Project 709: J. R. Paterson, D. C. García-Bellido, G. D. Edgecombe. 2012. New Artiopodan Arthropods from the Early Cambrian Emu Bay Shale Konservat-Lagerstätte of South Australia. Journal of Paleontology. 86 (2):340-357.
Specimen: † Squamacula buckorum (SAMP:15347a)

Abstract

The Emu Bay Shale Konservat-Lagerstätte (Cambrian Series 2, Stage 4) on Kangaroo Island, South Australia, is the source of two new non-biomineralized artiopodan arthropods. Squamacula buckorum n. sp. is the first record outside of China of a genus otherwise known only from its type species, S. clypeata, from the Chengjiang biota. The Australian species displays the long cephalic doublure and spiniform exopod setae that are apomorphic for this genus, provides new information on the alimentary tract and midgut glands (the latter preserved as three-dimensional, permineralized structures), and indicates interspecific variability in trunk segment numbers. The distribution of Squamacula strengthens the biogeographic connections between early Cambrian “Burgess Shale-type” biotas of Australia and South China. Australimicola spriggi n. gen. n. sp. represents a monotypic genus resolved in a cladistic analysis of Cambro-Ordovician artiopodans as most closely related to or within Conciliterga (a clade containing Helmetia, Kuamaia, Kwanyinaspis, Rhombicalvaria, Saperion, Skioldia, and Tegopelte). Compared with other members of this clade from Chengjiang and the Burgess Shale, the new genus is diagnosed by an elongate trunk with 23 thoracic tergites having spatulate pleural tips and a small pygidium possessing a single, elongate pair of pleural spines, with specimens also showing a hypostome attached to an anterior (or prehypostomal) sclerite, antennae, short endopods, an annulated alimentary tract, and a series of three-dimensional, permineralized midgut glands. An alternative relationship between Australimicola and the Early Ordovician–Early Devonian Cheloniellida explains the shared anterior flexure of trunk pleurae but forces dubious homologies in other characters, such as dorsally-articulated furcae versus spines.


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Project DOI: 10.7934/P709, http://dx.doi.org/10.7934/P709
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 1 Document
  • 31 Taxa
  • 1 Specimen
  • 38 Characters
Total size of project's media files: 4 MB

Download Project SDD File
Total scored cells: 950
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 38
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 86
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:38/0
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MorphoBank Project 709


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    MorphoBank Curator
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    Last logged in 04/23/12
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    Maureen Admin
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    John Paterson
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    Taxonomic Overview for Matrix 'M1626' (31 Taxa)

    taxon unscored
    cells
    scored
    cells
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    "-" cellscell images labels on
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    [1] † Aglaspis
    Last Modified in 12/05/13
    7282704003
    [2] † Emeraldella
    Last Modified in 12/05/13
    0333305003
    [3] † Sidneyia
    Last Modified in 12/05/13
    1313106003
    [4] † Australimicola
    Last Modified in 12/05/13
    11272601003
    [5] † Buenaspis
    Last Modified in 12/05/13
    13252401003
    [6] † Cindarella
    Last Modified in 12/05/13
    0363602003
    [7] † Emucaris
    Last Modified in 12/05/13
    10262602003
    [8] † Eoredlichia
    Last Modified in 12/05/13
    0373701003
    [9] † Helmetia
    Last Modified in 12/05/13
    6313101003
    [10] † Kangacaris
    Last Modified in 12/05/13
    9272702003
    [11] † Kuamaia
    Last Modified in 12/05/13
    3343401003
    [12] † Kwanyinaspis
    Last Modified in 12/05/13
    7303001003
    [13] † Liwia
    Last Modified in 12/05/13
    19181801003
    [14] † Misszhouia
    Last Modified in 12/05/13
    4303004003
    [15] † Naraoia compacta
    Last Modified in 12/05/13
    5302904003
    [16] † Naraoia spinosa
    Last Modified in 12/05/13
    4303004003
    [17] † Olenoides
    Last Modified in 12/05/13
    1363601003
    [18] † Panlongia
    Last Modified in 12/05/13
    11282601003
    [19] † Phytophilaspis
    Last Modified in 12/05/13
    15212102003
    [20] † Pygmaclypeatus
    Last Modified in 12/05/13
    10272701003
    [21] † Retifacies
    Last Modified in 12/05/13
    1353502003
    [22] † Saperion
    Last Modified in 12/05/13
    7262605003
    [23] † Sinoburius
    Last Modified in 12/05/13
    8292901003
    [24] † Siriocaris
    Last Modified in 12/05/13
    11252403003
    [25] † Skioldia
    Last Modified in 12/05/13
    14191905003
    [26] † Soomaspis
    Last Modified in 12/05/13
    14242301003
    [27] † Squamacula clypeata
    Last Modified in 12/05/13
    1333304003
    [28] † Squamacula buckorum
    Last Modified in 12/05/13
    17212100003
    [29] † Tariccoia
    Last Modified in 12/05/13
    14242301003
    [30] † Tegopelte
    Last Modified in 12/05/13
    14191905003
    [31] † Xandarella
    Last Modified in 12/05/13
    0363602003


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