Project 3216: M. I. Yavorskaya, E. Anton, P. Jałoszyński, A. Polilov, R. G. Beutel. 2018. Cephalic anatomy of Sphaeriusidae and a morphology-based phylogeny of the suborder Myxophaga (Coleoptera). Systematic Entomology. 43 (4):777-797.
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