Project 964: S. D. Sampson, E. K. Lund, M. A. Loewen, A. A. Farke, K. E. Clayton. 2013. A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia. Proceedings of the Royal Society B: Biological Sciences. 280 (1766).
Abstract
The fossil record of centrosaurine ceratopsids is largely restricted to the northern region of western North America (Alberta, Montana, Alaska). Exceptions consist of single taxa from Utah (Diabloceratops) and China (Sinoceratops), plus otherwise fragmentary remains from southern Laramidia. Here we describe a remarkable new taxon, Nasutoceratops titusi n. gen. et sp., from the late Campanian Kaiparowits Formation of Utah, represented by multiple specimens, including a nearly complete skull and partial postcranial skeleton. Autapomorphies include an enlarged narial region, pneumatic nasal ornamentation, abbreviated snout, and elongate, rostrolaterally directed supraorbital horncores. The subrectangular parietosquamosal frill is relatively unadorned and broadest in the mid-region. A phylogenetic analysis indicates that Nasutoceratops is the sister taxon to Avaceratops, and that a previously unknown subclade of centrosaurines branched off early in the group’s history and persisted for several million years during the late Campanian. As the first well-represented southern centrosaurine comparable in age to the bulk of northern forms, Nasutoceratops provides strong support for the provincialism hypothesis, which posits that Laramidia—the western landmass formed by inundation of the central region of North America by the Western Interior Seaway—hosted at least two coeval dinosaur communities for over a million years of late Campanian time.Read the article »
Project DOI: 10.7934/P964, http://dx.doi.org/10.7934/P964
This project contains | Matrices |
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Download Project SDD File ![]() | Total scored cells: 1684 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 98 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 396 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:98/0 |
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MorphoBank Project 964
MorphoBank Project 964
- Creation Date:
03 June 2013 - Publication Date:
10 September 2013 - Project views: 38197
- Media downloads: 1
- Matrix downloads: 16
Authors' Institutions
- Ohio University
- University of Utah
- Denver Museum of Nature and Science
- Natural History Museum of Utah
- Raymond Alf Museum of Paleontology (The Webb Schools)
Members
member name | taxa ![]() |
specimens ![]() |
media ![]() | media notes | chars ![]() | character
| cell scorings (scored, NPA, "-") | cell
| rules ![]() | ||||||||||||
Mark Loewen Project Administrator | 24 | 1 | 1 | 1 | 98 | 0 | 0 | 0 | 0 | 1684 (1480, 0, 204) | 0 | 0 | 0 | 0 | |||||||
Maureen Admin Full membership ![]() | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | 0 | |||||||
MorphoBank Curator Full membership ![]() | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M2151' (24 Taxa)
taxon | unscored cells |
scored cells ![]() |
no cell support ![]() |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Leptoceratops gracilis Last Modified in 12/05/13 | 1 | 60 | 60 | 0 | 37 | 0 | 0 | 3 ![]() |
[2] † Protoceratops andrewsi Last Modified in 12/05/13 | 2 | 66 | 66 | 0 | 30 | 0 | 0 | 3 ![]() |
[3] † Magnirostris dodsoni Last Modified in 12/05/13 | 33 | 35 | 35 | 0 | 30 | 0 | 0 | 3 ![]() |
[4] † Bagaceratops rozhdestvenskyi Last Modified in 12/05/13 | 26 | 38 | 38 | 0 | 34 | 0 | 0 | 3 ![]() |
[5] † Turanoceratops tardabilis Last Modified in 12/05/13 | 80 | 14 | 14 | 0 | 4 | 0 | 0 | 3 ![]() |
[6] † Zuniceratops christopheri Last Modified in 12/05/13 | 25 | 53 | 53 | 0 | 20 | 0 | 0 | 3 ![]() |
[7] † Diabloceratops eatoni Last Modified in 12/05/13 | 31 | 66 | 66 | 0 | 1 | 0 | 0 | 3 ![]() |
[8] † Albertaceratops nesmoi Last Modified in 12/05/13 | 37 | 59 | 59 | 0 | 2 | 0 | 0 | 3 ![]() |
[9] † Rubeosaurus ovatus Last Modified in 12/05/13 | 62 | 35 | 35 | 0 | 1 | 0 | 0 | 3 ![]() |
[10] † Styracosaurus albertensis Last Modified in 12/05/13 | 2 | 94 | 94 | 0 | 2 | 0 | 0 | 3 ![]() |
[11] † Spinops sternbergorum Last Modified in 12/05/13 | 62 | 34 | 34 | 0 | 2 | 0 | 0 | 3 ![]() |
[12] † Centrosaurus apertus Last Modified in 12/05/13 | 0 | 96 | 96 | 0 | 2 | 0 | 0 | 3 ![]() |
[13] † Coronosaurus brinkmani Last Modified in 12/05/13 | 13 | 83 | 83 | 0 | 2 | 0 | 0 | 3 ![]() |
[14] † Xenoceratops foremostensis Last Modified in 12/05/13 | 73 | 24 | 24 | 0 | 1 | 0 | 0 | 3 ![]() |
[15] † Sinoceratops zhuchengensis Last Modified in 12/05/13 | 47 | 49 | 48 | 0 | 3 | 0 | 0 | 3 ![]() |
[16] † Einiosaurus procurvicornis Last Modified in 12/05/13 | 17 | 79 | 78 | 0 | 3 | 0 | 0 | 3 ![]() |
[17] † Achelousaurus horneri Last Modified in 12/05/13 | 26 | 69 | 69 | 0 | 3 | 0 | 0 | 3 ![]() |
[18] † Pachyrhinosaurus canadensis Last Modified in 12/05/13 | 36 | 59 | 59 | 0 | 3 | 0 | 0 | 3 ![]() |
[19] † Pachyrhinosaurus lakustai Last Modified in 12/05/13 | 8 | 87 | 87 | 0 | 3 | 0 | 0 | 3 ![]() |
[20] † Pachyrhinosaurus perotorum Last Modified in 12/05/13 | 43 | 53 | 52 | 0 | 3 | 0 | 0 | 3 ![]() |
[21] † Chasmosaurus belli Last Modified in 12/05/13 | 1 | 93 | 92 | 0 | 5 | 0 | 0 | 3 ![]() |
[22] † Pentaceratops sternbergii Last Modified in 12/05/13 | 1 | 92 | 92 | 0 | 5 | 0 | 0 | 3 ![]() |
[23] † Avaceratops lammersi Last Modified in 12/05/13 | 19 | 74 | 74 | 0 | 5 | 0 | 0 | 3 ![]() |
[24] † Nasutoceratops titusi Last Modified in 12/05/13 | 27 | 68 | 68 | 0 | 3 | 0 | 0 | 3 ![]() |
Project views 
type | number of views | Individual items viewed (where applicable) |
Total project views | 38197 | |
Project overview | 4390 | |
Matrix views | 2830 | Matrix landing page (2488 views); Nasutoceratops phylogeny (342 views); |
Taxon list | 18045 | |
Media views | 6439 | Media search (5569 views); M307812 (870 views); |
Specimen list | 3104 | |
Bibliography | 1238 | |
Documents list | 2151 |
Project downloads 
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 401 | |
Project downloads | 376 | |
Matrix downloads | 16 | Nasutoceratops phylogeny (16 downloads); |
Document downloads | 8 | Nasutoceratops characters (4 downloads); Nasutoceratops matrix Excel (2 downloads); Nasutoceratops matrix TNT (2 downloads); |
Media downloads | 1 | M307812 (1 download); |