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Project 4306: F. J. Degrange, D. T. Ksepka, C. P. Tambussi. 2018. Redescription of the oldest crown clade penguin: cranial osteology, jaw myology, neuroanatomy, and phylogenetic affinities of Madrynornis mirandus. Journal of Vertebrate Paleontology. 38 (2):e1445636.
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Degrange, F. J., Ksepka, D. T. and Tambussi, C. P. 2018. Redescription of the oldest crown clade penguin: cranial osteology, jaw myology, neuroanatomy, and phylogenetic affinities of Madrynornis mirandus. Journal of Vertebrate Paleontology. Vol. 38(2), p. e1445636.