Project 979: G. Han,L. M. Chiappe,S. Ji,M. Habib,A. H. Turner,A. Chinsamy,X. Liu,L. Han. 2014. A new raptorial dinosaur with exceptionally long feathering provides insights into dromaeosaurid flight performance. Nature Communications. 5.
Specimen: Changyuraptor (HG:B016)

Abstract

Microraptorines are a group of predatory dromaeosaurid theropod dinosaurs with aero- dynamic capacity. These close relatives of birds are essential for testing hypotheses explaining the origin and early evolution of avian flight. Here we describe a new ‘four-winged’ microraptorine, Changyuraptor yangi, from the Early Cretaceous Jehol Biota of China. With tail feathers that are nearly 30 cm long, roughly 30% the length of the skeleton, the new fossil possesses the longest known feathers for any non-avian dinosaur. Furthermore, it is the largest theropod with long, pennaceous feathers attached to the lower hind limbs (that is, ‘hindwings’). The lengthy feathered tail of the new fossil provides insight into the flight performance of microraptorines and how they may have maintained aerial competency at larger body sizes. We demonstrate how the low-aspect-ratio tail of the new fossil would have acted as a pitch control structure reducing descent speed and thus playing a key role in landing.


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Project DOI: 10.7934/P979, http://dx.doi.org/10.7934/P979
This project containsMatrices
  • 1 Media
  • 1 Matrix
  • 1 Document
  • 119 Taxa
  • 1 Specimen
  • 477 Characters


Download Project SDD File
Total scored cells: 20977
Total media associated with cells: 0
Total labels associated with cell media: 0
Characters
Total characters: 477
Total characters with associated media: 0
Total characters with media with labels: 0
Total character states: 1127
Total character states with associated media: 0
Total character states with media with labels:0
Total unordered/ordered characters:477/0
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MorphoBank Project 979


    Members

    member name taxa specimens media chars character
    media
    labels
    cell scorings
    (scored, NPA, "-")
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    medialabels
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    Alan Turner
    Project Administrator
    Last logged in 07/16/14
    119114770020977
    (19380, 0, 1597)
    000


    Taxonomic Overview for Matrix 'M2175' (119 Taxa)

    taxon unscored
    cells
    scored
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    [1] Allosaurus fragilis
    Last Modified in 12/05/13
    96361359022001
    [2] Sinraptor dongi
    Last Modified in 12/05/13
    171294293013001
    [3] Ingenia yanshani
    Last Modified in 12/05/13
    257194193027001
    [4] Citipati osmolskae
    Last Modified in 12/05/13
    105342339033001
    [5] Rinchenia mongoliensis
    Last Modified in 12/05/13
    3589999020001
    [6] Oviraptor philoceratops
    Last Modified in 12/05/13
    316136136025001
    [7] Conchoraptor gracilis
    Last Modified in 12/05/13
    242214214021001
    [8] Incisivosaurus gauthieri
    Last Modified in 12/05/13
    31315615608001
    [9] Microvenator celer
    Last Modified in 12/05/13
    359107107011001
    [10] Chirostenotes pergracilis
    Last Modified in 12/05/13
    291163162024001
    [11] Caudipteryx zoui
    Last Modified in 12/05/13
    260192191026001
    [12] Dromaeosaurus albertensis
    Last Modified in 12/05/13
    33014214205001
    [13] Deinonychus antirrhopus
    Last Modified in 12/05/13
    171295294012001
    [14] Velociraptor mongoliensis
    Last Modified in 12/05/13
    83381379015001
    [15] Balaur bondoc
    Last Modified in 12/05/13
    36810410207001
    [16] Atrociraptor marshalli
    Last Modified in 12/05/13
    439393800001
    [17] Utahraptor
    Last Modified in 12/05/13
    391858402001
    [18] Adasaurus mongoliensis
    Last Modified in 12/05/13
    28518718507001
    [19] Achillobator giganticus
    Last Modified in 12/05/13
    36311011004001
    [20] Tsaagan mangas
    Last Modified in 12/05/13
    27819119108001
    [21] Saurornitholestes langsto
    Last Modified in 12/05/13
    35112512501001
    [22] Bambiraptor feinbergorum
    Last Modified in 12/05/13
    141328326010001
    [23] Tianyuraptor ostromi
    Last Modified in 12/05/13
    34113013006001
    [24] Sinornithosaurus millenii
    Last Modified in 12/05/13
    250217216011001
    [25] Microraptor zhaoianus
    Last Modified in 12/05/13
    25821521405001
    [26] Changyuraptor
    Last Modified in 07/16/14
    398797900001
    [27] Graciliraptor lujiatunensis
    Last Modified in 12/05/13
    428464504001
    [28] Hesperonychus elizabethae
    Last Modified in 12/05/13
    461161600001
    [29] Pyroraptor olympius
    Last Modified in 12/05/13
    465121200001
    [30] Rahonavis ostromi
    Last Modified in 12/05/13
    35811811702001
    [31] Buitreraptor gonzalozorum
    Last Modified in 12/05/13
    26121120709001
    [32] Neuquenraptor + unenlagia
    Last Modified in 12/05/13
    37410010003001
    [33] Austroraptor
    Last Modified in 12/05/13
    414636102001
    [34] Unenlagia
    Last Modified in 12/05/13
    396797902001
    [35] Unenlagia comahuensis
    Last Modified in 12/05/13
    400757502001
    [36] Unenlagia paynemili
    Last Modified in 12/05/13
    457201901001
    [37] Neuquenraptor argentinus
    Last Modified in 12/05/13
    452242401001
    [38] Shanag ashile
    Last Modified in 12/05/13
    439363602001
    [39] Mahakala omnogovae
    Last Modified in 12/05/13
    33014113908001
    [40] Mononykus olecranus
    Last Modified in 12/05/13
    316135135026001
    [41] Shuvuuia deserti
    Last Modified in 12/05/13
    126319317034001
    [42] Patagonykus puertai
    Last Modified in 12/05/13
    3848381012001
    [43] Albinykus
    Last Modified in 12/05/13
    443323202001
    [44] Alvarezsaurus calvoi
    Last Modified in 12/05/13
    423545301001
    [45] Proceratosaurus bradleyi
    Last Modified in 12/05/13
    395808002001
    [46] Ornitholestes hermanni
    Last Modified in 12/05/13
    230233231016001
    [47] Coelurus fragilis
    Last Modified in 12/05/13
    393818103001
    [48] Archaeopteryx lithographi
    Last Modified in 12/05/13
    15431931805001
    [49] Avimimus portentosus
    Last Modified in 12/05/13
    309152152016001
    [50] Confuciusornis sanctus
    Last Modified in 12/05/13
    208260257012001
    [51] Struthiomimus altus
    Last Modified in 12/05/13
    164282281032001
    [52] Gallimimus bullatus
    Last Modified in 12/05/13
    116325323038001
    [53] Garudimimus brevipes
    Last Modified in 12/05/13
    282173173022001
    [54] Pelecanimimus polydon
    Last Modified in 12/05/13
    3837878016001
    [55] Harpymimus okladnikovi
    Last Modified in 12/05/13
    342114113022001
    [56] Troodon formosus
    Last Modified in 12/05/13
    29917517503001
    [57] Saurornithoides mongolien
    Last Modified in 12/05/13
    32114714709001
    [58] Saurornithoides junior
    Last Modified in 12/05/13
    32914314206001
    [59] Xixiasaurus
    Last Modified in 12/05/13
    413616103001
    [60] Byronosaurus jaffei
    Last Modified in 12/05/13
    36210910906001
    [61] Sinornithoides youngi
    Last Modified in 12/05/13
    285174173019001
    [62] Sinovenator changii
    Last Modified in 12/05/13
    192276274011001
    [63] Mei long
    Last Modified in 12/05/13
    200265264013001
    [64] EK troodontid (igm 100/44
    Last Modified in 12/05/13
    444333300001
    [65] Jinfengopteryx elegans
    Last Modified in 12/05/13
    36510610606001
    [66] IGM 100/1126
    Last Modified in 12/05/13
    242223222013001
    [67] IGM 100/1323
    Last Modified in 12/05/13
    328140139010001
    [68] † Aurornis xui
    Last Modified in 12/05/13
    380959502001
    [69] Anchiornis huxleyi
    Last Modified in 12/05/13
    236219218023001
    [70] Xiaotingia
    Last Modified in 12/05/13
    31216516500001
    [71] Segnosaurus galbinensis
    Last Modified in 12/05/13
    380929205001
    [72] Erlikosaurus andrewsi
    Last Modified in 12/05/13
    296160160021001
    [73] Alxasaurus elesitaiensis
    Last Modified in 12/05/13
    353115114010001
    [74] Tyrannosaurus rex
    Last Modified in 12/05/13
    102357356019001
    [75] Tarbosaurus baatar
    Last Modified in 12/05/13
    407656505001
    [76] Albertosaurus sacrophagus
    Last Modified in 12/05/13
    18228828807001
    [77] Gorgosaurus libratus
    Last Modified in 12/05/13
    125341341011001
    [78] Daspletosaurus
    Last Modified in 12/05/13
    15531631606001
    [79] Eotyrannus lengi
    Last Modified in 12/05/13
    390818106001
    [80] Dilong paradoxus
    Last Modified in 12/05/13
    191272271015001
    [81] Shenzhousaurus orientalis
    Last Modified in 12/05/13
    303136134040001
    [82] Ornithomimus edmonticus
    Last Modified in 12/05/13
    152287286039001
    [83] Archaeornithomimus asiati
    Last Modified in 12/05/13
    315143141021001
    [84] Anserimimus planinychus
    Last Modified in 12/05/13
    404707003001
    [85] Huaxiagnathus orientalis
    Last Modified in 12/05/13
    273184183021001
    [86] Sinosauropteryx prima
    Last Modified in 12/05/13
    264196195018001
    [87] Compsognathus longipes
    Last Modified in 12/05/13
    222232230025001
    [88] Juravenator starki
    Last Modified in 12/05/13
    270181181026001
    [89] Jeholornis prima
    Last Modified in 12/05/13
    252200200025001
    [90] Jixiangornis orientalis
    Last Modified in 12/05/13
    225227226026001
    [91] Yanornis martini
    Last Modified in 12/05/13
    31815915702001
    [92] Apsaravis ukhaana
    Last Modified in 12/05/13
    303156154020001
    [93] Yixianornis
    Last Modified in 12/05/13
    239229227011001
    [94] Sapeornis
    Last Modified in 12/05/13
    219234233025001
    [95] Neuquenornis volans
    Last Modified in 12/05/13
    384929102001
    [96] Patagopteryx deferrariisi
    Last Modified in 12/05/13
    36710710604001
    [97] Cathayornis
    Last Modified in 12/05/13
    33314113707001
    [98] Concornis
    Last Modified in 12/05/13
    3751029903001
    [99] Gobipteryx
    Last Modified in 12/05/13
    403656509001
    [100] Vorona
    Last Modified in 12/05/13
    443333301001
    [101] Songlingornis
    Last Modified in 12/05/13
    432464401001
    [102] Pengornis houi
    Last Modified in 12/05/13
    323141140014001
    [103] Hesperornis
    Last Modified in 12/05/13
    192235235050001
    [104] Baptornis
    Last Modified in 12/05/13
    430484700001
    [105] Ichthyornis
    Last Modified in 12/05/13
    237225222018001
    [106] Iaceornis marshii
    Last Modified in 12/05/13
    419575602001
    [107] Limenavis patagonica
    Last Modified in 12/05/13
    4689900001
    [108] Crypturellus undulatus
    Last Modified in 12/05/13
    92328327058001
    [109] Lithornis
    Last Modified in 12/05/13
    218234229030001
    [110] Gallus gallus
    Last Modified in 12/05/13
    93332330054001
    [111] Crax pauxi
    Last Modified in 12/05/13
    97336336044001
    [112] Anas platyrhynchus
    Last Modified in 12/05/13
    106320319052001
    [113] Chauna torquata
    Last Modified in 12/05/13
    95339338044001
    [114] Hongshanornis longicrest
    Last Modified in 12/05/13
    304152148025001
    [115] Liaoningornis longidigitu
    Last Modified in 12/05/13
    441353501001
    [116] Pedopenna
    Last Modified in 12/05/13
    460161601001
    [117] Epidendrosaurus
    Last Modified in 12/05/13
    418545405001
    [118] Epidexipteryx
    Last Modified in 12/05/13
    355113112010001
    [119] Haplocheirus
    Last Modified in 12/05/13
    243221221013001


    Project views

    type number of views Individual items viewed (where applicable)
    Total project views24782
    Project overview3154
    Matrix views2336Matrix landing page (2111 views); Changyuraptor (225 views);
    Media views4447Media search (3870 views); M334034 (577 views);
    Taxon list11316
    Documents list1752
    Specimen list1777




    Project downloads

    type number of downloads Individual items downloaded (where applicable)
    Total downloads from project322
    Matrix downloads32Changyuraptor (32 downloads);
    Project downloads284
    Document downloads6Analysis README file (6 downloads);