Project 793: R. J. Garwood,J. A. Dunlop,G. Giribet,M. D. Sutton. 2011. Anatomically modern Carboniferous harvestmen demonstrate early cladogenesis and stasis in Opiliones. Nature Communications. 2 (444).
Specimen: † Hastocularis argus (unvouchered)

Abstract

Harvestmen, the third most-diverse arachnid order, are an ancient group found on all continental landmasses, except Antarctica. However, a terrestrial mode of life and leathery, poorly mineralized exoskeleton makes preservation unlikely, and their fossil record is limited. The few Palaeozoic species discovered to date appear surprisingly modern, but are too poorly preserved to allow unequivocal taxonomic placement. Here, we use high-resolution X-ray micro-tomography to describe two new harvestmen from the Carboniferous (~305 Myr) of France. The resulting computer models allow the first phylogenetic analysis of any Palaeozoic Opiliones, explicitly resolving both specimens as members of different extant lineages, and providing corroboration for molecular estimates of an early Palaeozoic radiation within the order. Furthermore, remarkable similarities between these fossils and extant harvestmen implies extensive morphological stasis in the order. Compared with other arachnids—and terrestrial arthropods generally—harvestmen are amongst the first groups to evolve fully modern body plans. This project builds upon MorphoBank Project 631, but includes new fossils.


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Project DOI: 10.7934/P793, http://dx.doi.org/10.7934/P793
This project containsMatrices
  • 3 Media
  • 1 Matrix
  • 2 Documents
  • 51 Taxa
  • 3 Specimens
  • 253 Characters
Total size of project's media files: 2 MB

Download Project SDD File
Total scored cells: 9850
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 253
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 542
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:253/0
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MorphoBank Project 793


    Members

    member name taxa specimens media chars character
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    media
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    (scored, NPA, "-")
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    Gonzalo Giribet
    Project Administrator
    Last logged in 04/12/14
    70100001036
    (610, 327, 99)
    000
    MorphoBank Curator
    Full membership
    Last logged in 04/24/14
    03200000
    (0, 0, 0)
    000
    Jason Dunlop
    Full membership
    Last logged in Never
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    Russell Garwood
    Full membership
    Last logged in 08/15/13
    00000000
    (0, 0, 0)
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    Prashant Sharma
    Full membership
    Last logged in 09/18/12
    00000000
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    Jo Wolfe
    Full membership
    Last logged in 04/11/14
    44002530008814
    (7617, 0, 1197)
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    Taxonomic Overview for Matrix 'M1802' (51 Taxa)

    taxon unscored
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    [1] Limulus polyphemus
    Last Modified in 12/05/13
    42161161050006
    [2] Carcinoscorpius rotundicauda
    Last Modified in 12/05/13
    44159159050006
    [3] Androctonus australis
    Last Modified in 12/05/13
    32176176045006
    [4] Belisarius xambeui
    Last Modified in 12/05/13
    38166166049006
    [5] Siro rubens
    Last Modified in 12/05/13
    33189189130006
    [6] Parasiro coiffaiti
    Last Modified in 12/05/13
    48175175030006
    [7] Chileogovea oedipus
    Last Modified in 12/05/13
    1409898114006
    [8] Ogovea cameroonensis
    Last Modified in 12/05/13
    1419797015006
    [9] Huitaca ventralis
    Last Modified in 12/05/13
    1419696016006
    [10] Meghalaya sp. 19a
    Last Modified in 12/05/13
    129108107214006
    [11] Fangensis insulanus
    Last Modified in 12/05/13
    47178178127006
    [12] Leptopsalis sp. 3 gunung bondeh
    Last Modified in 12/05/13
    48177177127006
    [13] Dalquestia formosa
    Last Modified in 12/05/13
    48186186019006
    [14] Odiellus troguloides
    Last Modified in 12/05/13
    35196196022006
    [15] Phalangium opilio
    Last Modified in 12/05/13
    34197197022006
    [16] Opilio parietinus
    Last Modified in 12/05/13
    47187187019006
    [17] Astrobunus grallator
    Last Modified in 12/05/13
    47187187019006
    [18] Nelima silvatica
    Last Modified in 12/05/13
    47187187019006
    [19] Leiobunum sp.
    Last Modified in 12/05/13
    32199199022006
    [20] Hadrobunus maculosus
    Last Modified in 12/05/13
    47187187019006
    [21] Caddo agilis
    Last Modified in 12/05/13
    48186186019006
    [22] Ischyropsalis luteipes
    Last Modified in 12/05/13
    35191191027006
    [23] Hesperonemastoma modestum
    Last Modified in 12/05/13
    47180180026006
    [24] Ceratolasma tricantha
    Last Modified in 12/05/13
    48181181024006
    [25] Sabacon cavicolens
    Last Modified in 12/05/13
    47182182024006
    [26] Taracus sp.
    Last Modified in 12/05/13
    47182182024006
    [27] Dicranolasma soerenseni
    Last Modified in 12/05/13
    47180180026006
    [28] Centetostoma dubium
    Last Modified in 12/05/13
    50177177026006
    [29] Ortholasma sp.
    Last Modified in 12/05/13
    48180180025006
    [30] Nemastoma bimaculatum
    Last Modified in 12/05/13
    33191191029006
    [31] Nipponopsalis abei
    Last Modified in 12/05/13
    48180180025006
    [32] Trogulus nepaeformis
    Last Modified in 12/05/13
    34190190029006
    [33] † Halitherses grimaldii
    Last Modified in 12/05/13
    12986861820006
    [34] † Ameticos scolos
    Last Modified in 12/05/13
    14558583317006
    [35] † Macrogyion cronus
    Last Modified in 12/05/13
    11564645915006
    [36] † Eophalangium sheari
    Last Modified in 12/05/13
    11651517511006
    [37] † Hastocularis argus
    Last Modified in 12/05/13
    1047271717006
    [38] † Mesobunus dunlopi
    Last Modified in 12/05/13
    1216060657006
    [39] Equitius doriae
    Last Modified in 12/05/13
    56175175022006
    [40] Triaenobunus sp.
    Last Modified in 12/05/13
    58173173022006
    [41] Zuma acuta
    Last Modified in 12/05/13
    58173173022006
    [42] Sandokan cf. alticeps
    Last Modified in 12/05/13
    47184183023006
    [43] Gnomulus sp.
    Last Modified in 12/05/13
    47184183023006
    [44] Scotolemon lespesi
    Last Modified in 12/05/13
    35192192026006
    [45] Maiorerus randoi
    Last Modified in 12/05/13
    38187187028006
    [46] Bishopella laciniosa
    Last Modified in 12/05/13
    45183183025006
    [47] Gnidia holmbergi
    Last Modified in 12/05/13
    43185185025006
    [48] Pachyloides thorellii
    Last Modified in 12/05/13
    34193193026006
    [49] Hoplobunus sp.
    Last Modified in 12/05/13
    45181181027006
    [50] Solifugae
    Last Modified in 12/05/13
    34157157062006
    [51] Pseudoscorpiones
    Last Modified in 12/05/13
    34163163056006


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    Taxon list8884
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    Bibliography636
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    Documents list1260
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    Matrix downloads8Anatomically modern Carboniferous harvestmen (8 downloads);
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