Project 784: A. Zieritz, A. F. Sartori, A. E. Bogan, D. C. Aldridge. 2014. Reconstructing the evolution of umbonal sculptures in the Unionida. Journal of Zoological Systematics and Evolutionary Research. 53 (1):76-86.
Abstract
‘Umbonal sculptures’ of freshwater mussels (Unionida), which ornament the early ontogenetic shell, have long been used for species identification. Specificity of these sculptures to higher taxonomic levels and their value for phylogenetic reconstruction are still under considerable scientific debate. In particular, the distribution of beak sculpture morphotypes across the unionoid phylogeny and, consequently, evolution of this character remain poorly understood. Based on an examination of 187 taxa, covering five of the six extant unionoid families, this study presents a new model of charac- ter evolution of umbonal sculptures in the order. Ten morphotypes were recognized and conceptualized into the cladistic characters sculpture presence and category. Optimization of sculpture presence on two recent hypotheses of palaeoheterodont phylogenetic relationships using the program Mesquite indicates a sculptured common ancestor of the extant Unionida, with multiple losses of the umbonal ornament occurring subsequently within the clade. Reconstruction of changes in sculpture category is ambiguous and demonstrates the need for further research into the evolutionary relationships of freshwater mussels in general and of their early ontogenetic sculptures in particular. Ambiguity is reduced in analyses applying a model with unequal costs of transformation between character states, which was derived from observations on intermediate forms and polymorphisms. These analyses sug- gest ‘V-shaped’ or ‘nodulous’ sculpture as the plesiomorphic sculptural category for Unionida. The relatively low levels of homoplasy inferred for V-shaped, pseudo-radial and double-looped sculptures suggest that these types may comprise useful guides to relationships within Unionida. The high degree of homoplasy of W-shaped, pseudo-concentric, wrinkled and single-looped sculptures, on the other hand, renders these sculpture types less fit for such purposes.Read the article »
Article DOI: 10.1111/jzs.12077
Project DOI: 10.7934/P784, http://dx.doi.org/10.7934/P784
This project contains | Matrices |
---|---|
Download Project SDD File | Total scored cells: 485 Total media associated with cells: 132 Total labels associated with cell media: 0 |
Characters | |
Total characters: 2 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 11 Total character states with associated media: 10 Total character states with media with labels:0 Total unordered/ordered characters:2/0 |
Currently Viewing:
MorphoBank Project 784
MorphoBank Project 784
- Creation Date:
30 August 2012 - Publication Date:
20 July 2014 - Project views: 167329
- Media downloads: 21
- Matrix downloads: 4
This research
supported by
Authors' Institutions
- North Carolina State University
- University of Cambridge
- Field Museum of Natural History
Members
member name | taxa | specimens | media | media notes | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Andre Sartori Project Administrator | 228 | 77 | 76 | 1 | 2 | 10 | 0 | 485 (464, 0, 21) | 132 | 0 | 0 | ||||
MorphoBank Curator Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Alexandra Zieritz Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M1879' (188 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Neotrigonia margaritacea Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[2] Acuticosta chinensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[3] Cuneopsis celtiformis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[4] Cuneopsis pisciculus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[5] Nodularia douglasiae Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
[6] Nodularia schoedei Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[7] Rhombuniopsis sp. Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[8] Unio caffer Last Modified in 03/20/14 | 0 | 3 | 1 | 0 | 0 | 1 | 0 | 3 |
[9] Unio delphinus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[10] Unio gibbus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[11] Unio mancus Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
[12] Unio pictorum Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[13] Unio ravoisieri Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[14] Unio terminalis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[15] Unio tigridis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[16] Unio tumidiformis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[17] Unio tumidus Last Modified in 03/20/14 | 0 | 5 | 0 | 0 | 0 | 2 | 0 | 3 |
[18] Alasmidonta arcula Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[19] Alasmidonta heterodon Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[20] Alasmidonta marginata Last Modified in 03/19/14 | 0 | 3 | 1 | 0 | 0 | 1 | 0 | 3 |
[21] Alasmidonta undulata Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[22] Alasmidonta varicosa Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[23] Alasmidonta viridis Last Modified in 03/20/14 | 0 | 4 | 1 | 0 | 0 | 1 | 0 | 3 |
[24] Anodonta anatina Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 4 | 0 | 3 |
[25] Anodonta cygnea Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[26] Anodonta kennerlyi Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[27] Anodontoides ferussacianus Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[28] Arcidens confragosus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[29] Cristaria plicata Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[30] Lasmigona complanata Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[31] Lasmigona compressa Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[32] Lasmigona costata Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[33] Lasmigona decorata Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[34] Lasmigona holstonia Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[35] Lasmigona subviridis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[36] Pegias fabula Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[37] Pseudanodonta complanata Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[38] Pyganodon cataracta Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
[39] Pyganodon implicata Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[40] Pyganodon grandis Last Modified in 03/20/14 | 0 | 4 | 1 | 0 | 0 | 1 | 0 | 3 |
[41] Pyganodon lacustris Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[42] Simpsonaias ambigua Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[43] Sinanodonta woodiana Last Modified in 03/20/14 | 0 | 3 | 1 | 0 | 0 | 2 | 0 | 3 |
[44] Strophitus undulatus Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[45] Utterbackia imbecillis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[46] Megalonaias nervosa Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[47] Quadrula cylindrica Last Modified in 03/20/14 | 0 | 5 | 0 | 0 | 0 | 2 | 0 | 3 |
[48] Quadrula pustulosa Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 2 | 0 | 3 |
[49] Quadrula quadrula Last Modified in 03/20/14 | 0 | 4 | 1 | 0 | 0 | 1 | 0 | 3 |
[50] Quadrula verrucosa Last Modified in 03/20/14 | 0 | 5 | 0 | 0 | 0 | 2 | 0 | 3 |
[51] Cyclonaias tuberculata Last Modified in 03/20/14 | 0 | 3 | 1 | 0 | 0 | 1 | 0 | 3 |
[52] Elliptio arctata Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[53] Elliptio complanata Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[54] Elliptio dilatata Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[55] Elliptio jayensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[56] Fusconaia flava Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 2 | 0 | 3 |
[57] Pleurobema sintoxia (=coccineum) Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[58] Uniomerus tetralasmus Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[59] Amblema plicata Last Modified in 03/20/14 | 0 | 3 | 1 | 0 | 0 | 1 | 0 | 3 |
[60] Actinonaias ligamentina Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[61] Actinonaias ligamentina (=carinata) Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[62] Actinonaias pectorosa Last Modified in 03/19/14 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 3 |
[63] Delphinonaias delphinulus Last Modified in 03/20/14 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 3 |
[64] Epioblasma brevidens Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[65] Epioblasma torulosa Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[66] Lampsilis cardium Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[67] Lampsilis fasciola Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 4 | 0 | 3 |
[68] Lampsilis ovata Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[69] Lampsilis radiata Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[70] Lampsilis siliquoidea Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[71] Lampsilis teres Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[72] Ligumia nasuta Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[73] Ligumia recta Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[74] Ligumia subrostrata Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[75] Obliquaria reflexa Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[76] Ptychobranchus fasciolaris Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[77] Toxolasma cylindrellus Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[78] Toxolasma parvus Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[79] Toxolasma paulus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[80] Truncilla donaciformis Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[81] Venustaconcha ellipsiformis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[82] Villosa fabalis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[83] Villosa iris Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 6 | 0 | 3 |
[84] Villosa taeniata Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[85] Gonidea angulata Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[86] Leguminaia saulcyi Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[87] Leguminaia wheatleyi Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[88] Potomida littoralis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[89] Brazzaea sp. Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[90] Coelatura aegyptiaca Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 3 | 0 | 3 |
[91] Coelatura bakeri Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[92] Coelatura hauttecoeuri Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[93] Coelatura horei Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[94] Coelatura hypsiprymna Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[95] Coelatura leopoldvillensis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[96] Coelatura lobensis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[97] Coelatura ratidota Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[98] Coelatura stuhlmanni Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[99] Grandidiera burtoni Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[100] Nitia acuminata Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[101] Nitia teretiuscula Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[102] Nyassunio nyassaensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[103] Nyassunio ujijiensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[104] Pseudospatha tanganyicensis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[105] Caudiculatus caudiculatus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[106] Elongaria orientalis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[107] Harmandia castelneaui Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[108] Harmandia somboriensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[109] Hyriopsis cumingii Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[110] Hyriopsis schlegelii Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[111] Pronodularia japanensis Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
[112] Lamellidens marginalis Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[113] Lamprotula leai Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[114] Parreysia caerulea Last Modified in 03/20/14 | 0 | 5 | 0 | 0 | 0 | 3 | 0 | 3 |
[115] Parreysia corbis Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 2 | 0 | 3 |
[116] Parreysia corrugata Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 3 | 0 | 3 |
[117] Parreysia favidens Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
[118] Parreysia lima Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[119] Parreysia occata Last Modified in 12/05/13 | 0 | 5 | 2 | 0 | 0 | 0 | 0 | 3 |
[120] Parreysia rajahensis Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 2 | 0 | 3 |
[121] Parreysia smaragdites Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[122] Parreysia tavoyensis Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 2 | 0 | 3 |
[123] Parreysia triembola Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[124] Physunio superbus Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
[125] Pilsbryoconcha compressa Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 2 | 0 | 3 |
[126] Pilsbryoconcha exilis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[127] Pressidens exanthematicus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[128] Protunio messageri Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[129] Pseudodon inoscularis Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[130] Pseudodon omiensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[131] Pseudodon vondembuschianus Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[132] Ptychorhynchus murinum Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[133] Scabies crispata Last Modified in 03/20/14 | 0 | 5 | 1 | 0 | 0 | 2 | 0 | 3 |
[134] Scabies nucleus Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
[135] Trapezoideus exolescens Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[136] Uniandra sp. Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[137] Uniandra contradens Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[138] Unionetta sp. Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[139] Cumberlandia monodonta Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[140] Margaritifera margaritifera Last Modified in 03/20/14 | 0 | 4 | 0 | 0 | 0 | 2 | 0 | 3 |
[141] Triplodon chodo Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[142] Triplodon corrugatus Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[143] Triplodon stevensii Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[144] Triplodon sp. Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[145] Castalia ambigua Last Modified in 03/20/14 | 0 | 3 | 1 | 0 | 0 | 1 | 0 | 3 |
[146] Castalia stevensi Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[147] Castalia psammoica Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[148] Castalia sp. Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[149] Diplodon chilensis Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[150] Diplodon deceptus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[151] Diplodon flucki Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[152] Diplodon fluctiger Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[153] Diplodon multistriatus Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[154] Diplodon rhuacoicus Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[155] Echyridella menziesii Last Modified in 03/20/14 | 0 | 3 | 0 | 0 | 0 | 2 | 0 | 3 |
[156] Hyridella australis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[157] Hyridella depressa Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[158] Hyridella drapeta Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[159] Hyridella glenelgensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[160] Hyridella narracanensis Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[161] Virgus beccarianus Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[162] Lortiella froggattii Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[163] Lortiella rugata Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[164] Velesunio ambiguus Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[165] Velesunio angasi Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[166] Mycetopoda siliquosa Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[167] Anodontites elongata Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[168] Anodontites guaranensis Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[169] Anodontites trapesialis Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[170] Anodontites trigonus Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[171] Monocondylaea minuana Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[172] Chelidonopsis hirundo Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[173] Mutela dubia Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[174] Mutela nilotica Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[175] Mutela rostrata Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[176] Pleiodon ovata Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[177] Pleiodon spekii Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[178] Aspatharia chaiziana Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[179] Aspatharia divaricata Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[180] Aspatharia pfeifferiana Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[181] Aspatharia rugifera Last Modified in 12/05/13 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 3 |
[182] Chambardia bourguignati Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[183] Chambardia nyassaensis Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[184] Chambardia wahlbergi Last Modified in 12/05/13 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 3 |
[185] Alathyria pertexta Last Modified in 03/20/14 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 3 |
[186] Lortiella froggatti Last Modified in 03/20/14 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 3 |
[187] Velesunio wilsonii Last Modified in 12/05/13 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 |
[188] Westralunio carteri Last Modified in 12/05/13 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 3 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 167329 | |
Project overview | 6178 | |
Media views | 66364 | Media search (7142 views); M328229 (840 views); M328230 (826 views); M328231 (923 views); M328232 (836 views); M328233 (836 views); M328234 (836 views); M328235 (813 views); M328236 (854 views); M328237 (873 views); M328238 (794 views); M328239 (779 views); M328240 (803 views); M328241 (777 views); M328242 (849 views); M328243 (767 views); M328244 (847 views); M328245 (861 views); M328246 (843 views); M328247 (805 views); M328248 (867 views); M328249 (834 views); M328250 (806 views); M328251 (836 views); M328252 (820 views); M328253 (814 views); M328254 (755 views); M328255 (800 views); M328256 (786 views); M328257 (759 views); M328258 (842 views); M328259 (774 views); M328260 (831 views); M328261 (753 views); M328262 (805 views); M328263 (796 views); M328264 (843 views); M328265 (818 views); M328266 (850 views); M328267 (782 views); M328268 (795 views); M328269 (785 views); M328270 (794 views); M328271 (744 views); M328272 (755 views); M328273 (756 views); M328274 (763 views); M328275 (768 views); M328276 (747 views); M328277 (783 views); M328278 (694 views); M328279 (843 views); M328280 (701 views); M328281 (750 views); M328282 (703 views); M328283 (689 views); M328284 (794 views); M328285 (724 views); M328286 (722 views); M328287 (765 views); M328288 (793 views); M328289 (759 views); M328290 (729 views); M328291 (743 views); M328292 (703 views); M328293 (674 views); M328294 (653 views); M328295 (686 views); M328296 (705 views); M328297 (726 views); M328298 (688 views); M328299 (773 views); M328300 (732 views); M328301 (697 views); M328302 (708 views); M328303 (689 views); M328310 (756 views); |
Matrix views | 3497 | Matrix landing page (2994 views); Umbonal sculpture (503 views); |
Taxon list | 26305 | |
Views for media list | 2219 | |
Specimen list | 62200 | |
Documents list | 34 | |
Bibliography | 532 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 385 | |
Project downloads | 360 | |
Matrix downloads | 4 | Umbonal sculpture (4 downloads); |
Media downloads | 21 | M328281 (3 downloads); M328230 (1 download); M328233 (1 download); M328244 (1 download); M328250 (1 download); M328253 (1 download); M328258 (2 downloads); M328259 (1 download); M328260 (1 download); M328261 (1 download); M328264 (1 download); M328278 (1 download); M328280 (1 download); M328283 (1 download); M328284 (1 download); M328285 (1 download); M328291 (1 download); M328293 (1 download); |