Project 964: S. D. Sampson, E. K. Lund, M. A. Loewen, A. A. Farke, K. E. Clayton. 2013. A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia. Proceedings of the Royal Society B: Biological Sciences. 280 (1766).
Abstract
The fossil record of centrosaurine ceratopsids is largely restricted to the northern region of western North America (Alberta, Montana, Alaska). Exceptions consist of single taxa from Utah (Diabloceratops) and China (Sinoceratops), plus otherwise fragmentary remains from southern Laramidia. Here we describe a remarkable new taxon, Nasutoceratops titusi n. gen. et sp., from the late Campanian Kaiparowits Formation of Utah, represented by multiple specimens, including a nearly complete skull and partial postcranial skeleton. Autapomorphies include an enlarged narial region, pneumatic nasal ornamentation, abbreviated snout, and elongate, rostrolaterally directed supraorbital horncores. The subrectangular parietosquamosal frill is relatively unadorned and broadest in the mid-region. A phylogenetic analysis indicates that Nasutoceratops is the sister taxon to Avaceratops, and that a previously unknown subclade of centrosaurines branched off early in the group’s history and persisted for several million years during the late Campanian. As the first well-represented southern centrosaurine comparable in age to the bulk of northern forms, Nasutoceratops provides strong support for the provincialism hypothesis, which posits that Laramidia—the western landmass formed by inundation of the central region of North America by the Western Interior Seaway—hosted at least two coeval dinosaur communities for over a million years of late Campanian time.Read the article »
Project DOI: 10.7934/P964, http://dx.doi.org/10.7934/P964
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 1684 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 98 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 396 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:98/0 |
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MorphoBank Project 964
MorphoBank Project 964
- Creation Date:
03 June 2013 - Publication Date:
10 September 2013 - Media downloads: 1
- Matrix downloads: 17
Authors' Institutions
- Ohio University
- University of Utah
- Denver Museum of Nature and Science
- Natural History Museum of Utah
- Raymond Alf Museum of Paleontology (The Webb Schools)
Members
member name | taxa | specimens | media | media notes | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||||||||
Mark Loewen Project Administrator | 24 | 1 | 1 | 1 | 98 | 0 | 0 | 0 | 0 | 1684 (1480, 0, 204) | 0 | 0 | 0 | 0 | |||||||
Maureen Admin Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | 0 | |||||||
MorphoBank Curator Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M2151' (24 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Leptoceratops gracilis Taxon name last Modified on 12/05/13 | 1 | 60 | 60 | 0 | 37 | 0 | 0 | 3 |
[2] † Protoceratops andrewsi Taxon name last Modified on 12/05/13 | 2 | 66 | 66 | 0 | 30 | 0 | 0 | 3 |
[3] † Magnirostris dodsoni Taxon name last Modified on 12/05/13 | 33 | 35 | 35 | 0 | 30 | 0 | 0 | 3 |
[4] † Bagaceratops rozhdestvenskyi Taxon name last Modified on 12/05/13 | 26 | 38 | 38 | 0 | 34 | 0 | 0 | 3 |
[5] † Turanoceratops tardabilis Taxon name last Modified on 12/05/13 | 80 | 14 | 14 | 0 | 4 | 0 | 0 | 3 |
[6] † Zuniceratops christopheri Taxon name last Modified on 12/05/13 | 25 | 53 | 53 | 0 | 20 | 0 | 0 | 3 |
[7] † Diabloceratops eatoni Taxon name last Modified on 12/05/13 | 31 | 66 | 66 | 0 | 1 | 0 | 0 | 3 |
[8] † Albertaceratops nesmoi Taxon name last Modified on 12/05/13 | 37 | 59 | 59 | 0 | 2 | 0 | 0 | 3 |
[9] † Rubeosaurus ovatus Taxon name last Modified on 12/05/13 | 62 | 35 | 35 | 0 | 1 | 0 | 0 | 3 |
[10] † Styracosaurus albertensis Taxon name last Modified on 12/05/13 | 2 | 94 | 94 | 0 | 2 | 0 | 0 | 3 |
[11] † Spinops sternbergorum Taxon name last Modified on 12/05/13 | 62 | 34 | 34 | 0 | 2 | 0 | 0 | 3 |
[12] † Centrosaurus apertus Taxon name last Modified on 12/05/13 | 0 | 96 | 96 | 0 | 2 | 0 | 0 | 3 |
[13] † Coronosaurus brinkmani Taxon name last Modified on 12/05/13 | 13 | 83 | 83 | 0 | 2 | 0 | 0 | 3 |
[14] † Xenoceratops foremostensis Taxon name last Modified on 12/05/13 | 73 | 24 | 24 | 0 | 1 | 0 | 0 | 3 |
[15] † Sinoceratops zhuchengensis Taxon name last Modified on 12/05/13 | 47 | 49 | 48 | 0 | 3 | 0 | 0 | 3 |
[16] † Einiosaurus procurvicornis Taxon name last Modified on 12/05/13 | 17 | 79 | 78 | 0 | 3 | 0 | 0 | 3 |
[17] † Achelousaurus horneri Taxon name last Modified on 12/05/13 | 26 | 69 | 69 | 0 | 3 | 0 | 0 | 3 |
[18] † Pachyrhinosaurus canadensis Taxon name last Modified on 12/05/13 | 36 | 59 | 59 | 0 | 3 | 0 | 0 | 3 |
[19] † Pachyrhinosaurus lakustai Taxon name last Modified on 12/05/13 | 8 | 87 | 87 | 0 | 3 | 0 | 0 | 3 |
[20] † Pachyrhinosaurus perotorum Taxon name last Modified on 12/05/13 | 43 | 53 | 52 | 0 | 3 | 0 | 0 | 3 |
[21] † Chasmosaurus belli Taxon name last Modified on 12/05/13 | 1 | 93 | 92 | 0 | 5 | 0 | 0 | 3 |
[22] † Pentaceratops sternbergii Taxon name last Modified on 12/05/13 | 1 | 92 | 92 | 0 | 5 | 0 | 0 | 3 |
[23] † Avaceratops lammersi Taxon name last Modified on 12/05/13 | 19 | 74 | 74 | 0 | 5 | 0 | 0 | 3 |
[24] † Nasutoceratops titusi Taxon name last Modified on 12/05/13 | 27 | 68 | 68 | 0 | 3 | 0 | 0 | 3 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 399 | |
Project downloads | 373 | |
Matrix downloads | 17 | Nasutoceratops phylogeny (17 downloads); |
Document downloads | 8 | Nasutoceratops characters (4 downloads); Nasutoceratops matrix Excel (2 downloads); Nasutoceratops matrix TNT (2 downloads); |
Media downloads | 1 | M307812 (1 download); |