0 | 9 | 0 | 0 | 1 |
[31] † Eskoharpes palanasus Last Modified in 07/21/15 | 29 | 71 | 71 | 0 | 14 | 0 | 0 | 1 |
[32] † Harpes macrocephalus Last Modified in 07/21/15 | 27 | 72 | 72 | 0 | 15 | 0 | 0 | 1 |
[33] † Eoharpes benignensis Last Modified in 07/21/15 | 18 | 79 | 79 | 0 | 17 | 0 | 0 | 1 |
[34] † Entomaspis radiatus Last Modified in 07/21/15 | 35 | 65 | 65 | 0 | 14 | 0 | 0 | 1 |
[35] † Holotrachelus punctillosus Last Modified in 07/21/15 | 18 | 80 | 80 | 0 | 16 | 0 | 0 | 1 |
[36] † Aulacopleura wulongensis Last Modified in 07/21/15 | 9 | 92 | 92 | 0 | 13 | 0 | 0 | 1 |
[37] † Aulacopleura konincki Last Modified in 07/21/15 | 11 | 90 | 90 | 0 | 13 | 0 | 0 | 1 |
[38] † Aulacopleura longecornuta Last Modified in 07/21/15 | 27 | 76 | 76 | 0 | 11 | 0 | 0 | 1 |
[39] † Cyphaspis dabrowni Last Modified in 07/21/15 | 10 | 94 | 94 | 0 | 10 | 0 | 0 | 1 |
[40] † Otarion huddyi Last Modified in 07/21/15 | 11 | 92 | 92 | 0 | 11 | 0 | 0 | 1 |
[41] † Maurotarion periergum Last Modified in 07/21/15 | 18 | 87 | 87 | 0 | 9 | 0 | 0 | 1 |
[42] † Maurotarion struszi Last Modified in 07/21/15 | 45 | 64 | 64 | 0 | 5 | 0 | 0 | 1 |
[43] † Brachymetopus germanicus Last Modified in 07/21/15 | 27 | 71 | 71 | 0 | 16 | 0 | 0 | 1 |
[44] † Rorringtonia kennedyi Last Modified in 07/21/15 | 27 | 75 | 75 | 0 | 12 | 0 | 0 | 1 |
[45] † Scharyia micropyga Last Modified in 07/21/15 | 9 | 91 | 91 | 0 | 14 | 0 | 0 | 1 |
[46] † Bathyurellus nitidus Last Modified in 07/21/15 | 20 | 78 | 78 | 0 | 16 | 0 | 0 | 1 |
[47] † Bathyurus ulu Last Modified in 07/21/15 | 11 | 86 | 86 | 0 | 17 | 0 | 0 | 1 |
[48] † Proetus talenti Last Modified in 07/21/15 | 7 | 92 | 92 | 0 | 15 | 0 | 0 | 1 |
[49] † Phillipsia belgica Last Modified in 07/21/15 | 18 | 84 | 84 | 0 | 12 | 0 | 0 | 1 |
[50] † Gerastos tuberculatus macrocensis Last Modified in 07/21/15 | 27 | 75 | 75 | 0 | 12 | 0 | 0 | 1 |
[51] † Tropidocoryphe bassei Last Modified in 07/21/15 | 27 | 71 | 71 | 0 | 16 | 0 | 0 | 1 |
[52] † Hintzecurus paragenalatus Last Modified in 07/21/15 | 14 | 89 | 89 | 0 | 11 | 0 | 0 | 1 |
[53] † Politicurus politus Last Modified in 07/21/15 | 27 | 76 | 76 | 0 | 11 | 0 | 0 | 1 |
[54] † Toernquistia sanchezae Last Modified in 07/21/15 | 5 | 100 | 100 | 0 | 9 | 0 | 0 | 1 |
[55] † Dimeropyge speyeri Last Modified in 07/21/15 | 5 | 93 | 93 | 0 | 16 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 34119 | |
Project overview | 3396 | |
Media views | 5898 | Media search (5076 views); M333749 (822 views); |
Matrix views | 2374 | Matrix landing page (2108 views); Proetide monophyly (266 views); |
Taxon list | 18495 | |
Specimen list | 2819 | |
Bibliography | 1127 | |
Documents list | 4 | |
Views for media list | 6 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 343 | |
Media downloads | 1 | M333749 (1 download); |
Matrix downloads | 14 | Proetide monophyly (14 downloads); |
Project downloads | 328 |