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Garzon-Orduña, I. J. 2012. Phylogenetic evidence for loss of sound production and a shift in sexual recognition signals in Hamadryas butterflies (Nymphalidae: Biblidinae). Systematic Entomology. Systematic Entomology. Vol. 37, pp. 84-101.
Garzón-Orduña, Ivonne J., Marini-Filho, Onildo, Johnson, Steve G. and Penz, Carla M. 2013. Phylogenetic relationships of Hamadryas (Nymphalidae: Biblidinae) based on the combined analysis of morphological and molecular data. Cladistics. Vol. 29(6), pp. 629-642.