Project 581: A. H. Turner, J. J. W. Sertich. 2010. Phylogenetic History of Simosuchus clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology. 30 (sp1):177-236.
Specimen: Simosuchus clarki (UA:8679)
View: dorsal

Abstract

The bizarre crocodyliform Simosuchus clarki shares several derived cranial characteristics with basal notosuchian taxa. In the original phylogenetic analysis assessing placement of Simosuchus among other crocodyliforms, it was recovered as a basal notosuchian. Additional preparation of the holotype skull and postcranium, and discovery of additional specimens, provides new information on the suite of morphologies in this notosuchian. Morphological analysis of all available material of S. clarki yields 60 autapomorphies distributed across the skeleton. To provide an updated test of the phylogeny of Notosuchia and placement of Simosuchus within the clade, we added seven new characters and expanded taxon sampling to include recently described notosuchians. A phylogenetic analysis of 301 characters and 84 crocodyliforms supports the position of Simosuchus as sister to Libycosuchus, more derived than Araripesuchus and Uruguaysuchus, and basal to other ziphosuchian notosuchians. We did not recover an Anatosuchus + Simosuchus clade, nor a monophyletic Araripesuchus, but did recover a peirosaurid clade including Peirosaurus, Lomasuchus, Uberabasuchus, and Hamadasuchus (but not Stolokrosaurus). We also found weak support for inclusion of Anatosuchus, Araripesuchus, Peirosauridae, and Mahajangasuchidae within Notosuchia and therefore for the conclusion that all basal mesoeucrocodylians from Gondwana, with the exception of Stolokrosuchus, are notosuchians. The results of this analysis demonstrate that basal mesoeucrocodylian relationships are currently in a state of flux and the need for caution in naming higher-level taxa until a more stable consensus topology is achieved. Similarly, large gaps in the notosuchian record render inconclusive the biogeographic origins of Simosuchus and the clade it represents on Madagascar.


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Project DOI: 10.7934/P581, http://dx.doi.org/10.7934/P581
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 86 Taxa
  • 1 Specimen
  • 301 Characters
Total size of project's media files: 3 MB

Download Project SDD File
Total scored cells: 15747
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 301
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 696
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:301/0
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MorphoBank Project 581

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    Taxonomic Overview for Matrix 'M1433' (85 Taxa)

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    [1] Gracilisuchus
    Last Modified in 12/05/13
    10819319300003
    [2] Terrestrisuchus
    Last Modified in 12/05/13
    15115115000003
    [3] Dibothrosuchus
    Last Modified in 12/05/13
    7722422400003
    [4] Protosuchus
    Last Modified in 12/05/13
    4725925400003
    [5] Hemiprotosuchus
    Last Modified in 12/05/13
    17512612600003
    [6] Orthosuchus
    Last Modified in 12/05/13
    8022222100003
    [7] Kayenta form
    Last Modified in 12/05/13
    13716516400003
    [8] Edentosuchus
    Last Modified in 12/05/13
    18012712100003
    [9] Zaraasuchus
    Last Modified in 12/05/13
    224807700003
    [10] Gobiosuchus
    Last Modified in 12/05/13
    8222421900003
    [11] Sichuanosuchus
    Last Modified in 12/05/13
    13117417000003
    [12] Shantungosuchus
    Last Modified in 12/05/13
    19510810600003
    [13] Zosuchus
    Last Modified in 12/05/13
    10619919500003
    [14] Fruita form
    Last Modified in 12/05/13
    11818418300003
    [15] Notosuchus
    Last Modified in 12/05/13
    2428127700003
    [16] Comahuesuchus
    Last Modified in 12/05/13
    15515014600003
    [17] Mariliasuchus
    Last Modified in 12/05/13
    10619919500003
    [18] Uruguaysuchus
    Last Modified in 12/05/13
    15814714300003
    [19] Chimaeresuchus
    Last Modified in 12/05/13
    206969500003
    [20] Malawisuchus
    Last Modified in 12/05/13
    7723022400003
    [21] Candidodon holo
    Last Modified in 12/05/13
    2956600003
    [22] Simosuchus clarki
    Last Modified in 12/05/13
    1529228501003
    [23] Yacarerani boliviensis
    Last Modified in 12/05/13
    9420920700003
    [24] Sphagesaurus hueni
    Last Modified in 12/05/13
    15214914900003
    [25] Sphagesaurus montealtensis
    Last Modified in 12/05/13
    10519819600003
    [26] Adamantinasuchus
    Last Modified in 12/05/13
    208939300003
    [27] Armadillosuchus arrudai
    Last Modified in 12/05/13
    19510710600003
    [28] Bretesuchus
    Last Modified in 12/05/13
    19611010500003
    [29] Baurusuchus
    Last Modified in 12/05/13
    8821521300003
    [30] Iberosuchus
    Last Modified in 12/05/13
    11219718900003
    [31] Libycosuchus
    Last Modified in 12/05/13
    12418317700003
    [32] Anatosuchus minor
    Last Modified in 12/05/13
    4525925600003
    [33] Araripesuchus gomesii
    Last Modified in 12/05/13
    1729128400003
    [34] Araripesuchus patagonicus
    Last Modified in 12/05/13
    7822822300003
    [35] Araripesuchus buitreraensi
    Last Modified in 12/05/13
    18412511700003
    [36] Araripesuchus wegeneri
    Last Modified in 12/05/13
    2827527201003
    [37] Araripesuchus tsangatsanga
    Last Modified in 12/05/13
    3726826301003
    [38] Lomasuchus
    Last Modified in 12/05/13
    11718618400003
    [39] Peirosaurus
    Last Modified in 12/05/13
    19710710400003
    [40] Hamadasuchus rebouli
    Last Modified in 12/05/13
    7822322300003
    [41] Uberabasuchus terrificus
    Last Modified in 12/05/13
    14216315900003
    [42] Montealtosuchus
    Last Modified in 12/05/13
    7322822800003
    [43] Theriosuchus
    Last Modified in 12/05/13
    7123523000003
    [44] Alligatorium
    Last Modified in 12/05/13
    231707000003
    [45] Goniopholis simus
    Last Modified in 12/05/13
    6723723400003
    [46] Goniopholis stovalli
    Last Modified in 12/05/13
    12417717700003
    [47] Eutretanonew
    Last Modified in 12/05/13
    11618718500003
    [48] Calsoyasuchus
    Last Modified in 12/05/13
    15115115000003
    [49] Sunosuchus
    Last Modified in 12/05/13
    18511611600003
    [50] Shamosuchus
    Last Modified in 12/05/13
    7223722900003
    [51] Bernissartia
    Last Modified in 12/05/13
    11818318300003
    [52] Hylaeochampsa
    Last Modified in 12/05/13
    231707000003
    [53] Allodaposuchus
    Last Modified in 12/05/13
    17712712400003
    [54] Glen rose form
    Last Modified in 12/05/13
    11218918900003
    [55] Borealosuchus
    Last Modified in 12/05/13
    4425825700003
    [56] Pristichampsus
    Last Modified in 12/05/13
    7422922700003
    [57] Eothoracosaurus
    Last Modified in 12/05/13
    9820920300003
    [58] Gavialis
    Last Modified in 12/05/13
    2328127800003
    [59] † Leidyosuchus canadensis
    Last Modified in 12/05/13
    6823523300003
    [60] Asiatosuchus germanicus
    Last Modified in 12/05/13
    8821421300003
    [61] Crocodylus
    Last Modified in 12/05/13
    1828428300003
    [62] Diplocynodon hantoniensis
    Last Modified in 12/05/13
    4126126000003
    [63] Alligator
    Last Modified in 12/05/13
    1728528400003
    [64] Hsisosuchus
    Last Modified in 12/05/13
    11219418900003
    [65] Pelagosaurus
    Last Modified in 12/05/13
    5924724200003
    [66] Steneosaurus
    Last Modified in 12/05/13
    4626425500003
    [67] Metriorhynchus superciliosus
    Last Modified in 12/05/13
    8022322100003
    [68] Metriorhynchus casamequel
    Last Modified in 12/05/13
    18711411400003
    [69] Geosaurus arauc
    Last Modified in 12/05/13
    10420019700003
    [70] Geosaurus suevi
    Last Modified in 12/05/13
    18312011800003
    [71] Dakosaurus maximus
    Last Modified in 12/05/13
    223787800003
    [72] † Dakosaurus andiniensis
    Last Modified in 12/05/13
    14216115900003
    [73] Rhabdognathus m
    Last Modified in 12/05/13
    12917517200003
    [74] Sokotosuchus
    Last Modified in 12/05/13
    232696900003
    [75] Dyrosaurus
    Last Modified in 12/05/13
    12317817800003
    [76] Hyposaurus
    Last Modified in 12/05/13
    19210910900003
    [77] Pholidosaurus
    Last Modified in 12/05/13
    16313913800003
    [78] Sarcosuchus
    Last Modified in 12/05/13
    6524323600003
    [79] Terminonaris robusta
    Last Modified in 12/05/13
    9221520900003
    [80] Pachycheilosuchus
    Last Modified in 12/05/13
    210949100003
    [81] Rugosuchus nonganensis
    Last Modified in 12/05/13
    12917617200003
    [82] Mahajangasuchus
    Last Modified in 12/05/13
    4026326100003
    [83] Kaprosuchus saharicus
    Last Modified in 12/05/13
    7922322200003
    [84] Stolokrosuchus
    Last Modified in 12/05/13
    8821521300003
    [85] Argochampsa
    Last Modified in 12/05/13
    14515815600003


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    Total downloads from project215
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