Project 5400: R. J. O’Flynn, M. Williams, Y. Liu, X.-G. Hou, J. Guo, G. D. Edgecombe. 2024. The early Cambrian Kuamaia lata, an artiopodan euarthropod with a raptorial frontal appendage. Journal of Paleontology. (In Press)
Abstract
Phylogenetic analysis demonstrates that Kuamaia lata Hou, 1987, a helmetiid euarthropod from the lower Cambrian (Series 2, Stage 3) Chengjiang Konservat-Lagerstätte, nests robustly within Artiopoda Hou and Bergström, 1997, the euarthropod clade including trilobitomorphs. Microtomography of new specimens of K. lata reveals details of morphology, notably a six-segmented head and raptorial frontal appendages, the latter contrasting with filiform antennae considered to be a diagnostic character of Artiopoda. Phylogenetic analyses demonstrate that a raptorial frontal appendage is a symplesiomorphy for upper stem-group euarthropods, retained across a swathe of tree space, but evolved secondarily in K. lata from an antenna within Artiopoda. The phylogenetic position of K. lata adds support to a six-segmented head being an ancestral state for upper stem- and crown-group euarthropods.Project DOI: 10.7934/P5400, http://dx.doi.org/10.7934/P5400
This project contains | Matrices |
---|---|
Download Project SDD File | Total scored cells: 22386 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 283 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 643 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:283/0 |
Currently Viewing:
MorphoBank Project 5400
MorphoBank Project 5400
- Creation Date:
21 July 2024 - Publication Date:
21 July 2024
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Robert O'Flynn Project Administrator | 85 | 1 | 1 | 283 | 0 | 0 | 22386 (14402, 0, 7984) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M29994' (85 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Priapulus caudatus Taxon name last Modified on 07/21/24 | 2 | 53 | 53 | 0 | 228 | 0 | 0 | 1 |
[2] † Cricocosmia jinningensis Taxon name last Modified on 07/21/24 | 7 | 54 | 54 | 0 | 222 | 0 | 0 | 1 |
[3] tardigrada Taxon name last Modified on 07/21/24 | 6 | 107 | 107 | 0 | 170 | 0 | 0 | 1 |
[4] † Microdictyon sinicum Taxon name last Modified on 07/21/24 | 14 | 116 | 116 | 0 | 153 | 0 | 0 | 1 |
[5] † Paucipodia inermis Taxon name last Modified on 07/21/24 | 16 | 109 | 109 | 0 | 158 | 0 | 0 | 1 |
[6] † Cardiodictyon catenulum Taxon name last Modified on 07/21/24 | 17 | 114 | 114 | 0 | 152 | 0 | 0 | 1 |
[7] † Hallucigenia sparsa Taxon name last Modified on 07/21/24 | 11 | 120 | 120 | 0 | 152 | 0 | 0 | 1 |
[8] † Hallucigenia fortis Taxon name last Modified on 07/21/24 | 14 | 117 | 117 | 0 | 152 | 0 | 0 | 1 |
[9] † Collinsium ciliosum Taxon name last Modified on 07/21/24 | 18 | 114 | 114 | 0 | 151 | 0 | 0 | 1 |
[10] † Luolishania longicruris Taxon name last Modified on 07/21/24 | 14 | 118 | 118 | 0 | 151 | 0 | 0 | 1 |
[11] † Onychodictyon ferox Taxon name last Modified on 07/21/24 | 14 | 117 | 117 | 0 | 152 | 0 | 0 | 1 |
[12] † Aysheaia pedunculata Taxon name last Modified on 07/21/24 | 12 | 114 | 114 | 0 | 157 | 0 | 0 | 1 |
[13] † Antennacanthopodia gracilis Taxon name last Modified on 07/21/24 | 13 | 112 | 112 | 0 | 158 | 0 | 0 | 1 |
[14] onychophora Taxon name last Modified on 07/21/24 | 5 | 117 | 117 | 0 | 161 | 0 | 0 | 1 |
[15] † Megadictyon haikouensis Taxon name last Modified on 07/21/24 | 17 | 108 | 108 | 0 | 158 | 0 | 0 | 1 |
[16] † Pambdelurion whittingtoni Taxon name last Modified on 07/21/24 | 40 | 109 | 109 | 0 | 134 | 0 | 0 | 1 |
[17] † Kerygmachela kierkegaardi Taxon name last Modified on 07/21/24 | 43 | 110 | 110 | 0 | 130 | 0 | 0 | 1 |
[18] † Opabinia regalis Taxon name last Modified on 07/21/24 | 43 | 107 | 107 | 0 | 133 | 0 | 0 | 1 |
[19] † Anomalocaris canadensis Taxon name last Modified on 07/21/24 | 23 | 142 | 142 | 0 | 118 | 0 | 0 | 1 |
[20] † Lyrarapax unguispinus Taxon name last Modified on 07/21/24 | 26 | 139 | 139 | 0 | 118 | 0 | 0 | 1 |
[21] † Amplectobelua symbrachiata Taxon name last Modified on 07/21/24 | 18 | 147 | 147 | 0 | 118 | 0 | 0 | 1 |
[22] † Peytoia nathorsti Taxon name last Modified on 07/21/24 | 25 | 140 | 140 | 0 | 118 | 0 | 0 | 1 |
[23] † Stanleycaris hirpex Taxon name last Modified on 07/21/24 | 29 | 137 | 137 | 0 | 117 | 0 | 0 | 1 |
[24] † Aegirocassis benmoulai Taxon name last Modified on 07/21/24 | 67 | 102 | 102 | 0 | 114 | 0 | 0 | 1 |
[25] † Hurdia victoria Taxon name last Modified on 07/21/24 | 31 | 134 | 134 | 0 | 118 | 0 | 0 | 1 |
[26] † Schinderhannes bartelsi Taxon name last Modified on 07/21/24 | 46 | 121 | 121 | 0 | 116 | 0 | 0 | 1 |
[27] † Fengzhengia mamingae Taxon name last Modified on 07/21/24 | 45 | 168 | 168 | 0 | 70 | 0 | 0 | 1 |
[28] † Kylinxia zhangi Taxon name last Modified on 07/21/24 | 12 | 203 | 203 | 0 | 68 | 0 | 0 | 1 |
[29] † Kiisortoqia soperi Taxon name last Modified on 07/21/24 | 32 | 172 | 172 | 0 | 79 | 0 | 0 | 1 |
[30] † Bushizheia yangi Taxon name last Modified on 07/21/24 | 32 | 175 | 175 | 0 | 76 | 0 | 0 | 1 |
[31] † Habelia optata Taxon name last Modified on 07/21/24 | 14 | 198 | 198 | 0 | 71 | 0 | 0 | 1 |
[32] † Mollisonia plenovenatrix Taxon name last Modified on 07/21/24 | 14 | 200 | 200 | 0 | 69 | 0 | 0 | 1 |
[33] Limulus polyphemus Taxon name last Modified on 07/21/24 | 1 | 210 | 210 | 0 | 72 | 0 | 0 | 1 |
[34] † Dibasterium durgae Taxon name last Modified on 07/21/24 | 24 | 187 | 187 | 0 | 72 | 0 | 0 | 1 |
[35] † Offacolus kingi Taxon name last Modified on 07/21/24 | 24 | 187 | 187 | 0 | 72 | 0 | 0 | 1 |
[36] † Haikoucaris ercaiensis Taxon name last Modified on 07/21/24 | 17 | 197 | 197 | 0 | 69 | 0 | 0 | 1 |
[37] † Yawunik kootenayi Taxon name last Modified on 07/21/24 | 7 | 208 | 208 | 0 | 68 | 0 | 0 | 1 |
[38] † Alalcomenaeus cambricus Taxon name last Modified on 07/21/24 | 6 | 209 | 209 | 0 | 68 | 0 | 0 | 1 |
[39] † Leanchoilia illecebrosa Taxon name last Modified on 07/21/24 | 6 | 209 | 209 | 0 | 68 | 0 | 0 | 1 |
[40] † Leanchoilia superlata Taxon name last Modified on 07/21/24 | 7 | 208 | 208 | 0 | 68 | 0 | 0 | 1 |
[41] † Fortiforceps foliosa Taxon name last Modified on 07/21/24 | 7 | 206 | 206 | 0 | 70 | 0 | 0 | 1 |
[42] † Tanglangia caudata Taxon name last Modified on 07/21/24 | 40 | 175 | 175 | 0 | 68 | 0 | 0 | 1 |
[43] † Yohoia tenuis Taxon name last Modified on 07/21/24 | 8 | 205 | 205 | 0 | 70 | 0 | 0 | 1 |
[44] † Sklerolibyon maomima Taxon name last Modified on 07/21/24 | 11 | 202 | 202 | 0 | 70 | 0 | 0 | 1 |
[45] † Jianfengia multisegmentalis Taxon name last Modified on 07/21/24 | 8 | 205 | 205 | 0 | 70 | 0 | 0 | 1 |
[46] † Forfexicaris valida Taxon name last Modified on 07/21/24 | 89 | 135 | 135 | 0 | 59 | 0 | 0 | 1 |
[47] † Occacaris oviformis Taxon name last Modified on 07/21/24 | 63 | 161 | 161 | 0 | 59 | 0 | 0 | 1 |
[48] † Surusicaris elegans Taxon name last Modified on 07/21/24 | 27 | 191 | 191 | 0 | 65 | 0 | 0 | 1 |
[49] † Isoxys auritus Taxon name last Modified on 07/21/24 | 20 | 198 | 198 | 0 | 65 | 0 | 0 | 1 |
[50] † Isoxys volucris Taxon name last Modified on 07/21/24 | 67 | 154 | 154 | 0 | 62 | 0 | 0 | 1 |
[51] † Isoxys curvirostratus Taxon name last Modified on 07/21/24 | 19 | 199 | 199 | 0 | 65 | 0 | 0 | 1 |
[52] † Isoxys acutangulus Taxon name last Modified on 07/21/24 | 23 | 195 | 195 | 0 | 65 | 0 | 0 | 1 |
[53] † Retifacies abnormalis Taxon name last Modified on 07/21/24 | 7 | 199 | 199 | 0 | 77 | 0 | 0 | 1 |
[54] † Sinoburius lunaris Taxon name last Modified on 07/21/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[55] † Cindarella eucalla Taxon name last Modified on 07/21/24 | 5 | 202 | 202 | 0 | 76 | 0 | 0 | 1 |
[56] † Xandarella spectaculum Taxon name last Modified on 07/21/24 | 5 | 202 | 202 | 0 | 76 | 0 | 0 | 1 |
[57] † Olenoides serratus Taxon name last Modified on 07/21/24 | 4 | 203 | 203 | 0 | 76 | 0 | 0 | 1 |
[58] † Hongshiyanaspis yiliangensis Taxon name last Modified on 07/21/24 | 3 | 204 | 204 | 0 | 76 | 0 | 0 | 1 |
[59] † Eoredlichia intermedia Taxon name last Modified on 07/21/24 | 4 | 203 | 203 | 0 | 76 | 0 | 0 | 1 |
[60] † Misszhouia longicaudata Taxon name last Modified on 07/21/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[61] † Naraoia compacta Taxon name last Modified on 07/21/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[62] † Naraoia spinosa Taxon name last Modified on 07/21/24 | 15 | 192 | 192 | 0 | 76 | 0 | 0 | 1 |
[63] † Arthroaspis bergstroemi Taxon name last Modified on 07/21/24 | 13 | 195 | 195 | 0 | 75 | 0 | 0 | 1 |
[64] † Tegopelte gigas Taxon name last Modified on 07/21/24 | 10 | 197 | 197 | 0 | 76 | 0 | 0 | 1 |
[65] † Saperion glumaceum Taxon name last Modified on 07/21/24 | 6 | 201 | 201 | 0 | 76 | 0 | 0 | 1 |
[66] † Helmetia expansa Taxon name last Modified on 07/21/24 | 73 | 139 | 139 | 0 | 71 | 0 | 0 | 1 |
[67] † Kuamaia lata Taxon name last Modified on 07/21/24 | 7 | 202 | 202 | 0 | 74 | 0 | 0 | 1 |
[68] † Chengjiangocaris kunmingensis Taxon name last Modified on 07/21/24 | 6 | 204 | 204 | 0 | 73 | 0 | 0 | 1 |
[69] † Alacaris mirabilis Taxon name last Modified on 07/21/24 | 7 | 203 | 203 | 0 | 73 | 0 | 0 | 1 |
[70] † Liangwangshania biloba Taxon name last Modified on 07/21/24 | 10 | 199 | 199 | 0 | 74 | 0 | 0 | 1 |
[71] † Guangweicaris spinatus Taxon name last Modified on 07/21/24 | 12 | 198 | 198 | 0 | 73 | 0 | 0 | 1 |
[72] † Fuxianhuia protensa Taxon name last Modified on 07/21/24 | 3 | 207 | 207 | 0 | 73 | 0 | 0 | 1 |
[73] Narceus americanus Taxon name last Modified on 07/21/24 | 1 | 192 | 192 | 0 | 90 | 0 | 0 | 1 |
[74] Scutigera coleoptrata Taxon name last Modified on 07/21/24 | 3 | 190 | 190 | 0 | 90 | 0 | 0 | 1 |
[75] Artemia salina Taxon name last Modified on 07/21/24 | 2 | 212 | 212 | 0 | 69 | 0 | 0 | 1 |
[76] † Yicaris dianensis Taxon name last Modified on 07/21/24 | 7 | 207 | 207 | 0 | 69 | 0 | 0 | 1 |
[77] † Hesslandona unisulcata Taxon name last Modified on 07/21/24 | 17 | 197 | 197 | 0 | 69 | 0 | 0 | 1 |
[78] † Martinssonia elongata Taxon name last Modified on 07/21/24 | 10 | 204 | 204 | 0 | 69 | 0 | 0 | 1 |
[79] † Waptia fieldensis Taxon name last Modified on 07/21/24 | 7 | 211 | 211 | 0 | 65 | 0 | 0 | 1 |
[80] † Clypecaris serrata Taxon name last Modified on 07/21/24 | 48 | 173 | 173 | 0 | 62 | 0 | 0 | 1 |
[81] † Canadaspis perfecta Taxon name last Modified on 07/21/24 | 12 | 206 | 206 | 0 | 65 | 0 | 0 | 1 |
[82] † Tokummia katalepsis Taxon name last Modified on 07/21/24 | 16 | 203 | 203 | 0 | 64 | 0 | 0 | 1 |
[83] † Branchiocaris pretiosa Taxon name last Modified on 07/21/24 | 23 | 196 | 196 | 0 | 64 | 0 | 0 | 1 |
[84] † Odaraia alata Taxon name last Modified on 07/21/24 | 64 | 159 | 159 | 0 | 60 | 0 | 0 | 1 |
[85] † Jugatacaris agilis Taxon name last Modified on 07/21/24 | 30 | 191 | 191 | 0 | 62 | 0 | 0 | 1 |