Project 798: M. Murakami, C. Shimada, Y. Hikida, H. Hirano. 2012. Two new extinct basal phocoenids (Cetacea, Odontoceti, Delphinoidea), from the upper Miocene Koetoi Formation of Japan and their phylogenetic significance. Journal of Vertebrate Paleontology. 32 (5):1172-1185.
Abstract
Two new extinct porpoises—Archaeophocaena teshioensis, gen. et sp. nov., and Miophocaena nishinoi, gen. et sp. nov.—are described from the upper Miocene Koetoi Formation (5.5–6.4 Ma) of Hokkaido, Northern Japan. The holotype of the former is composed of a partial skull, whereas the holotype of the latter is composed of a partial skull, right periotic, right stylohyoid, and pelvis. Both species are assigned to Phocoenidae on the basis of a unique combination of phocoenid characters: presences of frontal bosses, nasal protuberances, premaxillary eminences, and fossae for the inferior vestibule. However, they do not have dorsally developed preorbital sinus fossae or high premaxillary eminences, unlike derived phocoenids. Furthermore, their premaxillae contact the nasals (or the right premaxilla alone contacts the right nasal), unlike all other known phocoenids except Pterophocaena. A comprehensive morphological cladistic analysis indicates that the two new extinct species are the second-most basal phocoenids next to Pterophocaena and that Phocoenidae is the sister group of Delphinidae. The cranial morphology of the two new extinct species is intermediate in form between that of phocoenids and delphinids, supporting the hypothesis of a sister relationship for the two groups. Thus, these new species fill not only the morphological gap between Phocoenidae and other families of Delphinoidea but also a temporal gap in the fossil record of phocoenids in the western North Pacific.Read the article »
Article DOI: 10.1080/02724634.2012.694337
Project DOI: 10.7934/P798, http://dx.doi.org/10.7934/P798
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 15797 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 282 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1414 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:282/0 |
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MorphoBank Project 798
MorphoBank Project 798
- Creation Date:
28 September 2012 - Publication Date:
23 March 2022 - Project views: 8764
- Matrix downloads: 2
Authors' Institutions
- Waseda University
- Geological Survey of Japan
- Akita University
- Nakagawa Museum of Natural History
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 2 | 2 | 4 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Maureen Admin Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Mizuki Murakami Full membership | 71 | 0 | 2 | 282 | 0 | 0 | 15797 (15442, 0, 355) | 0 | 0 | 0 | ||||
Mizuki Murakami Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M1872' (73 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Georgiacetus vogtlensis Last Modified in 12/05/13 | 133 | 127 | 127 | 0 | 22 | 0 | 0 | 4 |
[2] † Zygorhiza kochii Last Modified in 12/05/13 | 68 | 195 | 192 | 0 | 22 | 0 | 0 | 4 |
[3] † Archaeodelphis patrius Last Modified in 12/05/13 | 168 | 105 | 105 | 0 | 9 | 0 | 0 | 4 |
[4] † Xenorophus sloanii Last Modified in 12/05/13 | 191 | 88 | 88 | 0 | 3 | 0 | 0 | 4 |
[5] † Agorophius pygmaeus Last Modified in 12/05/13 | 181 | 98 | 97 | 0 | 4 | 0 | 0 | 4 |
[6] † Patriocetus kazakhstanicus Last Modified in 12/05/13 | 164 | 117 | 116 | 0 | 2 | 0 | 0 | 4 |
[7] † Orycterocetus crocodilinus Last Modified in 12/05/13 | 93 | 185 | 180 | 0 | 9 | 0 | 0 | 4 |
[8] Physeter catodon Last Modified in 12/05/13 | 15 | 257 | 253 | 0 | 14 | 0 | 0 | 4 |
[9] Kogia breviceps Last Modified in 12/05/13 | 13 | 248 | 246 | 0 | 23 | 0 | 0 | 4 |
[10] † Waipatia maerewhenua Last Modified in 12/05/13 | 83 | 199 | 195 | 0 | 4 | 0 | 0 | 4 |
[11] † Squalodon calvertensis Last Modified in 12/05/13 | 70 | 212 | 209 | 0 | 3 | 0 | 0 | 4 |
[12] † Notocetus vanbenedeni Last Modified in 12/05/13 | 81 | 202 | 197 | 0 | 4 | 0 | 0 | 4 |
[13] † Prosqualodon davidis Last Modified in 12/05/13 | 104 | 179 | 173 | 0 | 5 | 0 | 0 | 4 |
[14] † Squaloziphius emlongi Last Modified in 12/05/13 | 151 | 128 | 127 | 0 | 4 | 0 | 0 | 4 |
[15] † Zarhachis flagellator Last Modified in 12/05/13 | 74 | 211 | 205 | 0 | 3 | 0 | 0 | 4 |
[16] † Pomatodelphis inaequalis Last Modified in 12/05/13 | 103 | 175 | 175 | 0 | 4 | 0 | 0 | 4 |
[17] Platanista gangetica Last Modified in 12/05/13 | 19 | 264 | 257 | 0 | 6 | 0 | 0 | 4 |
[18] Tasmacetus shepherdi Last Modified in 12/05/13 | 12 | 270 | 267 | 0 | 3 | 0 | 0 | 4 |
[19] Berardius bairdii Last Modified in 12/05/13 | 18 | 262 | 257 | 0 | 7 | 0 | 0 | 4 |
[20] Ziphius cavirostris Last Modified in 12/05/13 | 10 | 272 | 263 | 0 | 9 | 0 | 0 | 4 |
[21] Mesoplodon ginkgodens Last Modified in 12/05/13 | 32 | 245 | 243 | 0 | 7 | 0 | 0 | 4 |
[22] † Xiphiacetus bossi Last Modified in 12/05/13 | 53 | 234 | 226 | 0 | 3 | 0 | 0 | 4 |
[23] Lipotes vexillifer Last Modified in 12/05/13 | 6 | 275 | 275 | 0 | 1 | 0 | 0 | 4 |
[24] † Parapontoporia sternbergi Last Modified in 12/05/13 | 135 | 149 | 145 | 0 | 2 | 0 | 0 | 4 |
[25] Inia geoffrensis Last Modified in 12/05/13 | 5 | 282 | 269 | 0 | 8 | 0 | 0 | 4 |
[26] Pontoporia blainvillei Last Modified in 12/05/13 | 4 | 280 | 275 | 0 | 3 | 0 | 0 | 4 |
[27] † Pliopontos littoralis Last Modified in 12/05/13 | 108 | 177 | 172 | 0 | 2 | 0 | 0 | 4 |
[28] † Brachydelphis mazeasi Last Modified in 12/05/13 | 92 | 197 | 189 | 0 | 1 | 0 | 0 | 4 |
[29] † Kentriodon pernix Last Modified in 12/05/13 | 65 | 221 | 215 | 0 | 2 | 0 | 0 | 4 |
[30] † Atocetus iquensis Last Modified in 12/05/13 | 63 | 220 | 217 | 0 | 2 | 0 | 0 | 4 |
[31] † Hadrodelphis calvertense Last Modified in 12/05/13 | 121 | 159 | 159 | 0 | 2 | 0 | 0 | 4 |
[32] † Albireo whistleri Last Modified in 12/05/13 | 53 | 226 | 225 | 0 | 4 | 0 | 0 | 4 |
[33] † Denebola brachycephala Last Modified in 12/05/13 | 131 | 149 | 149 | 0 | 2 | 0 | 0 | 4 |
[34] Delphinapterus leucas Last Modified in 12/05/13 | 2 | 283 | 275 | 0 | 5 | 0 | 0 | 4 |
[35] Monodon monoceros Last Modified in 12/05/13 | 12 | 270 | 261 | 0 | 9 | 0 | 0 | 4 |
[36] † Odobenocetops peruvianus Last Modified in 12/05/13 | 106 | 156 | 156 | 0 | 20 | 0 | 0 | 4 |
[37] † Australithax intermedia Last Modified in 12/05/13 | 167 | 113 | 113 | 0 | 2 | 0 | 0 | 4 |
[38] † Lomacetus ginsburgi Last Modified in 12/05/13 | 165 | 115 | 115 | 0 | 2 | 0 | 0 | 4 |
[39] † Haborophocoena toyoshimai Last Modified in 12/05/13 | 110 | 170 | 170 | 0 | 2 | 0 | 0 | 4 |
[40] † Piscolithax boreios Last Modified in 12/05/13 | 125 | 154 | 154 | 0 | 3 | 0 | 0 | 4 |
[41] † Piscolithax longirostris Last Modified in 12/05/13 | 80 | 201 | 200 | 0 | 2 | 0 | 0 | 4 |
[42] † Piscolithax tedfordi Last Modified in 12/05/13 | 102 | 178 | 178 | 0 | 2 | 0 | 0 | 4 |
[43] † Salumiphocaena stocktoni Last Modified in 12/05/13 | 148 | 132 | 132 | 0 | 2 | 0 | 0 | 4 |
[44] † Septemtriocetus bosselaersi Last Modified in 12/05/13 | 195 | 84 | 84 | 0 | 3 | 0 | 0 | 4 |
[45] Phocoena sinus Last Modified in 12/05/13 | 16 | 267 | 263 | 0 | 3 | 0 | 0 | 4 |
[46] Phocoena spinipinnis Last Modified in 12/05/13 | 12 | 276 | 267 | 0 | 3 | 0 | 0 | 4 |
[47] Phocoena dioptrica Last Modified in 12/05/13 | 67 | 220 | 212 | 0 | 3 | 0 | 0 | 4 |
[48] Phocoena phocoena Last Modified in 12/05/13 | 1 | 289 | 277 | 0 | 4 | 0 | 0 | 4 |
[49] Phocoenoides dalli Last Modified in 12/05/13 | 5 | 289 | 274 | 0 | 3 | 0 | 0 | 4 |
[50] Neophocaena phocaenoides Last Modified in 12/05/13 | 5 | 284 | 273 | 0 | 4 | 0 | 0 | 4 |
[51] Lissodelphis borealis Last Modified in 12/05/13 | 12 | 276 | 267 | 0 | 3 | 0 | 0 | 4 |
[52] Cephalorhynchus hectori Last Modified in 12/05/13 | 9 | 274 | 270 | 0 | 3 | 0 | 0 | 4 |
[53] Lagenorhynchus acutus Last Modified in 12/05/13 | 19 | 260 | 259 | 0 | 4 | 0 | 0 | 4 |
[54] Orcaella brevirostris Last Modified in 12/05/13 | 20 | 260 | 257 | 0 | 5 | 0 | 0 | 4 |
[55] Orcinus orca Last Modified in 12/05/13 | 16 | 270 | 261 | 0 | 5 | 0 | 0 | 4 |
[56] Feresa attenuata Last Modified in 12/05/13 | 5 | 276 | 272 | 0 | 5 | 0 | 0 | 4 |
[57] Globicephala macrorhynchus Last Modified in 12/05/13 | 14 | 265 | 263 | 0 | 5 | 0 | 0 | 4 |
[58] Grampus griseus Last Modified in 12/05/13 | 10 | 272 | 266 | 0 | 6 | 0 | 0 | 4 |
[59] † Hemisyntrachelus cortesii Last Modified in 12/05/13 | 93 | 186 | 185 | 0 | 4 | 0 | 0 | 4 |
[60] Peponocephala electra Last Modified in 12/05/13 | 10 | 272 | 267 | 0 | 5 | 0 | 0 | 4 |
[61] Pseudorca crassidens Last Modified in 12/05/13 | 13 | 274 | 264 | 0 | 5 | 0 | 0 | 4 |
[62] Delphinus delphis Last Modified in 12/05/13 | 9 | 277 | 269 | 0 | 4 | 0 | 0 | 4 |
[63] Lagenodelphis hosei Last Modified in 12/05/13 | 6 | 279 | 272 | 0 | 4 | 0 | 0 | 4 |
[64] Sousa chinensis Last Modified in 12/05/13 | 23 | 264 | 255 | 0 | 4 | 0 | 0 | 4 |
[65] Stenella attenuata Last Modified in 12/05/13 | 7 | 283 | 271 | 0 | 4 | 0 | 0 | 4 |
[66] Tursiops truncatus Last Modified in 12/05/13 | 1 | 294 | 277 | 0 | 4 | 0 | 0 | 4 |
[67] Sotalia fluviatilis Last Modified in 12/05/13 | 8 | 277 | 270 | 0 | 4 | 0 | 0 | 4 |
[68] Steno bredanensis Last Modified in 12/05/13 | 8 | 280 | 270 | 0 | 4 | 0 | 0 | 4 |
[69] † Pterophocaena nishinoi Last Modified in 12/05/13 | 136 | 146 | 146 | 0 | 0 | 0 | 0 | 4 |
[70] † Stenella kabatensis Last Modified in 12/05/13 | 162 | 120 | 120 | 0 | 0 | 0 | 0 | 4 |
[71] † Miophocaena nishinoi Last Modified in 12/05/13 | 197 | 83 | 83 | 0 | 2 | 0 | 0 | 4 |
[72] † Archaeophocaena teshioensis Last Modified in 12/05/13 | 228 | 52 | 52 | 0 | 2 | 0 | 0 | 4 |
[73] † Haborophocoena minutus Last Modified in 12/05/13 | 168 | 113 | 112 | 0 | 2 | 0 | 0 | 4 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 8764 | |
Documents list | 330 | |
Media views | 2251 | Media search (1080 views); M151145 (180 views); M149653 (215 views); M149651 (194 views); M149652 (196 views); M149650 (198 views); M151146 (188 views); |
Project overview | 538 | |
Matrix views | 365 | Matrix landing page (323 views); Matrix of Murakami et al., 2012b nexus file (42 views); |
Bibliography | 239 | |
Specimen list | 961 | |
Views for media list | 640 | |
Taxon list | 3440 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 13 | |
Project downloads | 11 | |
Matrix downloads | 2 | Matrix of Murakami et al., 2012b nexus file (2 downloads); |