Project 638: L. Prendini. 2000. Phylogeny and Classification of the Superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): An Exemplar Approach. Cladistics. 16 (1):1-78.
Specimen: Pandinus imperator (unvouchered)
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Abstract

S. A. Stockwell (1989, “Revision of the Phylogeny and Higher Classification of Scorpions (Chelicerata).” Univ. of California, Berkeley) proposed a cladogram and revised classification for the superfamily Scorpionoidea Latreille 1802 (comprising the families Bothriuridae, Diplocentridae, Heteroscorpionidae, Ischnuridae, and Scorpionidae), based on 47 morphological characters and 35 supraspecific terminal taxa, representing genera whose monophyly was implicitly assumed. Given the widespread practice of defining scorpion genera on the basis of plesiomorphic character states, the assumption of monophyly implicit in supraspecific terminal taxa reduces confidence in Stockwell's cladistic findings and, consequently, his revised suprageneric classification. A re-investigation of scorpionoid phylogeny is presented here, based on 115 morphological characters (including the characters used by Stockwell) and 71 exemplar species. The criterion of “maximal morphological diversity” was employed for exemplar selection. This approach provides a stronger test of monophyly than random exemplar selection. Sixteen cladistic analyses were performed on the scorpionoid data matrix, which varied in the use of equal, successive, or implied weights and in the additive or nonadditive treatment of multistate characters. The preferred hypothesis, a single most parsimonious tree obtained by analysis with equal weights and 13 ordered multistate characters, yielded the scheme of relationships: (Bothriuridae ((Heteroscorpionidae Urodacinae) ((Hemiscorpiinae Ischnuridae) (Diplocentridae Scorpioninae)))). On the basis of these results, revisions are proposed to the existing suprageneric classification of the Scorpionoidea, including new diagnoses, new descriptions, and an illustrated key to the families and subfamilies. Familial status is provided for the scorpionid subfamilies Hemiscorpiinae and Urodacinae.


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Project DOI: 10.7934/P638, http://dx.doi.org/10.7934/P638
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 71 Taxa
  • 1 Specimen
  • 115 Characters
Total size of project's media files: 1 MB

Download Project SDD File
Total scored cells: 8132
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 115
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 266
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:115/0
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MorphoBank Project 638


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    MorphoBank Curator
    Project Administrator
    Last logged in 03/02/12
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    Maureen Admin
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    Last logged in 01/20/12
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    Lorenzo Prendini
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    Taxonomic Overview for Matrix 'M1512' (71 Taxa)

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    [1] Centruroides gracilis
    Last Modified in 12/05/13
    09898017003
    [2] Chaerilus granosus
    Last Modified in 12/05/13
    0102102013003
    [3] Bothriurus bonariensis
    Last Modified in 12/05/13
    011311302003
    [4] Bothriurus coriaceus
    Last Modified in 12/05/13
    011311302003
    [5] Centromachetes obscurus
    Last Modified in 12/05/13
    011311302003
    [6] Centromachetes pocockii
    Last Modified in 12/05/13
    011311302003
    [7] Cercophonius queenslandae
    Last Modified in 12/05/13
    011211203003
    [8] Cercophonius squama
    Last Modified in 12/05/13
    011211203003
    [9] Lisposoma elegans
    Last Modified in 12/05/13
    011111104003
    [10] Lisposoma josehermana
    Last Modified in 12/05/13
    011111104003
    [11] Orobothriurus alticola
    Last Modified in 12/05/13
    011311302003
    [12] Orobothriurus crassimanus
    Last Modified in 12/05/13
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    [13] Phoniocercus pictus
    Last Modified in 12/05/13
    011111104003
    [14] Phoniocercus sanmartini
    Last Modified in 12/05/13
    011111104003
    [15] Tehuankea moyanoi
    Last Modified in 12/05/13
    310910903003
    [16] Thestylus glasioui
    Last Modified in 12/05/13
    011111104003
    [17] Timogenes dorbignyi
    Last Modified in 12/05/13
    011011005003
    [18] Timogenes mapuche
    Last Modified in 12/05/13
    011011005003
    [19] Urophonius iheringii
    Last Modified in 12/05/13
    011211203003
    [20] Urophonius granulatus
    Last Modified in 12/05/13
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    [21] Brachistosternus ehrenbergii
    Last Modified in 12/05/13
    011011005003
    [22] Brachistosternus alienus
    Last Modified in 12/05/13
    011011005003
    [23] Brachistosternus ferrugineus
    Last Modified in 12/05/13
    011011005003
    [24] Vachonia martinezi
    Last Modified in 12/05/13
    110910905003
    [25] Bioculus caboensis
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    [26] Bioculus comondae
    Last Modified in 12/05/13
    011011005003
    [27] Cazierius gundlachii
    Last Modified in 12/05/13
    011011005003
    [28] Cazierius scaber
    Last Modified in 12/05/13
    011011005003
    [29] Didymocentrus hasethi
    Last Modified in 12/05/13
    011011005003
    [30] Didymocentrus lesueurii
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    011011005003
    [31] Diplocentrus gertschi
    Last Modified in 12/05/13
    011011005003
    [32] Diplocentrus mexicanus
    Last Modified in 12/05/13
    011011005003
    [33] Heternebo granti
    Last Modified in 12/05/13
    011011005003
    [34] Heteronebo jamaicae
    Last Modified in 12/05/13
    011011005003
    [35] Oiclus purvesii
    Last Modified in 12/05/13
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    [36] Tarsoporosus flavus
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    [37] Tarsoporosus kugleri
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    [38] Nebo hierichonticus
    Last Modified in 12/05/13
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    [39] Nebo poggesii
    Last Modified in 12/05/13
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    [40] Heteroscorpion goodmani
    Last Modified in 12/05/13
    011111104003
    [41] Heteroscorpion opisthacanthoides
    Last Modified in 12/05/13
    011111104003
    [42] Cheloctonus crassimanus
    Last Modified in 12/05/13
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    [43] Cheloctonus jonesii
    Last Modified in 12/05/13
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    [44] Chiromachetes fergusoni
    Last Modified in 12/05/13
    14999902003
    [45] Chiromachetes tirupati
    Last Modified in 12/05/13
    15989802003
    [46] Chiromachus ochropus
    Last Modified in 12/05/13
    011211203003
    [47] Hadogenes tityrus
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    [48] Hadogenes troglodytes
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    [49] Iomachus laeviceps
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    111111103003
    [50] Iomachus politus
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    [51] Liocheles australasiae
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    011211203003
    [52] Liocheles waigiensis
    Last Modified in 12/05/13
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    [53] Opisthacanthus elatus
    Last Modified in 12/05/13
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    [54] Opisthacanthus madagascariensis
    Last Modified in 12/05/13
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    [55] Opisthacanthus validus
    Last Modified in 12/05/13
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    [56] Palaeocheloctonus pauliani
    Last Modified in 12/05/13
    011211203003
    [57] Habibiella gaillardi
    Last Modified in 12/05/13
    011211203003
    [58] Hemiscorpius lepturus
    Last Modified in 12/05/13
    011211203003
    [59] Hemiscorpius maindroni
    Last Modified in 12/05/13
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    [60] Heterometrus spinifer
    Last Modified in 12/05/13
    011411401003
    [61] Heterometrus swammerdami
    Last Modified in 12/05/13
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    [62] Opistophthalmus boehmi
    Last Modified in 12/05/13
    011311302003
    [63] Opistophthalmus capensis
    Last Modified in 12/05/13
    011311302003
    [64] Opistophthalmus holmi
    Last Modified in 12/05/13
    011111104003
    [65] Pandinus cavimanus
    Last Modified in 12/05/13
    011311302003
    [66] Pandinus dictator
    Last Modified in 12/05/13
    011411401003
    [67] Pandinus imperator
    Last Modified in 12/05/13
    011311302003
    [68] Scorpio maurus kruglovi
    Last Modified in 12/05/13
    011411401003
    [69] Scorpio maurus mogadorensis
    Last Modified in 12/05/13
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    [70] Urodacus novaehollandiae
    Last Modified in 12/05/13
    011111104003
    [71] Urodacus yaschenkoi
    Last Modified in 12/05/13
    011111104003


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