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Ksepka, D. T., Field, D. J., Heath, T. A., Pett, W., Thomas, D. B., Giovanardi, S. and Tennyson, A. J. 2023. Largest-known fossil penguin provides insight into the early evolution of sphenisciform body size and flipper anatomy. Journal of Paleontology. Vol. 97(2), pp. 434-453.