Project 964: S. D. Sampson, E. K. Lund, M. A. Loewen, A. A. Farke, K. E. Clayton. 2013. A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia. Proceedings of the Royal Society B: Biological Sciences. 280 (1766).
Specimen: † Nasutoceratops titusi (UMNH/VP:16800)

Abstract

The fossil record of centrosaurine ceratopsids is largely restricted to the northern region of western North America (Alberta, Montana, Alaska). Exceptions consist of single taxa from Utah (Diabloceratops) and China (Sinoceratops), plus otherwise fragmentary remains from southern Laramidia. Here we describe a remarkable new taxon, Nasutoceratops titusi n. gen. et sp., from the late Campanian Kaiparowits Formation of Utah, represented by multiple specimens, including a nearly complete skull and partial postcranial skeleton. Autapomorphies include an enlarged narial region, pneumatic nasal ornamentation, abbreviated snout, and elongate, rostrolaterally directed supraorbital horncores. The subrectangular parietosquamosal frill is relatively unadorned and broadest in the mid-region. A phylogenetic analysis indicates that Nasutoceratops is the sister taxon to Avaceratops, and that a previously unknown subclade of centrosaurines branched off early in the group’s history and persisted for several million years during the late Campanian. As the first well-represented southern centrosaurine comparable in age to the bulk of northern forms, Nasutoceratops provides strong support for the provincialism hypothesis, which posits that Laramidia—the western landmass formed by inundation of the central region of North America by the Western Interior Seaway—hosted at least two coeval dinosaur communities for over a million years of late Campanian time.


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Project DOI: 10.7934/P964, http://dx.doi.org/10.7934/P964
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 3 Documents
  • 24 Taxa
  • 1 Specimen
  • 98 Characters
Total size of project's media files: 2 MB

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Total scored cells: 1684
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 98
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 396
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:98/0
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MorphoBank Project 964


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    Mark Loewen
    Project Administrator
    Last logged in 09/25/13
    241119800001684
    (1480, 0, 204)
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    Maureen Admin
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    Last logged in 10/02/13
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    MorphoBank Curator
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    Taxonomic Overview for Matrix 'M2151' (24 Taxa)

    taxon unscored
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    [1] † Leptoceratops gracilis
    Last Modified in 12/05/13
    16060037003
    [2] † Protoceratops andrewsi
    Last Modified in 12/05/13
    26666030003
    [3] † Magnirostris dodsoni
    Last Modified in 12/05/13
    333535030003
    [4] † Bagaceratops rozhdestvenskyi
    Last Modified in 12/05/13
    263838034003
    [5] † Turanoceratops tardabilis
    Last Modified in 12/05/13
    80141404003
    [6] † Zuniceratops christopheri
    Last Modified in 12/05/13
    255353020003
    [7] † Diabloceratops eatoni
    Last Modified in 12/05/13
    31666601003
    [8] † Albertaceratops nesmoi
    Last Modified in 12/05/13
    37595902003
    [9] † Rubeosaurus ovatus
    Last Modified in 12/05/13
    62353501003
    [10] † Styracosaurus albertensis
    Last Modified in 12/05/13
    2949402003
    [11] † Spinops sternbergorum
    Last Modified in 12/05/13
    62343402003
    [12] † Centrosaurus apertus
    Last Modified in 12/05/13
    0969602003
    [13] † Coronosaurus brinkmani
    Last Modified in 12/05/13
    13838302003
    [14] † Xenoceratops foremostensis
    Last Modified in 12/05/13
    73242401003
    [15] † Sinoceratops zhuchengensis
    Last Modified in 12/05/13
    47494803003
    [16] † Einiosaurus procurvicornis
    Last Modified in 12/05/13
    17797803003
    [17] † Achelousaurus horneri
    Last Modified in 12/05/13
    26696903003
    [18] † Pachyrhinosaurus canadensis
    Last Modified in 12/05/13
    36595903003
    [19] † Pachyrhinosaurus lakustai
    Last Modified in 12/05/13
    8878703003
    [20] † Pachyrhinosaurus perotorum
    Last Modified in 12/05/13
    43535203003
    [21] † Chasmosaurus belli
    Last Modified in 12/05/13
    1939205003
    [22] † Pentaceratops sternbergii
    Last Modified in 12/05/13
    1929205003
    [23] † Avaceratops lammersi
    Last Modified in 12/05/13
    19747405003
    [24] † Nasutoceratops titusi
    Last Modified in 12/05/13
    27686803003


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    Total project views22772
    Project overview3037
    Matrix views2016Matrix landing page (1789 views); Nasutoceratops phylogeny (227 views);
    Taxon list9302
    Media views4184Media search (3648 views); M307812 (536 views);
    Specimen list1936
    Bibliography736
    Documents list1561




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    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project226
    Project downloads204
    Matrix downloads13Nasutoceratops phylogeny (13 downloads);
    Document downloads8Nasutoceratops characters (4 downloads); Nasutoceratops matrix Excel (2 downloads); Nasutoceratops matrix TNT (2 downloads);
    Media downloads1M307812 (1 download);