Project 610: I. M. Vea, D. A. Grimaldi. 2012. Phylogeny of ensign scale insects (Hemiptera: Coccoidea: Ortheziidae) based on the morphology of Recent and fossil females. Systematic Entomology. 37 (4):758-783.
Specimen: † Burmorthezia kotejai (AMNH Bu1094)

Abstract

The Ortheziidae (ensign scale insects) is a morphologically well-defined
family. The morphology and occurrence in the fossil record suggests a probable early origin of the family in scale insect evolution. The present phylogenetic analysis – based on 69 morphological characters of female ortheziids, using 39 exemplar Recent species – provides the first analytical assessment of relationships among living genera of the family, as well as the relationships of eight fossil species, based on complete, well-preserved specimens in amber. Monophyly of the subfamilies
Newsteadiinae, Ortheziinae and Ortheziolinae is supported, but Nipponortheziinae is found to be paraphyletic by inclusion of the Ortheziolinae. Thus, the subfamily Ortheziolinae is reduced in rank to tribe Ortheziolini stat.n., which now includes Matileortheziola Kozár & Foldi, Ortheziolacoccus Kozár, Ortheziolamameti Kozár and Ortheziola Sulc. Consequently, the tribes Matileortheziolini, Ortheziolacoccini and Ortheziolamametini are synonymized (syn.n.) here under Ortheziolini. Five new species and one new genus of fossil ensign scales are described from three amber deposits: Burmorthezia gen.n. with type species Burmorthezia kotejai sp.n. and also B. insolita sp.n., both in mid-Cretaceous Burmese amber (98 Ma) and Arctorthezia baltica sp.n. in Eocene Baltic amber (c. 43 Ma) based on second-instar nymphs; Mixorthezia kozari sp.n. and M. dominicana sp.n. in Miocene Dominican
amber (c. 17 Ma) based on adult females. Fossil placements are unambiguous, with Burmorthezia forming a stem to crown-group (Recent and Tertiary) Ortheziidae.
A summary of described fossil ortheziids is provided.


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Project DOI: 10.7934/P610, http://dx.doi.org/10.7934/P610
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 52 Taxa
  • 6 Specimens
  • 75 Characters
Total size of project's media files: 1 MB

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Total scored cells: 2969
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 69
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 172
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:69/0
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MorphoBank Project 610


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    member name taxa specimens media chars character
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    Isabelle Vea
    Project Administrator
    Last logged in 09/13/12
    526169069002969
    (2920, 0, 49)
    00000


    Taxonomic Overview for Matrix 'M1475' (47 Taxa)

    taxon unscored
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    [1] Matsucoccus gallicolus
    Last Modified in 12/05/13
    2585809001
    [2] Pseudococcus longispinus
    Last Modified in 12/05/13
    0616108001
    [3] Icerya purchasi
    Last Modified in 12/05/13
    0626207001
    [4] Puto yuccae
    Last Modified in 12/05/13
    0636306001
    [5] Newsteadia floccosa
    Last Modified in 12/05/13
    0696900001
    [6] Newsteadia morrisoni
    Last Modified in 12/05/13
    0696900001
    [7] Newsteadia monikae
    Last Modified in 12/05/13
    0696900001
    [8] Newsteadia caledoniensis
    Last Modified in 12/05/13
    1686800001
    [9] Newsteadia minima
    Last Modified in 12/05/13
    0696900001
    [10] Newsteadia americana
    Last Modified in 12/05/13
    0696900001
    [11] Newsteadia africana
    Last Modified in 12/05/13
    1686800001
    [12] Jermycoccus boliviensis
    Last Modified in 12/05/13
    0676702001
    [13] Neomixorthezia brazilana
    Last Modified in 12/05/13
    0696900001
    [14] Neomixorthezia machupicchui
    Last Modified in 12/05/13
    0696900001
    [15] Neonipponorthezia regina
    Last Modified in 12/05/13
    9606000001
    [16] Nipponorthezia ardisiae
    Last Modified in 12/05/13
    0696900001
    [17] Nipponorthezia obscura
    Last Modified in 12/05/13
    0696900001
    [18] † Nipponorthezinella
    Last Modified in 12/05/13
    30393900001
    [19] Nipponorthezinella guadalcanalia
    Last Modified in 12/05/13
    8616100001
    [20] Arctorthezia occidentalis
    Last Modified in 12/05/13
    0696900001
    [21] Arctorthezia pseudoccidentalis
    Last Modified in 12/05/13
    0696900001
    [22] Arctorthezia cataphracta
    Last Modified in 12/05/13
    0696900001
    [23] Graminorthezia graminis
    Last Modified in 12/05/13
    0696900001
    [24] Graminorthezia tillandsiae
    Last Modified in 12/05/13
    8616100001
    [25] Graminorthezia minor
    Last Modified in 12/05/13
    13565600001
    [26] Orthezia urticae
    Last Modified in 12/05/13
    0696900001
    [27] Orthezia ambrosicola
    Last Modified in 12/05/13
    0696900001
    [28] Orthezia annae
    Last Modified in 12/05/13
    0696900001
    [29] Orthezia graminicola
    Last Modified in 12/05/13
    0696900001
    [30] Orthezia newcomeri
    Last Modified in 12/05/13
    0696900001
    [31] Praelongorthezia artemisiae
    Last Modified in 12/05/13
    3666600001
    [32] Praelongorthezia praelonga
    Last Modified in 12/05/13
    0696900001
    [33] Matilorthezia angolaensis
    Last Modified in 12/05/13
    1676701001
    [34] Insignorthezia insignis
    Last Modified in 12/05/13
    0686801001
    [35] Ortheziolacoccus madecassa
    Last Modified in 12/05/13
    6636300001
    [36] Ortheziolamameti guineensis
    Last Modified in 12/05/13
    3666600001
    [37] Ortheziola vejdovskyi
    Last Modified in 12/05/13
    1686800001
    [38] Mixorthexia reynei
    Last Modified in 12/05/13
    9606000001
    [39] Mixorthezia monticola
    Last Modified in 12/05/13
    4656500001
    [40] Acropygorthezia williamsi
    Last Modified in 12/05/13
    15757011001
    [41] † Mixorthezia dominicana
    Last Modified in 12/05/13
    26434300001
    [42] † Mixorthezia kozari2
    Last Modified in 12/05/13
    25444400001
    [43] † Newsteadia succini
    Last Modified in 12/05/13
    25444400001
    [44] † Arctorthezia antiqua
    Last Modified in 12/05/13
    24454500001
    [45] † Arctorthezia baltica
    Last Modified in 12/05/13
    25444400001
    [46] † Burmorthezia kotejai
    Last Modified in 12/05/13
    24434302001
    [47] † Burmorthezia insolita
    Last Modified in 12/05/13
    25424202001


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