Project 568: J. C. Lamsdell, İ. Hoşgör, P. A. Selden. 2013. A New Ordovician Eurypterid (Arthropoda: Chelicerata) from South East Turkey: Evidence for a Cryptic Ordovician Record of Eurypterida. Gondwana Research. 23:354–366.
Specimen: † Paraeurypterus anatoliensis (MTANHMSETR:10-Ä°Z-01-1)

Abstract

A new species of eurypterid, Paraeurypterus anatoliensis gen. et sp. nov., is described from the Upper Ordovician (Katian) Şort Tepe Formation of southeast Turkey. The single specimen, preserving the carapace, mesosoma and fragments of appendages, appears morphologically intermediate between the eurypteroid families Dolichopteridae and Eurypteridae. P. anatoliensis retains the plesiomorphic conditions of crescentic eyes with enlarged palpebral lobes and a quadrate carapace with ornamentation consisting of small pustules but also displaying the derived characteristics of genal facets and a row of large acicular scales across the posterior of each tergite. Phylogenetic analysis incorporating each of the major eurypterine clades and all Eurypterina having a three-segmented genital operculum (the triploperculate condition) resolves eurypteroids to be an unnatural group, with Dolichopteridae and Eurypteridae forming part of a grade leading to diploperculate Eurypterina. P. anatoliensis is intermediate between the two eurypteroid families, as is ‘Eurypterus’ minor from the Pentland Hills of Scotland, which is shown to be a distinct genus and assigned to Pentlandopterus gen. nov. Using the phylogenetic topology to infer ghost ranges for each of the major eurypterid clades reveals that the majority of eurypterid superfamilies must have originated by the Katian, indicating a largely unsampled record of Ordovician eurypterids. The occurrence of poor dispersers such as Paraeurypterus in the Ordovician of Gondwana is puzzling, and it has been suggested that they dispersed to the continent during periods of sea level lowstand in the Sandbian and Hirnantian, however this does not explain the lack of Ordovician species in North America and Europe, given the well-sampled nature of these continents, and an alternative is proposed whereby eurypterids originated in Gondwana and radiated out to Laurentia and Baltica in the late Ordovician and early Silurian, thus explaining their sudden appearance in the European and North American rock record.


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Project DOI: 10.7934/P568, http://dx.doi.org/10.7934/P568
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 45 Taxa
  • 1 Specimen
  • 81 Characters
Total size of project's media files: 70 MB

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Total scored cells: 2970
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 81
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 195
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:81/0
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MorphoBank Project 568


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    James Lamsdell
    Project Administrator
    Last logged in 11/28/12
    451108100002970
    (2842, 0, 128)
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    Taxonomic Overview for Matrix 'M1387' (45 Taxa)

    taxon unscored
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    [1] † Weinbergina opitzi
    Last Modified in 12/05/13
    16767013001
    [2] † Brachyopterus stubblefieldi
    Last Modified in 12/05/13
    13606008001
    [3] † Parastylonurus ornatus
    Last Modified in 12/05/13
    1757505001
    [4] † Stoermeropterus conicus
    Last Modified in 12/05/13
    0797902001
    [5] † Stoermeropterus latus
    Last Modified in 12/05/13
    14656502001
    [6] † Stoermeropterus nodosus
    Last Modified in 12/05/13
    36444401001
    [7] † Vinetopterus struvei
    Last Modified in 12/05/13
    4757502001
    [8] † Vinetopterus martini
    Last Modified in 12/05/13
    17626202001
    [9] † Moselopterus ancylotelson
    Last Modified in 12/05/13
    1787802001
    [10] † Moselopterus elongatus
    Last Modified in 12/05/13
    4757502001
    [11] † Onychopterella augusti
    Last Modified in 12/05/13
    15636303001
    [12] † Onychopterella kokomoensis
    Last Modified in 12/05/13
    5737303001
    [13] † Dolichopterus macrocheirus
    Last Modified in 12/05/13
    0808001001
    [14] † Eurypterus remipes
    Last Modified in 12/05/13
    0808001001
    [15] † Hughmilleria socialis
    Last Modified in 12/05/13
    2767603001
    [16] † Tylopterella boylei
    Last Modified in 12/05/13
    46343401001
    [17] † Megalograptus ohioensis
    Last Modified in 12/05/13
    0757506001
    [18] † Mixopterus kiaeri
    Last Modified in 12/05/13
    3767602001
    [19] † Strobilopterus princetonii
    Last Modified in 12/05/13
    12686801001
    [20] † Erieopterus microphthalmus
    Last Modified in 12/05/13
    1797901001
    [21] † Rhenopterus diensti
    Last Modified in 12/05/13
    0757506001
    [22] † Octoberaspis ushakovi
    Last Modified in 12/05/13
    16575708001
    [23] † Loganamaraspis dunlopi
    Last Modified in 12/05/13
    31444406001
    [24] † Diploaspis casteri
    Last Modified in 12/05/13
    18565607001
    [25] † Paraeurypterus anatoliensis
    Last Modified in 12/05/13
    43383800001
    [26] † Pentlandopterus minor
    Last Modified in 12/05/13
    41404000001
    [27] † Chasmataspis laurencii
    Last Modified in 12/05/13
    214949011001
    [28] † Buffalopterus pustulosus
    Last Modified in 12/05/13
    34454502001
    [29] † Syntomopterella richardsoni
    Last Modified in 12/05/13
    47333301001
    [30] † Dolichopterus jewetti
    Last Modified in 12/05/13
    4767601001
    [31] † Ruedemannipterus stylonuroides
    Last Modified in 12/05/13
    38424201001
    [32] † Clarkeipterus testudineus
    Last Modified in 12/05/13
    46343401001
    [33] † Clarkeipterus otisius
    Last Modified in 12/05/13
    67131301001
    [34] † Erieopterus eriensis
    Last Modified in 12/05/13
    10717100001
    [35] † Eurypterus henkeni
    Last Modified in 12/05/13
    9717101001
    [36] † Eurypterus tetragonophalmus
    Last Modified in 12/05/13
    0808001001
    [37] † Eurypterus leopoldi
    Last Modified in 12/05/13
    36454500001
    [38] † Eurypterus pittsfordensis
    Last Modified in 12/05/13
    8737300001
    [39] † Eurypterus henningsmoeni
    Last Modified in 12/05/13
    11696901001
    [40] † Orcanopterus manitoulinensis
    Last Modified in 12/05/13
    13666602001
    [41] † Pterygotus anglicus
    Last Modified in 12/05/13
    1747406001
    [42] † Adelophthalmus sievertsi
    Last Modified in 12/05/13
    1757505001
    [43] † Eurypterus dekayi
    Last Modified in 12/05/13
    1797901001
    [44] † Carcinosoma newlini
    Last Modified in 12/05/13
    3757503001
    [45] † Nanahughmilleria norvegica
    Last Modified in 12/05/13
    1787802001


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    Total project views19828
    Project overview2657
    Matrix views1893Matrix landing page (1618 views); Turkish euypterid (275 views);
    Media views3873Media search (3384 views); M94106 (489 views);
    Specimen list2039
    Taxon list9365
    Documents list1




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    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project194
    Project downloads185
    Matrix downloads9Turkish euypterid (9 downloads);