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Project 3932: E. Vlachos, G. L. Georgalis, S. Roussiakis, M. Böhme, G. Theodorou. 2019. The Pikermian tortoises (Testudines, Testudinidae) from the late Miocene of the South Balkans. Journal of Vertebrate Paleontology. 39 (5):e1711520.
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M739118
† Hesperotestudo osborniana
ventral