Project 4306: F. J. Degrange, D. T. Ksepka, C. P. Tambussi. 2018. Redescription of the oldest crown clade penguin: cranial osteology, jaw myology, neuroanatomy, and phylogenetic affinities of Madrynornis mirandus. Journal of Vertebrate Paleontology. 38 (2):e1445636.

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