Project 797: Linda A. Tsuji, J. Muller, R. R. Reisz. 2012. Anatomy of Emeroleter levis and the Phylogeny of the Nycteroleter Parareptiles. Journal of Vertebrate Paleontology. 32 (1):45-67.
Specimen: † Emeroleter (:KPM uncat/E1)


The nycteroleter parareptiles have lately become the focus of increased attention, owing to their recently recognized sister-group relationship to pareiasaurs and the discovery an impedance-matching ear in members of the group. The Kotel’nich locality in central Russia dates to the early part of the Late Permian and preserves a diverse tetrapod fauna that includes the pareiasaur Deltavjatia vjatkensis and the nycteroleter Emeroleter levis. The latter taxon was previously known only from isolated crania, but recent excavations have produced additional material, including well-preserved postcrania, allowing a complete redescription. The skull of Emeroleter is typified by dermal sculpturing consisting of evenly spaced small, round pits and supratemporals that extend posterolaterally into broad horns. The quadratojugal is long and the posterior portion of the element curves dorsally into a pointed projection, an autapomorphy of the taxon. Postcranially, Emeroleter is very gracile with a long, sigmoid femur, and slender limbs, carpal, and tarsal elements. A reassessment of the Russian nycteroleters results in the synonymy of Tokosaurus perforatus with Macroleter poezicus. Phylogenetic analysis of parareptilian relationships using both parsimony and Bayesian inference yields a monophyly of ‘nycteroleters’ in parsimony, whereas the group is found to be paraphyletic with Bayesian inference. The genus Bashkyroleter is consistently paraphyletic. A clade consisting of the nycteroleters and pareiasaurs, here termed Pareiasauromorpha, is supported by both methods.

Read the article »

Project DOI: 10.7934/P797,
This project contains
  • 1 Media
  • 3 Documents
  • 30 Taxa
  • 1 Specimen
Total size of project's media files: 2.78M

Download Project SDD File
Currently Viewing:
MorphoBank Project 797

    Authors' Institutions

    • Museum für Naturkunde - Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin

    • University of Toronto


    member name taxa specimens media
    MorphoBank Curator
    Project Administrator
    Maureen Admin
    Full membership
    Linda Tsuji
    Full membership

    Project has no matrices defined.

    Project views

    type number of views Individual items viewed (where applicable)
    Total project views36703
    Project overview3640
    Media views6976Media search (5951 views); M149503 (1025 views);
    Specimen list3381
    Taxon list18743
    Documents list2217
    Matrix views276Matrix landing page (276 views);

    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project337
    Document downloads12Emeroleter Matrix (5 downloads); Copyright Permission (2 downloads); Character List (5 downloads);
    Media downloads1M149503 (1 download);
    Project downloads324